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<document id="F8DF0A313E3FC290D0C21A7C7743729F" ID-DOI="10.11646/zootaxa.5486.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="13209973" ID-ZooBank="F172702B-ED72-41FA-BF71-6515D2459003" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722788125309" checkinUser="plazi" docAuthor="Abhishek, Venkateshaiah, David, Karamankodu Jacob &amp; Pradeep, Shivanand" docDate="2024" docId="039487C7ED2EFF891C9EF85CFAEAF895" docLanguage="en" docName="zootaxa.5486.2.2.6.pdf" docOrigin="Zootaxa 5486 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5486.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bactrocera (Bactrocera) ettinabhuja Abhishek and David 2024, sp. nov." docType="treatment" docUuid="0709E585-C164-44A0-8CD7-BBB6F25F670E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="287" masterDocId="FFADFFBFED2DFF8D1C09FFCAFF8EFFE5" masterDocTitle="Two new species and a new record of Bactrocera Macquart (Diptera: Tephritidae: Dacinae: Dacini) from India with an updated key to species of subgenus Bactrocera" masterLastPageNumber="300" masterPageNumber="283" pageNumber="286" updateTime="1722788913681" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3842CDDAEA39D34326D9CED561EEC6F5">Two new species and a new record of Bactrocera Macquart (Diptera: Tephritidae: Dacinae: Dacini) from India with an updated key to species of subgenus Bactrocera</mods:title>
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<mods:namePart id="63E2EB323EA5EA1EFAAE1F84690C8C3B">Abhishek, Venkateshaiah</mods:namePart>
<mods:affiliation id="5D05F91F85AD69B212A85E6574013FFA">ICAR-National Bureau of Agricultural Insect Resources, Bengaluru- 560024, Karnataka, India. &amp; Keladi Shivappa Nayaka University of Agricultural and Horticultural Sciences, Shivamogga, Karnataka, India</mods:affiliation>
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<mods:namePart id="DD53BC3E74F1E35DC12754C17EAAFC21">David, Karamankodu Jacob</mods:namePart>
<mods:affiliation id="0472C93D543A016A7EC9EEE6110907EF">ICAR-National Bureau of Agricultural Insect Resources, Bengaluru- 560024, Karnataka, India.</mods:affiliation>
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<mods:namePart id="84D460A062E27F2A472395F839459EA7">Pradeep, Shivanand</mods:namePart>
<mods:affiliation id="1571070EED78A55EECB0AF5A90B576CE">ICAR-National Bureau of Agricultural Insect Resources, Bengaluru- 560024, Karnataka, India. &amp; Keladi Shivappa Nayaka University of Agricultural and Horticultural Sciences, Shivamogga, Karnataka, India &amp; ICAR-National Bureau of Agricultural Insect Resources, Bengaluru- 560024, Karnataka, India.</mods:affiliation>
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<mods:date id="604CA5B78D38571490DD0D9D2DADD64A">2024</mods:date>
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<paragraph id="8B8236D1ED2EFF8E1C9EF85CFBE0F84A" blockId="3.[151,1134,1941,2004]" box="[151,1134,1941,1968]" pageId="3" pageNumber="286">
<heading id="D0CA81BDED2EFF8E1C9EF85CFBE0F84A" bold="true" box="[151,1134,1941,1968]" fontSize="11" level="1" pageId="3" pageNumber="286" reason="1">
<emphasis id="B949EAC3ED2EFF8E1C9EF85CFBE0F84A" bold="true" box="[151,1134,1941,1968]" pageId="3" pageNumber="286">
<taxonomicName id="4C3D4D52ED2EFF8E1C9EF85CFCD9F84A" authority="Abhishek and David" authorityName="Abhishek and David" authorityYear="2024" box="[151,855,1941,1968]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="ettinabhuja" status="sp. nov." subGenus="Bactrocera">
<emphasis id="B949EAC3ED2EFF8E1C9EF85CFE93F855" bold="true" box="[151,285,1942,1968]" italics="true" pageId="3" pageNumber="286">Bactrocera</emphasis>
(
<emphasis id="B949EAC3ED2EFF8E1D27F85CFE3AF855" bold="true" box="[302,436,1942,1968]" italics="true" pageId="3" pageNumber="286">Bactrocera</emphasis>
)
<emphasis id="B949EAC3ED2EFF8E1DCCF85FFDDEF855" bold="true" box="[453,592,1941,1968]" italics="true" pageId="3" pageNumber="286">ettinabhuja</emphasis>
Abhishek and David
</taxonomicName>
,
<taxonomicNameLabel id="A27A57B8ED2EFF8E1F6AF85FFC4FF84A" box="[867,961,1941,1967]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13062A54ED2EFF8E1FD9F85FFBEBF84A" box="[976,1125,1941,1967]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="3" pageNumber="286">Figures 18</figureCitation>
)
</emphasis>
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</paragraph>
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<emphasis id="B949EAC3ED29FF891C9EFF5CFEE1FF55" bold="true" box="[151,367,150,176]" pageId="4" pageNumber="287">Description: Male</emphasis>
</paragraph>
<paragraph id="8B8236D1ED29FF891C9EFF70FBA9FF31" blockId="4.[151,1437,150,1905]" box="[151,1063,186,213]" pageId="4" pageNumber="287">
A medium-sized fly, (total body length
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), reddish-brown to black species.
</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFF14FF62FE11" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
Head (
<figureCitation id="13062A54ED29FF891D1AFF14FED6FF1D" box="[275,344,222,248]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Fig. 1</figureCitation>
): Face
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high and
<quantity id="4CC59B34ED29FF891E80FF14FD61FF1C" box="[649,751,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="4" pageNumber="287" unit="mm" value="1.88">1.88 mm</quantity>
wide, frons
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high and
<quantity id="4CC59B34ED29FF891852FF14FB4EFF1C" box="[1115,1216,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="287" unit="mm" value="0.58">0.58 mm</quantity>
wide, face entirely fulvous with the oral margin
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high and
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wide with elongate facial oval spots of
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high and
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wide, frons with three frontal setae and two orbital setae, bases of all setae black, ocellar triangle black, ocellus fuscous, outer vertical seta and medial vertical seta well developed; prominent fuscous markings on gena connected to eye margin, occiput black except posterior margin of eye and vertex, postocular setae thin, lunule fuscous; antennal segments dark fuscous on outer side, arista not plumose, combined length of antennal segments viz., scape (
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), pedicel (
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) and flagellomere (
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) almost equal to the vertical length (
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) of face.
</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFE34FDBCFD15" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
Thorax (
<figureCitation id="13062A54ED29FF891D2CFE34FE09FDFD" box="[293,391,510,536]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Figs 2, 3</figureCitation>
): Scutum reddish brown to black (
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high and
<quantity id="4CC59B34ED29FF891FD8FE34FBBDFDFC" box="[977,1075,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="4" pageNumber="287" unit="mm" value="2.05">2.05 mm</quantity>
wide) with lanceolate markings; lateral postsutural vitta narrow (
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) obscured beyond postsutural supra-alar seta, ending at or before intra-alar setae and medial postsutural vittae absent. Yellow markings as follows, notopleuron, postpronotal lobe, narrow mesopleural (anepisternal) stripe (
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high and
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wide), almost equal to notopleuron, continued as yellow transverse spot on katepisternum, 0.5 of katatergite and anatergite. Scutellum yellow (
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long,
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wide) with a narrow (
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) black basal band, subscutellum black with
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high and
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wide and with thin medial reddish-brown line.
</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFD30FB75FCDC" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">Chaetotaxy: 2 scapular setae; 1 anterior notopleural seta; 1 posterior notopleural seta; 1 mesopleural seta; 1 post sutural supra-alar seta; 1 prescutellar setae; 1 post-alar seta; 1 intra-alar seta; 1 apical scutellar setae.</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFC89FDA3FC65" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
Legs (
<figureCitation id="13062A54ED29FF891D04FC88FEDCFCB9" box="[269,338,834,860]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Fig. 3</figureCitation>
): Coxa, trochanter, tibia of fore and hind legs, 0.4 of tibia of mid legs, tarsus and pretarsus of mid legs are fuscous; remainder fulvous.
</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFC40FB3FFC2C" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
Wing (
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): (
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) predominantly hyaline, cell bc and c colourless, narrow costal band from cell sc to wing apex confluent with vein R
<subScript id="17B93494ED29FF891E0CFC0BFD91FC2A" attach="left" box="[517,543,961,975]" fontSize="6" pageId="4" pageNumber="287">2+3</subScript>
, extension of cell cup longer than cup, anal streak present.
</paragraph>
<paragraph id="8B8236D1ED29FF891CC7FC18FBF9FBBD" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
Abdomen (
<figureCitation id="13062A54ED29FF891D44FC18FE1AFC09" box="[333,404,978,1004]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Fig. 4</figureCitation>
): Abdomen oval (
<quantity id="4CC59B34ED29FF891E69FC18FD49FC08" box="[608,711,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.23" pageId="4" pageNumber="287" unit="mm" value="3.23">3.23 mm</quantity>
long;
<quantity id="4CC59B34ED29FF891F18FC18FCF9FC08" box="[785,887,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.41" pageId="4" pageNumber="287" unit="mm" value="2.41">2.41 mm</quantity>
wide) predominantly red-brown to black except fulvous band on tergites II transversely, pecten on tergite III and a brown medial patch gradually expanding over tergites III-V, with dense white hair over the medial region, and two large spots on tergite V, sternite I, II and V are fuscous and II and III are fulvous, sternite V of male with a deep posterior emargination.
</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFBA8FD43FAB0" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
Male genitalia (
<figureCitation id="13062A54ED29FF891D7EFBA8FE76FB99" box="[375,504,1122,1149]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Figs 6, 7, 8</figureCitation>
,
<figureCitation id="13062A54ED29FF891E0CFBA8FDADFB99" box="[517,547,1122,1148]" captionStart="FIGURES4144" captionStartId="12.[151,264,1204,1228]" captionTargetBox="[151,1436,181,1180]" captionTargetId="figure-24@12.[151,1436,181,1180]" captionTargetPageId="12" captionText="FIGURES4144. Comparisonbetween theepandrium(lateral view).41, Bactrocera(Bactrocera) ettinabhujaAbhishek &amp;David, sp. nov.; 42, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 43, Bactrocera (Bactrocera) paraosbeckiae Drew.; 44, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209993" httpUri="https://zenodo.org/record/13209993/files/figure.png" pageId="4" pageNumber="287">41</figureCitation>
,
<figureCitation id="13062A54ED29FF891E26FBA8FDC3FB98" box="[559,589,1122,1149]" captionStart="FIGURES 4548" captionStartId="13.[151,264,1761,1785]" captionTargetBox="[151,1436,182,1733]" captionTargetId="figure-17@13.[151,1436,181,1737]" captionTargetPageId="13" captionText="FIGURES 4548. Comparison between the epandrium (posterior view). 45, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 46, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 47, Bactrocera (Bactrocera) paraosbeckiae Drew.; 48, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209995" httpUri="https://zenodo.org/record/13209995/files/figure.png" pageId="4" pageNumber="287">45</figureCitation>
,
<figureCitation id="13062A54ED29FF891E50FBA8FDF4FB99" box="[601,634,1122,1148]" captionStart="FIGURES 4952" captionStartId="14.[151,264,994,1018]" captionTargetBox="[151,1437,181,970]" captionTargetId="figure-24@14.[151,1437,181,970]" captionTargetPageId="14" captionText="FIGURES 4952. Comparison between the glans of phallus. 49, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 50, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 51, Bactrocera (Bactrocera) paraosbeckiae Drew; 52, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209999" httpUri="https://zenodo.org/record/13209999/files/figure.png" pageId="4" pageNumber="287">49</figureCitation>
): Epandrium and surstyli inverted U-shaped in outline (posterior view), strongly sclerotised, epandrium quadrate, as long as wide (
<quantity id="4CC59B34ED29FF891F23FB4CFC1EFB44" box="[810,912,1158,1185]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="287" unit="mm" value="0.28">0.28 mm</quantity>
high;
<quantity id="4CC59B34ED29FF891FDEFB4CFBB2FB44" box="[983,1084,1158,1185]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="287" unit="mm" value="0.26">0.26 mm</quantity>
wide) equal to length of lateral surstylus (
<quantity id="4CC59B34ED29FF891D00FB60FEFCFB21" box="[265,370,1194,1220]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="287" unit="mm" value="0.27">0.27 mm</quantity>
), lateral surstylus broad (
<quantity id="4CC59B34ED29FF891E83FB60FD6AFB21" box="[650,740,1194,1220]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="287" unit="mm" value="0.1">0.1 mm</quantity>
), posterior lobe of lateral surstylus, pointed, curved back (
<quantity id="4CC59B34ED29FF891962FB60FF49FB0D" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="287" unit="mm" value="0.11">0.11 mm</quantity>
), not longer than anterior lobe (
<quantity id="4CC59B34ED29FF891E27FB04FD16FB0D" box="[558,664,1230,1257]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="287" unit="mm" value="0.25">0.25 mm</quantity>
); proctiger membraneous (
<quantity id="4CC59B34ED29FF891FCEFB04FBBFFB0D" box="[967,1073,1230,1256]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="287" unit="mm" value="0.06">0.06 mm</quantity>
), dome shaped and smaller than epandrium, medial surstylus as long as lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus elongate (
<quantity id="4CC59B34ED29FF891D0FFADCFEE0FAD5" box="[262,366,1302,1328]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" pageId="4" pageNumber="287" unit="mm" value="2.63">2.63 mm</quantity>
) with well-developed glans (
<quantity id="4CC59B34ED29FF891EBEFADCFC91FAD5" box="[695,799,1302,1328]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="287" unit="mm" value="0.43">0.43 mm</quantity>
) (
<figureCitation id="13062A54ED29FF891F3FFADCFCF1FAD5" box="[822,895,1302,1328]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Figs 8</figureCitation>
,
<figureCitation id="13062A54ED29FF891F84FADCFC23FAD5" box="[909,941,1302,1328]" captionStart="FIGURES 4952" captionStartId="14.[151,264,994,1018]" captionTargetBox="[151,1437,181,970]" captionTargetId="figure-24@14.[151,1437,181,970]" captionTargetPageId="14" captionText="FIGURES 4952. Comparison between the glans of phallus. 49, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 50, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 51, Bactrocera (Bactrocera) paraosbeckiae Drew; 52, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209999" httpUri="https://zenodo.org/record/13209999/files/figure.png" pageId="4" pageNumber="287">49</figureCitation>
); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present.
</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFA94FE70FA9D" blockId="4.[151,1437,150,1905]" box="[199,510,1374,1400]" pageId="4" pageNumber="287">Attractant: Methyl eugenol.</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFA49FE41FA79" blockId="4.[151,1437,150,1905]" box="[199,463,1410,1437]" pageId="4" pageNumber="287">Host plant: Not known.</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFA6CFF66FA01" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
<materialsCitation id="3B553C8CED29FF891CCEFA6CFF66FA01" collectingDate="2023-10-14" collectionCode="V" country="India" county="Chikkamagaluru" location="Abhishek" municipality="Ettinabhuja" pageId="4" pageNumber="287" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="holotype">
Material examined:
<typeStatus id="54868873ED29FF891DA2FA6CFD98FA25" box="[427,534,1446,1472]" pageId="4" pageNumber="287" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D3BFD58ED29FF891E2BFA6CFDC8FA25" box="[546,582,1446,1472]" count="1" pageId="4" pageNumber="287" type="male">1♂</specimenCount>
,
<collectingCountry id="F32A7641ED29FF891E5DFA6CFD28FA25" box="[596,678,1446,1472]" name="India" pageId="4" pageNumber="287">INDIA</collectingCountry>
,
<collectingRegion id="49F9F833ED29FF891EBBFA6CFCA6FA25" box="[690,808,1446,1472]" country="India" name="Karnataka" pageId="4" pageNumber="287">Karnataka</collectingRegion>
,
<collectingCounty id="62E34E5DED29FF891F3DFA6CFC7CFA25" box="[820,1010,1446,1472]" pageId="4" pageNumber="287">Chikkamagaluru</collectingCounty>
,
<collectingMunicipality id="6BE6ACABED29FF891FF4FA6CFB0FFA25" box="[1021,1153,1446,1472]" pageId="4" pageNumber="287">Ettinabhuja</collectingMunicipality>
,
<date id="FF831011ED29FF891884FA6CFA8EFA25" box="[1165,1280,1446,1472]" pageId="4" pageNumber="287" value="2023-10-14">
<collectingDate id="EFC7E9F9ED29FF891884FA6CFA8EFA25" box="[1165,1280,1446,1472]" pageId="4" pageNumber="287" value="2023-10-14">14.x.2023</collectingDate>
</date>
,
<location id="8EE2600AED29FF891902FA6CFAF7FA25" LSID="urn:lsid:plazi:treatment:039487C7ED2EFF891C9EF85CFAEAF895:8EE2600AED29FF891902FA6CFAF7FA25" box="[1291,1401,1446,1472]" country="India" county="Chikkamagaluru" municipality="Ettinabhuja" name="Abhishek" pageId="4" pageNumber="287" stateProvince="Karnataka">Abhishek</location>
,
<collectionCode id="ED2CAE14ED29FF89198CFA6DFA17FA24" box="[1413,1433,1447,1473]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="287" type="Museum">V</collectionCode>
. (NIM).
</materialsCitation>
</paragraph>
<paragraph id="8B8236D1ED29FF891CCEFA24FEBDF9C9" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
Etymology: This species is named after the
<typeStatus id="54868873ED29FF891EE2FA25FC95F9EC" box="[747,795,1519,1545]" pageId="4" pageNumber="287">type</typeStatus>
locality of the species and is a noun in apposition (Ettinabhuja).
</paragraph>
<paragraph id="8B8236D1ED29FF891CCEF9FCFAEAF895" blockId="4.[151,1437,150,1905]" pageId="4" pageNumber="287">
Diagnosis:
<taxonomicName id="4C3D4D52ED29FF891D4EF9FDFD6AF9B5" authorityName="Abhishek &amp; David &amp; Pradeep" authorityYear="2024" box="[327,740,1590,1616]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="ettinabhuja" status="sp. nov." subGenus="Bactrocera">
<emphasis id="B949EAC3ED29FF891D4EF9FDFE4AF9B5" box="[327,452,1591,1616]" italics="true" pageId="4" pageNumber="287">Bactrocera</emphasis>
(
<emphasis id="B949EAC3ED29FF891DDCF9FDFDDCF9B5" box="[469,594,1591,1616]" italics="true" pageId="4" pageNumber="287">Bactrocera</emphasis>
)
<emphasis id="B949EAC3ED29FF891E6DF9FCFD6AF9B5" box="[612,740,1590,1616]" italics="true" pageId="4" pageNumber="287">ettinabhuja</emphasis>
</taxonomicName>
<emphasis id="B949EAC3ED29FF891EE2F9FCFCC8F9B5" bold="true" box="[747,838,1590,1616]" pageId="4" pageNumber="287">
<taxonomicNameLabel id="A27A57B8ED29FF891EE2F9FCFCC8F9B5" box="[747,838,1590,1616]" pageId="4" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C3D4D52ED29FF891FD5F9FDFF5BF990" authority="Drew" authorityName="Drew, Drew and Raghu" authorityYear="2002" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="paraosbeckiae" subGenus="Bactrocera">
<emphasis id="B949EAC3ED29FF891FD5F9FDFBD7F9B5" box="[988,1113,1591,1616]" italics="true" pageId="4" pageNumber="287">Bactrocera</emphasis>
(
<emphasis id="B949EAC3ED29FF891862F9FDFB66F9B5" box="[1131,1256,1591,1616]" italics="true" pageId="4" pageNumber="287">Bactrocera</emphasis>
)
<emphasis id="B949EAC3ED29FF8918F0F9FCFA12F9B5" box="[1273,1436,1590,1616]" italics="true" pageId="4" pageNumber="287">paraosbeckiae</emphasis>
Drew
</taxonomicName>
in possessing reddish brown scutum (
<figureCitation id="13062A54ED29FF891E84F990FD56F991" box="[653,728,1626,1652]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Figs 2</figureCitation>
,
<figureCitation id="13062A54ED29FF891EEFF990FC8DF991" box="[742,771,1626,1652]" captionStart="FIGURES 1724" captionStartId="8.[151,261,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-24@8.[151,1436,181,1585]" captionTargetPageId="8" captionText="FIGURES 1724. Bactrocera (Bactrocera) paraosbeckiae Drew. 17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209983" httpUri="https://zenodo.org/record/13209983/files/figure.png" pageId="4" pageNumber="287">18</figureCitation>
) and elongate (
<quantity id="4CC59B34ED29FF891FBFF990FB91F991" box="[950,1055,1626,1652]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="287" unit="mm" value="0.27">0.27 mm</quantity>
) lateral surstylus (
<figureCitation id="13062A54ED29FF8918FDF990FAB1F991" box="[1268,1343,1626,1652]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Figs 6</figureCitation>
,
<figureCitation id="13062A54ED29FF891944F990FAE2F991" box="[1357,1388,1626,1652]" captionStart="FIGURES 1724" captionStartId="8.[151,261,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-24@8.[151,1436,181,1585]" captionTargetPageId="8" captionText="FIGURES 1724. Bactrocera (Bactrocera) paraosbeckiae Drew. 17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209983" httpUri="https://zenodo.org/record/13209983/files/figure.png" pageId="4" pageNumber="287">22</figureCitation>
,
<figureCitation id="13062A54ED29FF891970F990FF3AF97D" captionStart="FIGURES4144" captionStartId="12.[151,264,1204,1228]" captionTargetBox="[151,1436,181,1180]" captionTargetId="figure-24@12.[151,1436,181,1180]" captionTargetPageId="12" captionText="FIGURES4144. Comparisonbetween theepandrium(lateral view).41, Bactrocera(Bactrocera) ettinabhujaAbhishek &amp;David, sp. nov.; 42, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 43, Bactrocera (Bactrocera) paraosbeckiae Drew.; 44, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209993" httpUri="https://zenodo.org/record/13209993/files/figure.png" pageId="4" pageNumber="287">41, 43</figureCitation>
) but can be differentiated by black subscutellum (
<figureCitation id="13062A54ED29FF891F09F9B4FCD9F97C" box="[768,855,1662,1689]" captionStart="FIGURES 5356" captionStartId="15.[151,264,1215,1239]" captionTargetBox="[177,1411,181,1191]" captionTargetId="figure-17@15.[177,1411,181,1191]" captionTargetPageId="15" captionText="FIGURES 5356. Comparison between the subscutellum. 53, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 54, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 55, Bactrocera (Bactrocera) paraosbeckiae Drew; 56, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13210001" httpUri="https://zenodo.org/record/13210001/files/figure.png" pageId="4" pageNumber="287">Fig. 53</figureCitation>
); lateral vitta narrow (
<quantity id="4CC59B34ED29FF891857F9B4FB46F97D" box="[1118,1224,1662,1688]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="287" unit="mm" value="0.12">0.12 mm</quantity>
) obscured beyond postsutural supra-alar seta, ending at or before intra-alar setae (
<figureCitation id="13062A54ED29FF891F5FF968FC14F959" box="[854,922,1698,1724]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Fig. 2</figureCitation>
); narrow costal band confluent with R
<subScript id="17B93494ED29FF89194FF97FFAD1F926" attach="left" box="[1350,1375,1717,1731]" fontSize="6" pageId="4" pageNumber="287">2+3</subScript>
(
<figureCitation id="13062A54ED29FF891966F968FF29F904" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Fig. 5</figureCitation>
); ceromata reddish brown (
<quantity id="4CC59B34ED29FF891DD2F90CFDCFF904" box="[475,577,1734,1761]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="287" unit="mm" value="0.33">0.33 mm</quantity>
high,
<quantity id="4CC59B34ED29FF891E8EF90CFD65F904" box="[647,747,1734,1761]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="287" unit="mm" value="0.62">0.62 mm</quantity>
wide) (
<figureCitation id="13062A54ED29FF891F49F90CFC07F905" box="[832,905,1734,1760]" captionStart="FIGURES 18" captionStartId="5.[151,264,1534,1558]" captionTargetBox="[164,1423,166,1506]" captionTargetId="figure-17@5.[164,1423,166,1510]" captionTargetPageId="5" captionText="FIGURES 18. Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov. 1, head; 2, scutum (dorsal view); 3, thorax (lateral view); 4, abdomen; 5, wing; 6, epandrium and surstyli (lateral view); 7, epandrium and surstyli (posterior view); 8, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209975" httpUri="https://zenodo.org/record/13209975/files/figure.png" pageId="4" pageNumber="287">Figs 4</figureCitation>
,
<figureCitation id="13062A54ED29FF891F9CF90CFC3BF904" box="[917,949,1734,1761]" captionStart="FIGURES 5760" captionStartId="15.[151,264,1941,1965]" captionTargetBox="[170,1416,1335,1917]" captionTargetId="figure-58@15.[170,1416,1335,1917]" captionTargetPageId="15" captionText="FIGURES 5760. Comparison between the ceromata. 57, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 58, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 59, Bactrocera (Bactrocera) paraosbeckiae Drew; 60, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13210003" httpUri="https://zenodo.org/record/13210003/files/figure.png" pageId="4" pageNumber="287">57</figureCitation>
); abdominal sternites I, II and V fuscous, II and III fulvous and males are attracted to Methyl eugenol. In contrast,
<taxonomicName id="4C3D4D52ED29FF891FADF921FBEBF8E1" authorityName="Drew, Drew and Raghu" authorityYear="2002" box="[932,1125,1770,1796]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="paraosbeckiae">
<emphasis id="B949EAC3ED29FF891FADF921FBEBF8E1" box="[932,1125,1770,1796]" italics="true" pageId="4" pageNumber="287">B. paraosbeckiae</emphasis>
</taxonomicName>
possess black subscutellum with medial brown patch posteriorly (
<figureCitation id="13062A54ED29FF891E48F8C4FD1BF8CC" box="[577,661,1806,1833]" captionStart="FIGURES 5356" captionStartId="15.[151,264,1215,1239]" captionTargetBox="[177,1411,181,1191]" captionTargetId="figure-17@15.[177,1411,181,1191]" captionTargetPageId="15" captionText="FIGURES 5356. Comparison between the subscutellum. 53, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 54, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 55, Bactrocera (Bactrocera) paraosbeckiae Drew; 56, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13210001" httpUri="https://zenodo.org/record/13210001/files/figure.png" pageId="4" pageNumber="287">Fig. 56</figureCitation>
); lateral vitta tapering posteriorly (
<quantity id="4CC59B34ED29FF891814F8C4FB0AF8CD" box="[1053,1156,1806,1832]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="287" unit="mm" value="0.13">0.13 mm</quantity>
) ending at intra-alar seta (
<figureCitation id="13062A54ED29FF891C96F8F8FF7BF8A9" box="[159,245,1842,1868]" captionStart="FIGURES 1724" captionStartId="8.[151,261,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-24@8.[151,1436,181,1585]" captionTargetPageId="8" captionText="FIGURES 1724. Bactrocera (Bactrocera) paraosbeckiae Drew. 17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209983" httpUri="https://zenodo.org/record/13209983/files/figure.png" pageId="4" pageNumber="287">Fig. 18</figureCitation>
); costal band overlapping vein R
<subScript id="17B93494ED29FF891E79F88FFD07F8B6" attach="left" box="[624,649,1861,1875]" fontSize="6" pageId="4" pageNumber="287">2+3</subScript>
(
<figureCitation id="13062A54ED29FF891E9EF8F8FD62F8A9" box="[663,748,1842,1868]" captionStart="FIGURES 1724" captionStartId="8.[151,261,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-24@8.[151,1436,181,1585]" captionTargetPageId="8" captionText="FIGURES 1724. Bactrocera (Bactrocera) paraosbeckiae Drew. 17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209983" httpUri="https://zenodo.org/record/13209983/files/figure.png" pageId="4" pageNumber="287">Fig. 21</figureCitation>
); ceromata predominantly fuscous (
<quantity id="4CC59B34ED29FF89188CF8F8FB63F8A8" box="[1157,1261,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="287" unit="mm" value="0.37">0.37 mm</quantity>
high,
<quantity id="4CC59B34ED29FF89193EF8F8FA13F8A8" box="[1335,1437,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="287" unit="mm" value="0.56">0.56 mm</quantity>
wide) (
<figureCitation id="13062A54ED29FF891CEFF89CFEB0F895" box="[230,318,1878,1904]" captionStart="FIGURES 1724" captionStartId="8.[151,261,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-24@8.[151,1436,181,1585]" captionTargetPageId="8" captionText="FIGURES 1724. Bactrocera (Bactrocera) paraosbeckiae Drew. 17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209983" httpUri="https://zenodo.org/record/13209983/files/figure.png" pageId="4" pageNumber="287">Figs 20</figureCitation>
,
<figureCitation id="13062A54ED29FF891D40F89CFEE7F894" box="[329,361,1878,1905]" captionStart="FIGURES 5760" captionStartId="15.[151,264,1941,1965]" captionTargetBox="[170,1416,1335,1917]" captionTargetId="figure-58@15.[170,1416,1335,1917]" captionTargetPageId="15" captionText="FIGURES 5760. Comparison between the ceromata. 57, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 58, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 59, Bactrocera (Bactrocera) paraosbeckiae Drew; 60, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13210003" httpUri="https://zenodo.org/record/13210003/files/figure.png" pageId="4" pageNumber="287">59</figureCitation>
); abdominal sternites I, III, IV and V fuscous, II fulvous and males are attracted to cuelure.
</paragraph>
</subSubSection>
</treatment>
</document>