treatments-xml/data/03/C2/87/03C287F5FFA9FF80DFCDF952265AFD66.xml

<document id="3AD17BAFE1D9D8AA7B96B0C8A7BD32C0" ID-CLB-Dataset="32538" ID-DOI="10.11646/zootaxa.4294.3.9" ID-GBIF-Dataset="33e106cc-ce9e-4961-a008-618db6baa57f" ID-ISSN="1175-5326" ID-Zenodo-Dep="840186" ID-ZooBank="78C13075-B932-4424-AFAD-9547F7D013B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502206309044" checkinUser="plazi" docAuthor="Saldaitis, Aidas, Yakovlev, Roman V. &amp; Truuverk, Andro" docDate="2017" docId="03C287F5FFA9FF80DFCDF952265AFD66" docLanguage="en" docName="zootaxa.4294.3.9.pdf" docOrigin="Zootaxa 4294 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kerzhnerocossus tannuolus Saldaitis &amp; Yakovlev &amp; Truuverk 2017, sp. nov." docType="treatment" docUuid="0CC93CED-CBF3-4EA5-8F87-55EB40911857" docUuidSource="ZooBank" docVersion="8" lastPageNumber="393" masterDocId="FFFBFF8DFFA9FF84DF5AFFEC252EFFCE" masterDocTitle="Review of the genus Kerzhnerocossus Yakovlev, 2011 (Lepidoptera: Cossidae) with descriptions of two new species from Russia and Mongolia" masterLastPageNumber="394" masterPageNumber="389" pageNumber="389" updateTime="1698466344396" updateUser="ExternalLinkService">
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<mods:title id="64B471E9E98611D7AB19532893C11957">Review of the genus Kerzhnerocossus Yakovlev, 2011 (Lepidoptera: Cossidae) with descriptions of two new species from Russia and Mongolia</mods:title>
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<mods:namePart id="A2A2734FA80B8322DFBA94C8A9657260">Saldaitis, Aidas</mods:namePart>
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<paragraph id="8BD436E3FFA9FF84DFCDF9522764F916" blockId="0.[151,586,1726,1786]" box="[151,586,1726,1752]" pageId="0" pageNumber="389">
<heading id="D09C818FFFA9FF84DFCDF9522764F916" bold="true" box="[151,586,1726,1752]" fontSize="11" level="1" pageId="0" pageNumber="389" reason="1">
<emphasis id="B91FEAF1FFA9FF84DFCDF9522764F916" bold="true" box="[151,586,1726,1752]" pageId="0" pageNumber="389">
<taxonomicName id="4C6B4D60FFA9FF84DFCDF95224CBF916" authority="Saldaitis &amp; Yakovlev &amp; Truuverk, 2017" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[151,485,1726,1752]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="tannuolus" status="sp. nov.">
<emphasis id="B91FEAF1FFA9FF84DFCDF95224CBF916" bold="true" box="[151,485,1726,1752]" italics="true" pageId="0" pageNumber="389">Kerzhnerocossus tannuolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C578AFFA9FF84DEB6F9522764F916" box="[492,586,1726,1752]" pageId="0" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
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(
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, 
<figureCitation id="13502A66FFA9FF84DE71F90D2478F937" box="[299,342,1761,1786]" captionStart="FIGURES 6 – 9" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[159,1433,379,1884]" captionTargetId="figure@2.[151,1436,373,1894]" captionTargetPageId="2" captionText="FIGURES 6 – 9. Kerzhnerocossus ssp. male genitalia. 6. K. tannuolus, male, holotype, OP 3497 m (ASV / WIGJ); 7. K. tannuolus, male, paratype, OP 3459 m (OP); 8. K. kamelini, male, holotype (ZISP); 9. K. sambainu, male, paratype (ZISP)." httpUri="https://zenodo.org/record/840190/files/figure.png" pageId="0" pageNumber="389">6, 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3716568FFA9FF84DFCDF8C5211EF838" pageId="0" pageNumber="389" type="materials_examined">
<paragraph id="8BD436E3FFA9FF84DFCDF8C52111F844" blockId="0.[151,1437,1833,2038]" pageId="0" pageNumber="389">
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<emphasis id="B91FEAF1FFA9FF84DFCDF8C52411F88C" bold="true" box="[151,319,1833,1858]" pageId="0" pageNumber="389">
<typeStatus id="54D08841FFA9FF84DFCDF8C525FCF88C" box="[151,210,1833,1858]" pageId="0" pageNumber="389">Type</typeStatus>
material
</emphasis>
. 
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<typeStatus id="54D08841FFA9FF84DE17F8C52490F88C" box="[333,446,1833,1858]" pageId="0" pageNumber="389" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (
<figureCitation id="13502A66FFA9FF84DD49F8C5277BF88F" box="[531,597,1833,1858]" captionStart="FIGURES 1 – 5" captionStartId="1.[151,264,1590,1613]" captionTargetBox="[340,1247,446,1569]" captionTargetId="figure@1.[340,1247,445,1569]" captionTargetPageId="1" captionText="FIGURES 1 – 5. Kerzhnerocossus spp. adults &amp; distribution map. 1. K. tannuolus, male, holotype, [Russia], Tuva, OP 3497 m (ASV / WIGJ); 2. K. tannuolus, male, paratype, [Russia], Tuva, OP 3459 m (OP); 3. K. kamelini, male, holotype, Mongolia, Hovd aimak (ZISP); 4. K. sambainu, male, paratype, Mongolia, East aimak (ZISP); 5. Distribution map of the genus Kerzhnerocossus." httpUri="https://zenodo.org/record/840188/files/figure.png" pageId="0" pageNumber="389">Fig. 1</figureCitation>
), S[outhern] 
<collectorName id="269E5335FFA9FF84DDB4F8C5264EF88C" box="[750,864,1833,1858]" pageId="0" pageNumber="389">Siberia, C</collectorName>
[entral] 
<collectingRegion id="49AFF801FFA9FF84DCE0F8C626DBF88F" box="[954,1013,1834,1857]" country="Russia" name="Tyva" pageId="0" pageNumber="389">Tuva</collectingRegion>
, 
<collectingCounty id="62B54E6FFFA9FF84DB58F8C52012F88F" box="[1026,1340,1833,1858]" pageId="0" pageNumber="389">Vostochnyi Tannu-Ola Mts.</collectingCounty>
, 
<collectingMunicipality id="6BB0AC99FFA9FF84DA12F8C620B2F88C" box="[1352,1436,1834,1858]" pageId="0" pageNumber="389">Durgen</collectingMunicipality>
r[iver]., 
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vill. env., 
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<quantity id="4C939B06FFA9FF84DEAAF8A12764F8AB" box="[496,586,1869,1894]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="0" pageNumber="389" unit="m" value="1250.0">1250 m</quantity>
</elevation>
, 12̄ 
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<date id="FFD51023FFA9FF84DDDBF8A12629F8A8" box="[641,775,1869,1894]" pageId="0" pageNumber="389" value="2004-07-13">13.vii.2004</date>
</collectingDate>
, 
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/ 
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, leg. 
<collectorName id="269E5335FFA9FF84DB46F8A2215AF8AB" box="[1052,1140,1870,1893]" pageId="0" pageNumber="389">J. Hron</collectorName>
, M. 
<collectorName id="269E5335FFA9FF84DBF7F8A12022F8A8" box="[1197,1292,1869,1894]" pageId="0" pageNumber="389">Česanek</collectorName>
&amp; J. 
<collectorName id="269E5335FFA9FF84DA09F8A120B2F8A8" box="[1363,1436,1869,1894]" pageId="0" pageNumber="389">Rekelj</collectorName>
(slide OP 
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<quantity id="4C939B06FFA9FF84DE5BF89D247FF847" box="[257,337,1905,1929]" metricMagnitude="3" metricUnit="m" metricValue="3.497" pageId="0" pageNumber="389" unit="m" value="3497.0">3497m</quantity>
</elevation>
, 
<collectorName id="269E5335FFA9FF84DE06F89D24E8F844" box="[348,454,1905,1930]" pageId="0" pageNumber="389">GenBank</collectorName>
accession code: 
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MF 
<date id="FFD51023FFA9FF84DDF1F89D262EF844" box="[683,768,1905,1930]" pageId="0" pageNumber="389" value="56-07-14">071456</date>
</accessionNumber>
) (
<figureCitation id="13502A66FFA9FF84DC42F89D2672F844" box="[792,860,1905,1930]" captionStart="FIGURES 6 – 9" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[159,1433,379,1884]" captionTargetId="figure@2.[151,1436,373,1894]" captionTargetPageId="2" captionText="FIGURES 6 – 9. Kerzhnerocossus ssp. male genitalia. 6. K. tannuolus, male, holotype, OP 3497 m (ASV / WIGJ); 7. K. tannuolus, male, paratype, OP 3459 m (OP); 8. K. kamelini, male, holotype (ZISP); 9. K. sambainu, male, paratype (ZISP)." httpUri="https://zenodo.org/record/840190/files/figure.png" pageId="0" pageNumber="389">
<collectorName id="269E5335FFA9FF84DC42F89D266BF844" box="[792,837,1905,1930]" pageId="0" pageNumber="389">Fig.</collectorName>
6
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), (coll.ASV/ 
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).
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</paragraph>
<paragraph id="8BD436E3FFA9FF84DF9DF879211EF838" blockId="0.[151,1437,1833,2038]" pageId="0" pageNumber="389">
<materialsCitation id="3B033CBEFFA9FF84DF9DF8792689F81C" ID-GBIF-Occurrence="1585957401" collectingDate="2002-07-21" collectorName="M. Cesanek" elevation="3459" location="GenBank" pageId="0" pageNumber="389" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54D08841FFA9FF84DF9DF8792411F860" box="[199,319,1941,1966]" pageId="0" pageNumber="389" type="paratype">
<emphasis id="B91FEAF1FFA9FF84DF9DF8792411F860" bold="true" box="[199,319,1941,1966]" pageId="0" pageNumber="389">Paratypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D6DFD6AFFA9FF84DE17F87A24B9F860" box="[333,407,1941,1966]" pageId="0" pageNumber="389" type="male">1 male</specimenCount>
(
<figureCitation id="13502A66FFA9FF84DEFCF87924C4F860" box="[422,490,1941,1966]" captionStart="FIGURES 1 – 5" captionStartId="1.[151,264,1590,1613]" captionTargetBox="[340,1247,446,1569]" captionTargetId="figure@1.[340,1247,445,1569]" captionTargetPageId="1" captionText="FIGURES 1 – 5. Kerzhnerocossus spp. adults &amp; distribution map. 1. K. tannuolus, male, holotype, [Russia], Tuva, OP 3497 m (ASV / WIGJ); 2. K. tannuolus, male, paratype, [Russia], Tuva, OP 3459 m (OP); 3. K. kamelini, male, holotype, Mongolia, Hovd aimak (ZISP); 4. K. sambainu, male, paratype, Mongolia, East aimak (ZISP); 5. Distribution map of the genus Kerzhnerocossus." httpUri="https://zenodo.org/record/840188/files/figure.png" pageId="0" pageNumber="389">Fig. 2</figureCitation>
), with the same data of locality as the 
<typeStatus id="54D08841FFA9FF84DCCEF87926D8F860" box="[916,1014,1941,1966]" pageId="0" pageNumber="389" type="holotype">holotype</typeStatus>
(slide OP 
<elevation id="0046D1D0FFA9FF84DB3CF8792198F860" box="[1126,1206,1941,1966]" metricMagnitude="3" metricUnit="m" metricValue="3.4589999999999996" pageId="0" pageNumber="389" unit="m" value="3459.0">
<quantity id="4C939B06FFA9FF84DB3CF8792198F860" box="[1126,1206,1941,1966]" metricMagnitude="3" metricUnit="m" metricValue="3.4589999999999996" pageId="0" pageNumber="389" unit="m" value="3459.0">3459m</quantity>
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, 
<location id="8EB46038FFA9FF84DB9BF8792005F860" LSID="urn:lsid:plazi:treatment:03C287F5FFA9FF80DFCDF952265AFD66:8EB46038FFA9FF84DB9BF8792005F860" box="[1217,1323,1941,1966]" name="GenBank" pageId="0" pageNumber="389">GenBank</location>
accession code: 
<accessionNumber id="9438AB00FFA9FF84DF87F8552472F81C" box="[221,348,1977,2002]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF071457" pageId="0" pageNumber="389">
MF 
<date id="FFD51023FFA9FF84DE5DF8552472F81C" box="[263,348,1977,2002]" pageId="0" pageNumber="389" value="57-07-14">071457</date>
</accessionNumber>
) (
<figureCitation id="13502A66FFA9FF84DE2FF8552493F81F" box="[373,445,1977,2002]" captionStart="FIGURES 6 – 9" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[159,1433,379,1884]" captionTargetId="figure@2.[151,1436,373,1894]" captionTargetPageId="2" captionText="FIGURES 6 – 9. Kerzhnerocossus ssp. male genitalia. 6. K. tannuolus, male, holotype, OP 3497 m (ASV / WIGJ); 7. K. tannuolus, male, paratype, OP 3459 m (OP); 8. K. kamelini, male, holotype (ZISP); 9. K. sambainu, male, paratype (ZISP)." httpUri="https://zenodo.org/record/840190/files/figure.png" pageId="0" pageNumber="389">Fig. 7</figureCitation>
), (coll. OP), 
<collectingDate id="EF91E9CBFFA9FF84DD0DF85527F4F81C" box="[599,730,1977,2002]" pageId="0" pageNumber="389" value="2002-07-21">
<date id="FFD51023FFA9FF84DD0DF85527F4F81C" box="[599,730,1977,2002]" pageId="0" pageNumber="389" value="2002-07-21">21.vii.2002</date>
</collectingDate>
, leg. 
<collectorName id="269E5335FFA9FF84DC41F8562689F81C" box="[795,935,1977,2002]" pageId="0" pageNumber="389">M. Česanek</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B033CBEFFA9FF84DCECF855211EF838" ID-GBIF-Occurrence="1585957400" collectingDate="1995-06-12" collectionCode="MZH" collectorName="Jalava &amp; Kullberg" country="Russia" latitude="50.016666" location="Tere-Khol lake" longLatPrecision="1099" longitude="95.05" pageId="0" pageNumber="389" specimenCount="1" specimenCount-male="1" stateProvince="Tuva" typeStatus="paratype">
<specimenCount id="9D6DFD6AFFA9FF84DCECF8552129F81C" box="[950,1031,1977,2002]" pageId="0" pageNumber="389" type="male">1 male</specimenCount>
, 
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, 
<collectingRegion id="49AFF801FFA9FF84DB2BF8562184F81F" box="[1137,1194,1978,2001]" country="Russia" name="Tyva" pageId="0" pageNumber="389">Tuva</collectingRegion>
Republic, 
<location id="8EB46038FFA9FF84DA70F85525E7F838" LSID="urn:lsid:plazi:treatment:03C287F5FFA9FF80DFCDF952265AFD66:8EB46038FFA9FF84DA70F85525E7F838" country="Russia" latitude="50.016666" longLatPrecision="1099" longitude="95.05" name="Tere-Khol lake" pageId="0" pageNumber="389" stateProvince="Tuva">Tere-Khol lake</location>
, sand dunes, 9̄ 
<collectingDate id="EF91E9CBFFA9FF84DE2DF83124DAF838" box="[375,500,2013,2038]" pageId="0" pageNumber="389" value="1995-06-12">
<date id="FFD51023FFA9FF84DE2DF83124DAF838" box="[375,500,2013,2038]" pageId="0" pageNumber="389" value="1995-06-12">12.vi.1995</date>
</collectingDate>
, 
<geoCoordinate id="EE5F5024FFA9FF84DD5BF831274FF838" box="[513,609,2013,2038]" direction="north" orientation="latitude" pageId="0" pageNumber="389" precision="925" value="50.016666">50°01´N</geoCoordinate>
<geoCoordinate id="EE5F5024FFA9FF84DD32F83127E6F838" box="[616,712,2013,2038]" direction="east" orientation="longitude" pageId="0" pageNumber="389" precision="925" value="95.05">95°03´E</geoCoordinate>
, 
<collectorName id="269E5335FFA9FF84DD8EF8312634F838" box="[724,794,2013,2038]" pageId="0" pageNumber="389">Jalava</collectorName>
&amp; 
<collectorName id="269E5335FFA9FF84DC64F831268FF838" box="[830,929,2013,2038]" pageId="0" pageNumber="389">Kullberg</collectorName>
leg. (
<collectionCode id="ED7AAE26FFA9FF84DCBAF832210AF838" box="[992,1060,2014,2038]" country="Finland" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="0" pageNumber="389">MZH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716568FFA8FF85DF9DFF7B2681FE46" pageId="1" pageNumber="390" type="diagnosis">
<paragraph id="8BD436E3FFA8FF85DF9DFF7B2681FE46" blockId="1.[151,1437,151,392]" pageId="1" pageNumber="390">
<emphasis id="B91FEAF1FFA8FF85DF9DFF7B246FFF7E" bold="true" box="[199,321,151,176]" pageId="1" pageNumber="390">Diagnosis.</emphasis>
<taxonomicName id="4C6B4D60FFA8FF85DE10FF742751FF7E" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[330,639,152,176]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFA8FF85DE10FF742751FF7E" box="[330,639,152,176]" italics="true" pageId="1" pageNumber="390">Kerzhnerocossus tannuolus</emphasis>
</taxonomicName>
differs from other members of the genus by its larger size (wingspan 28̄ 
<quantity id="4C939B06FFA8FF85DF9AFF50243BFF1A" box="[192,277,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="390" unit="mm" value="30.0">30 mm</quantity>
vs 24̄ 
<quantity id="4C939B06FFA8FF85DE31FF5024ECFF1A" box="[363,450,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="390" unit="mm" value="25.0">25 mm</quantity>
), different forewing pattern and undulated hindwing bands rather than the reticulated pattern as in 
<taxonomicName id="4C6B4D60FFA8FF85DE75FF0D24EEFF36" authorityName="Yakovlev" authorityYear="2011" box="[303,448,224,248]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="sambainu">
<emphasis id="B91FEAF1FFA8FF85DE75FF0D24EEFF36" box="[303,448,224,248]" italics="true" pageId="1" pageNumber="390">K. sambainu</emphasis>
</taxonomicName>
(
<figureCitation id="13502A66FFA8FF85DE88FF332736FF36" box="[466,536,223,248]" captionStart="FIGURES 1 – 5" captionStartId="1.[151,264,1590,1613]" captionTargetBox="[340,1247,446,1569]" captionTargetId="figure@1.[340,1247,445,1569]" captionTargetPageId="1" captionText="FIGURES 1 – 5. Kerzhnerocossus spp. adults &amp; distribution map. 1. K. tannuolus, male, holotype, [Russia], Tuva, OP 3497 m (ASV / WIGJ); 2. K. tannuolus, male, paratype, [Russia], Tuva, OP 3459 m (OP); 3. K. kamelini, male, holotype, Mongolia, Hovd aimak (ZISP); 4. K. sambainu, male, paratype, Mongolia, East aimak (ZISP); 5. Distribution map of the genus Kerzhnerocossus." httpUri="https://zenodo.org/record/840188/files/figure.png" pageId="1" pageNumber="390">Fig. 4</figureCitation>
) and 
<taxonomicName id="4C6B4D60FFA8FF85DD06FF0D27CEFF36" authority="Saldaitis &amp; Yakovlev &amp; Truuverk, 2017" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[604,736,224,248]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="kamelini" status="sp. nov.">
<emphasis id="B91FEAF1FFA8FF85DD06FF0D27CEFF36" box="[604,736,224,248]" italics="true" pageId="1" pageNumber="390">K. kamelini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C578AFFA8FF85DDB0FF332668FF36" box="[746,838,223,248]" pageId="1" pageNumber="390" rank="species">
<emphasis id="B91FEAF1FFA8FF85DDB0FF332668FF36" bold="true" box="[746,838,223,248]" pageId="1" pageNumber="390">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13502A66FFA8FF85DC02FF3326B1FF36" box="[856,927,223,248]" captionStart="FIGURES 1 – 5" captionStartId="1.[151,264,1590,1613]" captionTargetBox="[340,1247,446,1569]" captionTargetId="figure@1.[340,1247,445,1569]" captionTargetPageId="1" captionText="FIGURES 1 – 5. Kerzhnerocossus spp. adults &amp; distribution map. 1. K. tannuolus, male, holotype, [Russia], Tuva, OP 3497 m (ASV / WIGJ); 2. K. tannuolus, male, paratype, [Russia], Tuva, OP 3459 m (OP); 3. K. kamelini, male, holotype, Mongolia, Hovd aimak (ZISP); 4. K. sambainu, male, paratype, Mongolia, East aimak (ZISP); 5. Distribution map of the genus Kerzhnerocossus." httpUri="https://zenodo.org/record/840188/files/figure.png" pageId="1" pageNumber="390">Fig. 3</figureCitation>
). In its male genitalia the valva has a small semicircular process in the center of the costal edge. In 
<taxonomicName id="4C6B4D60FFA8FF85DC76FEE9269EFED2" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[812,944,260,284]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="kamelini">
<emphasis id="B91FEAF1FFA8FF85DC76FEE9269EFED2" box="[812,944,260,284]" italics="true" pageId="1" pageNumber="390">K. kamelini</emphasis>
</taxonomicName>
(
<figureCitation id="13502A66FFA8FF85DC9EFEE82124FED2" box="[964,1034,260,285]" captionStart="FIGURES 6 – 9" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[159,1433,379,1884]" captionTargetId="figure@2.[151,1436,373,1894]" captionTargetPageId="2" captionText="FIGURES 6 – 9. Kerzhnerocossus ssp. male genitalia. 6. K. tannuolus, male, holotype, OP 3497 m (ASV / WIGJ); 7. K. tannuolus, male, paratype, OP 3459 m (OP); 8. K. kamelini, male, holotype (ZISP); 9. K. sambainu, male, paratype (ZISP)." httpUri="https://zenodo.org/record/840190/files/figure.png" pageId="1" pageNumber="390">Fig. 8</figureCitation>
) the process on the costal edge is trapezoidal, the transtilla process is triangular, the angle of the transtilla process apex is about 80° (in 
<taxonomicName id="4C6B4D60FFA8FF85DA54FEC520B2FE8E" authorityName="Yakovlev" authorityYear="2011" box="[1294,1436,296,320]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="sambainu">
<emphasis id="B91FEAF1FFA8FF85DA54FEC520B2FE8E" box="[1294,1436,296,320]" italics="true" pageId="1" pageNumber="390">K. sambainu</emphasis>
</taxonomicName>
(
<figureCitation id="13502A66FFA8FF85DFC5FEA025CCFEAA" box="[159,226,332,357]" captionStart="FIGURES 6 – 9" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[159,1433,379,1884]" captionTargetId="figure@2.[151,1436,373,1894]" captionTargetPageId="2" captionText="FIGURES 6 – 9. Kerzhnerocossus ssp. male genitalia. 6. K. tannuolus, male, holotype, OP 3497 m (ASV / WIGJ); 7. K. tannuolus, male, paratype, OP 3459 m (OP); 8. K. kamelini, male, holotype (ZISP); 9. K. sambainu, male, paratype (ZISP)." httpUri="https://zenodo.org/record/840190/files/figure.png" pageId="1" pageNumber="390">Fig. 9</figureCitation>
) the angle is about 60°) and the phallus is slightly curved from the base through the middle while in both 
<taxonomicName id="4C6B4D60FFA8FF85DAD8FEA1242AFE46" authorityName="Yakovlev" authorityYear="2011" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="sambainu">
<emphasis id="B91FEAF1FFA8FF85DAD8FEA1242AFE46" italics="true" pageId="1" pageNumber="390">K. sambainu</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6B4D60FFA8FF85DE60FE9D2495FE46" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[314,443,368,392]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="kamelini">
<emphasis id="B91FEAF1FFA8FF85DE60FE9D2495FE46" box="[314,443,368,392]" italics="true" pageId="1" pageNumber="390">K. kamelini</emphasis>
</taxonomicName>
the phallus is strongly curved in the middle.
</paragraph>
</subSubSection>
<caption id="DF14666BFFA8FF85DFCDF9DA2467F967" httpUri="https://zenodo.org/record/840188/files/figure.png" pageId="1" pageNumber="390" targetBox="[340,1247,446,1569]" targetPageId="1">
<paragraph id="8BD436E3FFA8FF85DFCDF9DA2467F967" blockId="1.[151,1436,1590,1705]" pageId="1" pageNumber="390">
<emphasis id="B91FEAF1FFA8FF85DFCDF9DA2415F982" bold="true" box="[151,315,1590,1613]" pageId="1" pageNumber="390">FIGURES 1–5.</emphasis>
<taxonomicName id="4C6B4D60FFA8FF85DE19F9DB24C1F983" authorityName="Yakovlev" authorityYear="2011" box="[323,495,1591,1613]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAF1FFA8FF85DE19F9DB24C1F983" box="[323,495,1591,1613]" italics="true" pageId="1" pageNumber="390">Kerzhnerocossus</emphasis>
</taxonomicName>
spp. adults &amp; distribution map. 1. 
<taxonomicName id="4C6B4D60FFA8FF85DC00F9D426F3F983" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[858,989,1591,1613]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFA8FF85DC00F9D426F3F983" box="[858,989,1591,1613]" italics="true" pageId="1" pageNumber="390">K. tannuolus</emphasis>
</taxonomicName>
, male, holotype, [Russia], Tuva, OP3497m (ASV/WIGJ); 2. 
<taxonomicName id="4C6B4D60FFA8FF85DE19F9BB24EAF9A5" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[323,452,1621,1644]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFA8FF85DE19F9BB24EAF9A5" box="[323,452,1621,1644]" italics="true" pageId="1" pageNumber="390">K. tannuolus</emphasis>
</taxonomicName>
, male, paratype, [Russia], Tuva, OP3459m (OP); 3. 
<taxonomicName id="4C6B4D60FFA8FF85DC89F9BB2169F9A5" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[979,1095,1621,1644]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="kamelini">
<emphasis id="B91FEAF1FFA8FF85DC89F9BB2169F9A5" box="[979,1095,1621,1644]" italics="true" pageId="1" pageNumber="390">K. kamelini</emphasis>
</taxonomicName>
, male, holotype, Mongolia, Hovd aimak (ZISP); 4. 
<taxonomicName id="4C6B4D60FFA8FF85DE3CF99924DEF944" authorityName="Yakovlev" authorityYear="2011" box="[358,496,1652,1674]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="sambainu">
<emphasis id="B91FEAF1FFA8FF85DE3CF99924DEF944" box="[358,496,1652,1674]" italics="true" pageId="1" pageNumber="390">K. sambainu</emphasis>
</taxonomicName>
, male, paratype, Mongolia, East aimak (ZISP); 5. Distribution map of the genus 
<taxonomicName id="4C6B4D60FFA8FF85DFCDF97F246DF967" authorityName="Yakovlev" authorityYear="2011" box="[151,323,1683,1705]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAF1FFA8FF85DFCDF97F246DF967" box="[151,323,1683,1705]" italics="true" pageId="1" pageNumber="390">Kerzhnerocossus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3716568FFA8FF86DF9DF93924BEFE8E" lastPageId="2" lastPageNumber="391" pageId="1" pageNumber="390" type="description">
<paragraph id="8BD436E3FFA8FF85DF9DF93926AAF86C" blockId="1.[151,1437,1749,2026]" pageId="1" pageNumber="390">
<emphasis id="B91FEAF1FFA8FF85DF9DF9392477F920" bold="true" box="[199,345,1749,1774]" pageId="1" pageNumber="390">Description.</emphasis>
Wingspan 28̄ 
<quantity id="4C939B06FFA8FF85DEA5F93A277FF920" box="[511,593,1750,1774]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="390" unit="mm" value="30.0">30 mm</quantity>
(
<typeStatus id="54D08841FFA8FF85DD05F93927EDF920" box="[607,707,1749,1774]" pageId="1" pageNumber="390" type="holotype">holotype</typeStatus>
<quantity id="4C939B06FFA8FF85DD90F93A2633F920" box="[714,797,1750,1774]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="390" unit="mm" value="30.0">30 mm</quantity>
), length of forewing 13̄ 
<quantity id="4C939B06FFA8FF85DB6AF93A21ACF920" box="[1072,1154,1750,1774]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="390" unit="mm" value="14.0">14 mm</quantity>
(
<typeStatus id="54D08841FFA8FF85DBCAF93921DAF920" box="[1168,1268,1749,1774]" pageId="1" pageNumber="390" type="holotype">holotype</typeStatus>
<quantity id="4C939B06FFA8FF85DBA1F93A2060F920" box="[1275,1358,1750,1774]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="390" unit="mm" value="14.0">14 mm</quantity>
). Head very small, width 0.2 times width of thorax; antenna bipectinate, length of processes exceeding rod diameter by 1.5 times; body densely covered with pale grey scales; forewing cream, with thick reticulated pattern of brown undulated bands and wide transverse bands in the postdiscal and submarginal areas with a series of transverse brown strokes along costal edge, apex rounded; fringe mottled, dark at veins and pale between veins; hindwing cream, with thin reticulated pattern and sputtering of brown scales.
</paragraph>
<paragraph id="8BD436E3FFA8FF86DF9DF8412711FED3" blockId="1.[151,1437,1749,2026]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" lastPageNumber="391" pageId="1" pageNumber="390">
Male genitalia (
<figureCitation id="13502A66FFA8FF85DE22F8412490F808" box="[376,446,1965,1990]" captionStart="FIGURES 6 – 9" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[159,1433,379,1884]" captionTargetId="figure@2.[151,1436,373,1894]" captionTargetPageId="2" captionText="FIGURES 6 – 9. Kerzhnerocossus ssp. male genitalia. 6. K. tannuolus, male, holotype, OP 3497 m (ASV / WIGJ); 7. K. tannuolus, male, paratype, OP 3459 m (OP); 8. K. kamelini, male, holotype (ZISP); 9. K. sambainu, male, paratype (ZISP)." httpUri="https://zenodo.org/record/840190/files/figure.png" pageId="1" pageNumber="390">Fig. 6</figureCitation>
). Uncus triangular, smoothly narrowing to apex, apex blunt, semicircular; gnathos arms thin, long, in fusion form small gnathos with fine spikes on surface; valve relatively narrow with small semicircular process in middle third of costal edge; distal third of valva membranous, transtilla process triangular, angle of transtilla process apex about 80°; juxta small, trapezoidal; saccus small, semicircular; phallus shorter than valve, spoon-like, slightly curved on border of basal and middle thirds; vesica aperture in dorso-apical position half length of the phallus; vesica without cornuti.
</paragraph>
<paragraph id="8BD436E3FFABFF86DF9DFECB24BEFE8E" blockId="2.[151,1436,151,320]" box="[199,400,295,320]" pageId="2" pageNumber="391">Female unknown.</paragraph>
</subSubSection>
<caption id="DF14666BFFABFF86DFCDF8972045F87E" httpUri="https://zenodo.org/record/840190/files/figure.png" pageId="2" pageNumber="391" targetBox="[159,1433,379,1884]" targetPageId="2">
<paragraph id="8BD436E3FFABFF86DFCDF8972045F87E" blockId="2.[151,1437,1915,1968]" pageId="2" pageNumber="391">
<emphasis id="B91FEAF1FFABFF86DFCDF897246FF85F" bold="true" box="[151,321,1915,1938]" pageId="2" pageNumber="391">FIGURES 6–9.</emphasis>
<taxonomicName id="4C6B4D60FFABFF86DE15F89024D5F85C" authorityName="Yakovlev" authorityYear="2011" box="[335,507,1916,1938]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAF1FFABFF86DE15F89024D5F85C" box="[335,507,1916,1938]" italics="true" pageId="2" pageNumber="391">Kerzhnerocossus</emphasis>
</taxonomicName>
ssp. male genitalia. 6. 
<taxonomicName id="4C6B4D60FFABFF86DC5CF89126A1F85C" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[774,911,1916,1938]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFABFF86DC5CF89126A1F85C" box="[774,911,1916,1938]" italics="true" pageId="2" pageNumber="391">K. tannuolus</emphasis>
</taxonomicName>
, male, holotype, OP3497m (ASV/WIGJ); 7. 
<taxonomicName id="4C6B4D60FFABFF86DADEF89125D5F87E" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFABFF86DADEF89125D5F87E" italics="true" pageId="2" pageNumber="391">K. tannuolus</emphasis>
</taxonomicName>
, male, paratype, OP3459m (OP); 8. 
<taxonomicName id="4C6B4D60FFABFF86DD34F87727CCF87E" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[622,738,1946,1968]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="kamelini">
<emphasis id="B91FEAF1FFABFF86DD34F87727CCF87E" box="[622,738,1946,1968]" italics="true" pageId="2" pageNumber="391">K. kamelini</emphasis>
</taxonomicName>
, male, holotype (ZISP); 9. 
<taxonomicName id="4C6B4D60FFABFF86DCADF8772159F87E" authorityName="Yakovlev" authorityYear="2011" box="[1015,1143,1946,1968]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="sambainu">
<emphasis id="B91FEAF1FFABFF86DCADF8772159F87E" box="[1015,1143,1946,1968]" italics="true" pageId="2" pageNumber="391">K. sambainu</emphasis>
</taxonomicName>
, male, paratype (ZISP).
</paragraph>
</caption>
<subSubSection id="C3716568FFAAFF80DF9DFF7B21D7FE1E" lastPageId="4" lastPageNumber="393" pageId="3" pageNumber="392" type="discussion">
<paragraph id="8BD436E3FFAAFF87DF9DFF7B245FFED3" blockId="3.[151,1437,151,285]" pageId="3" pageNumber="392">
<emphasis id="B91FEAF1FFAAFF87DF9DFF7B2415FF7E" bold="true" box="[199,315,151,176]" pageId="3" pageNumber="392">Remarks.</emphasis>
One of the paratype specimens has a double uncus (
<figureCitation id="13502A66FFAAFF87DCDBFF7B26EBFF7E" box="[897,965,151,176]" captionStart="FIGURES 6 – 9" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[159,1433,379,1884]" captionTargetId="figure@2.[151,1436,373,1894]" captionTargetPageId="2" captionText="FIGURES 6 – 9. Kerzhnerocossus ssp. male genitalia. 6. K. tannuolus, male, holotype, OP 3497 m (ASV / WIGJ); 7. K. tannuolus, male, paratype, OP 3459 m (OP); 8. K. kamelini, male, holotype (ZISP); 9. K. sambainu, male, paratype (ZISP)." httpUri="https://zenodo.org/record/840190/files/figure.png" pageId="3" pageNumber="392">Fig. 7</figureCitation>
). Despite this unique aberration, the DNA results confirm that the specimen is conspecific with the holotype. There are slight differences in the appearance of the specimens. Paratype from Tere-Khol lake, sand dunes is more light with more expressed postdiscal and submarginal bands.
</paragraph>
<caption id="DF14666BFFAAFF87DFCDF9CF2451F9B2" httpUri="https://zenodo.org/record/840192/files/figure.png" pageId="3" pageNumber="392" targetBox="[340,1247,429,1550]" targetPageId="3">
<paragraph id="8BD436E3FFAAFF87DFCDF9CF2451F9B2" blockId="3.[151,1436,1571,1660]" pageId="3" pageNumber="392">
<emphasis id="B91FEAF1FFAAFF87DFCDF9CF2478F9F7" bold="true" box="[151,342,1571,1594]" pageId="3" pageNumber="392">FIGURES 10–11.</emphasis>
<taxonomicName id="4C6B4D60FFAAFF87DE3AF9C924C8F9F4" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[352,486,1572,1594]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFAAFF87DE3AF9C924C8F9F4" box="[352,486,1572,1594]" italics="true" pageId="3" pageNumber="392">K. tannuolus</emphasis>
</taxonomicName>
biotope. 10. Russia, Tuva, Vostochnyi Tannu-Ola Mts., Durgen river valley general view (photo Martin Česanek, 1997); 11. Russia, Tuva, Vostochnyi Tannu-Ola Mts., Durgen river valley more detailed view (photo Martin Česanek, 1997)
</paragraph>
</caption>
<paragraph id="8BD436E3FFAAFF87DF9DF9402053F8FE" blockId="3.[151,1436,1708,2021]" pageId="3" pageNumber="392">
<emphasis id="B91FEAF1FFAAFF87DF9DF9402480F90B" bold="true" box="[199,430,1708,1733]" pageId="3" pageNumber="392">Molecular analysis.</emphasis>
DNA was extracted from the legs of two 
<taxonomicName id="4C6B4D60FFAAFF87DCDAF9412121F90A" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[896,1039,1708,1732]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFAAFF87DCDAF9412121F90A" box="[896,1039,1708,1732]" italics="true" pageId="3" pageNumber="392">K. tannuolus</emphasis>
</taxonomicName>
specimens, using a High Pure PCR Template Preparation Kit' (Roche Diagnostics GmbH, Mannheim, Germany). The extraction was carried out following the manufacturer's instructions, with the exception that the first incubation step was 55°C for up to 3 h rather than 1 h. Extracted genomic DNA is housed in the Museum of Natural History of the University of Tartu.
</paragraph>
<paragraph id="8BD436E3FFAAFF80DF9DF8D020BAFF1B" blockId="3.[151,1436,1708,2021]" lastBlockId="4.[151,1437,151,680]" lastPageId="4" lastPageNumber="393" pageId="3" pageNumber="392">
Reaction mixtures for PCR and cycle sequencing followed protocol described by 
<bibRefCitation id="EFFA4B12FFAAFF87DB02F8D0201BF89B" box="[1112,1333,1852,1877]" pageId="3" pageNumber="392" refString="Ounap, E., Viidalepp, J. &amp; Truuverk, A. (2016) Phylogeny of the subfamily Larentiinae (Lepidoptera: Geometridae): integrating molecular data and traditional classifications. Systematic Entomology, 41 (4), 824 - 843. https: // doi. org / 10.1111 / syen. 12195" type="journal article">
Õunap 
<emphasis id="B91FEAF1FFAAFF87DBF3F8D121F7F89A" box="[1193,1241,1852,1876]" italics="true" pageId="3" pageNumber="392">et al</emphasis>
. (2016)
</bibRefCitation>
, with the number of cycles being 45 for PCR, and temperature of the annealing step being 47C for PCR and 45C for cycle sequencing. Sequencing the whole barcode fragment of the COI gene using one primer pair failed. Therefore this gene region was sequenced in two partially overlapping fragments, using primers cov1f (5’-TCG CTT ATT ATT CAG CCA TTT TAT T-3’, 
<bibRefCitation id="EFFA4B12FFAAFF87DE8CF8202781F82B" author="Ounap" box="[470,687,1996,2021]" pageId="3" pageNumber="392" refString="Ounap, E., Viidalepp, J. &amp; Saarma, U. (2008) Systematic position of Lythriini revised: transferred from Larentiinae to Sterrhinae (Lepidoptera, Geometridae). Zoologica Scripta, 37, 405 - 413." type="journal article" year="2008">
Õunap 
<emphasis id="B91FEAF1FFAAFF87DD70F821274BF82A" box="[554,613,1996,2020]" italics="true" pageId="3" pageNumber="392">et al.</emphasis>
, 2008
</bibRefCitation>
) and cov1r (5’-CTG CAC CAT TTT CTA CAA TTC TTC T-3’, 
<bibRefCitation id="EFFA4B12FFADFF80DFCDFF7B245CFF7E" author="Ounap" box="[151,370,151,176]" pageId="4" pageNumber="393" refString="Ounap, E., Viidalepp, J. &amp; Saarma, U. (2008) Systematic position of Lythriini revised: transferred from Larentiinae to Sterrhinae (Lepidoptera, Geometridae). Zoologica Scripta, 37, 405 - 413." type="journal article" year="2008">
Õunap 
<emphasis id="B91FEAF1FFADFF80DFB6FF752409FF7E" box="[236,295,152,176]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
, 2008
</bibRefCitation>
) for 5’ half, and primers ron (5’-GGA TCA CCT GAT ATA GCA TTC CC-3’, 
<bibRefCitation id="EFFA4B12FFADFF80DA46FF7B2410FF1A" author="Caterino" pageId="4" pageNumber="393" refString="Caterino, M. S. &amp; Sperling, F. A. H. (1999) Papilio phylogeny based on mitochondrial cytochrome oxidase I and II genes. Molecular Phylogenetics and Evolution, 11, 122 - 137." type="journal article" year="1999">Caterino &amp; Sperling, 1999</bibRefCitation>
) and nancy (5’-CCC GGT AAA ATT AAA ATA TAA ACT TC-3’, 
<bibRefCitation id="EFFA4B12FFADFF80DB61FF502023FF1B" author="Ounap" box="[1083,1293,188,213]" pageId="4" pageNumber="393" refString="Ounap, E., Viidalepp, J. &amp; Saarma, U. (2005) Phylogenetic evaluation of the taxonomic status of Timandra griseata and T. comae (Lepidoptera: Geometridae: Sterrhinae). European Journal of Entomology, 102, 607 - 615. https: // doi. org / 10.14411 / eje. 2005.085" type="journal article" year="2005">
Õunap 
<emphasis id="B91FEAF1FFADFF80DBD7FF5121EBFF1A" box="[1165,1221,188,212]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
, 2005
</bibRefCitation>
) for 3’ half.
</paragraph>
<paragraph id="8BD436E3FFADFF80DF9DFF0C21B8FEAB" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="393">
Consensus sequences were created in GENEIOUS R7.1.7 (http://www.geneious.com, 
<bibRefCitation id="EFFA4B12FFADFF80DB24FF0C2061FF36" author="Kearse" box="[1150,1359,224,248]" pageId="4" pageNumber="393" refString="Kearse, M., Moir, R., Wilson, A., Stones-Havas, S., Cheung, M., Sturrock, S., Buxton, S., Cooper, A., Markowitz, S., Duran, C., Thierer, T., Ashton, B., Mentjies, P. &amp; Drummond, A. (2012) Geneious Basic: an integrated and extendable desktop software platform for the organization and analysis of sequence data. Bioinformatics, 28 (12), 1647 - 1649. https: // doi. org / 10.1093 / bioinformatics / bts 199" type="journal article" year="2012">
Kearse 
<emphasis id="B91FEAF1FFADFF80DB8EFF0D2028FF36" box="[1236,1286,224,248]" italics="true" pageId="4" pageNumber="393">et al</emphasis>
. 2012
</bibRefCitation>
) using sequence data from both DNA strands. Sequences were aligned by eye and edited in BIOEDIT 7.2.5 (
<bibRefCitation id="EFFA4B12FFADFF80DA43FEE820BEFED2" author="Hall" box="[1305,1424,260,285]" pageId="4" pageNumber="393" refString="Hall, T. A. (1999) BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95 / 98 / NT. Nucleic Acids Symposium Series, 41, 95 - 98." type="journal article" year="1999">Hall 1999</bibRefCitation>
). Specimen ID with DNA voucher code and GenBank accession code of the COI sequences, are listed as follows: 
<taxonomicName id="4C6B4D60FFADFF80DFCDFEA02431FEAA" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[151,287,332,356]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFADFF80DFCDFEA02431FEAA" box="[151,287,332,356]" italics="true" pageId="4" pageNumber="393">K.tannuolus</emphasis>
</taxonomicName>
HT; 
<taxonomicName id="4C6B4D60FFADFF80DE02FEA024C7FEAB" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[344,489,332,357]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="tannuolus">K.tannuolus</taxonomicName>
1; 
<accessionNumber id="9438AB00FFADFF80DEA7FEA02754FEAB" box="[509,634,332,357]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF071456" pageId="4" pageNumber="393">
MF 
<date id="FFD51023FFADFF80DD7CFEA02754FEAB" box="[550,634,332,357]" pageId="4" pageNumber="393" value="56-07-14">071456</date>
</accessionNumber>
and 
<taxonomicName id="4C6B4D60FFADFF80DDEAFEA02616FEAA" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[688,824,332,356]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFADFF80DDEAFEA02616FEAA" box="[688,824,332,356]" italics="true" pageId="4" pageNumber="393">K.tannuolus</emphasis>
</taxonomicName>
PT; 
<taxonomicName id="4C6B4D60FFADFF80DC37FEA026D7FEAB" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[877,1017,332,357]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="tannuolus">K.tannuolus</taxonomicName>
2; 
<accessionNumber id="9438AB00FFADFF80DB48FEA021BCFEAB" box="[1042,1170,332,357]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF071457" pageId="4" pageNumber="393">
MF 
<date id="FFD51023FFADFF80DB66FEA021BCFEAB" box="[1084,1170,332,357]" pageId="4" pageNumber="393" value="57-07-14">071457</date>
</accessionNumber>
.
</paragraph>
<paragraph id="8BD436E3FFADFF80DF9DFE8321D7FE1E" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="393">
Molecular variation based on the Kimura two-parameter distance model (
<bibRefCitation id="EFFA4B12FFADFF80DCA4FE8321B2FE46" author="Kimura" box="[1022,1180,367,392]" pageId="4" pageNumber="393" refString="Kimura, M (1980) A simple method for estimating evolutionary rate of base substitutions through comparative studies of nucleotide sequences. Journal of Molecular Evolution, 16, 111 - 120." type="journal article" year="1980">Kimura, 1980</bibRefCitation>
) for 658 bp COI DNA barcodes between two specimens of 
<taxonomicName id="4C6B4D60FFADFF80DD1FFE7927F6FE62" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[581,728,404,428]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFADFF80DD1FFE7927F6FE62" box="[581,728,404,428]" italics="true" pageId="4" pageNumber="393">K. tannuolus</emphasis>
</taxonomicName>
was only 0.15 %. Therefore we conclude that the barcoded specimens of 
<taxonomicName id="4C6B4D60FFADFF80DE6BFE5524EEFE1E" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[305,448,440,464]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="tannuolus">
<emphasis id="B91FEAF1FFADFF80DE6BFE5524EEFE1E" box="[305,448,440,464]" italics="true" pageId="4" pageNumber="393">K. tannuolus</emphasis>
</taxonomicName>
belong to the same species, despite the aberrant genitalia of the 
<typeStatus id="54D08841FFADFF80DBCBFE5421DBFE1E" box="[1169,1269,440,464]" pageId="4" pageNumber="393" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716568FFADFF80DF9DFE302694FD4B" pageId="4" pageNumber="393" type="biology_ecology">
<paragraph id="8BD436E3FFADFF80DF9DFE302694FD4B" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="393">
<materialsCitation id="3B033CBEFFADFF80DF9DFE30261CFDD6" ID-GBIF-Occurrence="1585957405" collectingDate="9 - 12 June 1995, 12 - 13 July 2004 and 21 July 2002" location="Russian Siberia" pageId="4" pageNumber="393" specimenCount="1" stateProvince="Tuva">
<emphasis id="B91FEAF1FFADFF80DF9DFE3024DEFE3B" bold="true" box="[199,496,476,501]" pageId="4" pageNumber="393">Biology and distribution.</emphasis>
Three males were found during the day on 
<collectingDate id="EF91E9CBFFADFF80DC8DFE30242BFDD6" isEnumeration="true" pageId="4" pageNumber="393">
<date id="FFD51023FFADFF80DC8DFE3021ABFE3B" box="[983,1157,476,501]" pageId="4" pageNumber="393" value="1995-06-09" valueMax="1995-06-12" valueMin="1995-06-09">9-12 June 1995</date>
, 
<date id="FFD51023FFADFF80DBCAFE302064FE3A" box="[1168,1354,476,501]" pageId="4" pageNumber="393" value="2004-07-12" valueMax="2004-07-13" valueMin="2004-07-12">12–13 July 2004</date>
and 
<date id="FFD51023FFADFF80DADAFE30242BFDD6" pageId="4" pageNumber="393" value="2002-07-21">
21 July 
<quantity id="4C939B06FFADFF80DF97FDEC242BFDD6" box="[205,261,512,536]" metricMagnitude="1" metricUnit="m" metricValue="5.08508" pageId="4" pageNumber="393" unit="in" value="2002.0">2002</quantity>
</date>
</collectingDate>
in a remote part of 
<location id="8EB46038FFADFF80DEB2FE1327B5FDD6" LSID="urn:lsid:plazi:treatment:03C287F5FFA9FF80DFCDF952265AFD66:8EB46038FFADFF80DEB2FE1327B5FDD6" box="[488,667,511,536]" name="Russian Siberia" pageId="4" pageNumber="393" stateProvince="Tuva">Russian Siberia</location>
, 
<collectingRegion id="49AFF801FFADFF80DDFDFDEC27CEFDD9" box="[679,736,512,535]" country="Russia" name="Tyva" pageId="4" pageNumber="393">Tuva</collectingRegion>
region
</materialsCitation>
. 
<materialsCitation id="3B033CBEFFADFF80DC64FE132694FD4B" ID-GBIF-Occurrence="1585957403" county="The" location="East Tannu-Ola mountains" municipality="Durgen river" pageId="4" pageNumber="393" specimenCount="1" stateProvince="Tuva" typeStatus="holotype">
<collectingCounty id="62B54E6FFFADFF80DC64FE132644FDD6" box="[830,874,511,536]" pageId="4" pageNumber="393">The</collectingCounty>
species was collected at 1250 meters a.s.l. on the right bank of the 
<collectingMunicipality id="6BB0AC99FFADFF80DE0FFDC924CAFDF3" box="[341,484,548,573]" pageId="4" pageNumber="393">Durgen river</collectingMunicipality>
on north slope of the 
<location id="8EB46038FFADFF80DD8DFDC9212EFDF3" LSID="urn:lsid:plazi:treatment:03C287F5FFA9FF80DFCDF952265AFD66:8EB46038FFADFF80DD8DFDC9212EFDF3" box="[727,1024,548,573]" county="The" municipality="Durgen river" name="East Tannu-Ola mountains" pageId="4" pageNumber="393" stateProvince="Tuva">East Tannu-Ola mountains</location>
and sand dunes near 
<location id="8EB46038FFADFF80DBB7FDC820B6FDF3" LSID="urn:lsid:plazi:treatment:03C287F5FFA9FF80DFCDF952265AFD66:8EB46038FFADFF80DBB7FDC820B6FDF3" box="[1261,1432,548,573]" county="The" municipality="Durgen river" name="Tere-Khol lake" pageId="4" pageNumber="393" stateProvince="Tuva">Tere-Khol lake</location>
. 
<location id="8EB46038FFADFF80DFCDFDAB25EDFDAE" LSID="urn:lsid:plazi:treatment:03C287F5FFA9FF80DFCDF952265AFD66:8EB46038FFADFF80DFCDFDAB25EDFDAE" box="[151,195,583,608]" county="The" municipality="Durgen river" name="The" pageId="4" pageNumber="393" stateProvince="Tuva">The</location>
<typeStatus id="54D08841FFADFF80DF96FDA425D2FDAE" box="[204,252,584,608]" pageId="4" pageNumber="393">type</typeStatus>
locality of 
<taxonomicName id="4C6B4D60FFADFF80DED9FDA4273AFDAE" authorityName="Saldaitis &amp; Yakovlev &amp; Truuverk" authorityYear="2017" box="[387,532,583,608]" class="Insecta" family="Cossidae" genus="Kerzhnerocossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="tannuolus">K. tannuolus</taxonomicName>
is situated on flat river bank with 
<taxonomicName id="4C6B4D60FFADFF80DCFFFDA421B9FDAE" authority="Ledeb." authorityName="Ledeb." box="[933,1175,583,608]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="393" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="B91FEAF1FFADFF80DCFFFDA4216FFDAE" box="[933,1089,584,608]" italics="true" pageId="4" pageNumber="393">Larix sibirica</emphasis>
Ledeb.
</taxonomicName>
forest, with 
<taxonomicName id="4C6B4D60FFADFF80DA70FDA42437FD4B" authority="Pall." authorityName="Pall." class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="393" phylum="Tracheophyta" rank="species" species="jubata">
<emphasis id="B91FEAF1FFADFF80DA70FDA425F1FD4A" italics="true" pageId="4" pageNumber="393">Caragana jubata</emphasis>
Pall.
</taxonomicName>
shrubs and 
<taxonomicName id="4C6B4D60FFADFF80DEC4FD8024F8FD4A" box="[414,470,620,644]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="4" pageNumber="393" phylum="Chordata" rank="genus">
<emphasis id="B91FEAF1FFADFF80DEC4FD8024F8FD4A" box="[414,470,620,644]" italics="true" pageId="4" pageNumber="393">Salix</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6B4D60FFADFF80DD57FD802779FD4A" box="[525,599,620,644]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="393" phylum="Tracheophyta" rank="genus">
<emphasis id="B91FEAF1FFADFF80DD57FD802779FD4A" box="[525,599,620,644]" italics="true" pageId="4" pageNumber="393">Betula</emphasis>
</taxonomicName>
(pers. comm. Martin Česanek).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716568FFADFF80DF9DFD63265AFD66" box="[199,884,655,680]" pageId="4" pageNumber="393" type="etymology">
<paragraph id="8BD436E3FFADFF80DF9DFD63265AFD66" blockId="4.[151,1437,151,680]" box="[199,884,655,680]" pageId="4" pageNumber="393">
<emphasis id="B91FEAF1FFADFF80DF9DFD632460FD66" bold="true" box="[199,334,655,680]" pageId="4" pageNumber="393">Etymology.</emphasis>
The new species is named after the 
<typeStatus id="54D08841FFADFF80DDB9FD7C263DFD66" box="[739,787,656,680]" pageId="4" pageNumber="393">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>