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<document id="7169A151CF6FB0DF23A1CAAD1AF32716" ID="10.11646/zootaxa.4136.3.6" ID-DOI="10.11646/zootaxa.4136.3.6" ID-GBIF-Dataset="cafaba1e-ca02-4145-99ac-7fc197006b21" ID-ISSN="1175-5326" ID-Zenodo-Dep="271917" ID-ZooBank="0F911AA3-A875-425E-A196-221C7C1A4CB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1467880999355" checkinUser="plazi" docAuthor="Barrio-Amorós, Cesar L. &amp; Ortíz, Juan C." docDate="2016" docId="03C287EBFFE1FFA30AA32940F1F3FA17" docLanguage="en" docName="zootaxa.4136.3.6.pdf" docOrigin="Zootaxa 4136 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gonatodes petersi Donoso-Barros 1967" docType="treatment" docVersion="11" lastPageNumber="550" masterDocId="FFFBFF93FFEBFFAE0A342868F072FF8A" masterDocTitle="Venezuelan geckos (Gekkonidae, Phyllodactylidae, Sphaerodactylidae) in the collection of the Universidad de Concepción in Chile, with description of the type series of Gonatodes ligiae and Gonatodes petersi (Sphaerodactylidae)" masterLastPageNumber="552" masterPageNumber="537" pageNumber="547" updateTime="1698363896553" updateUser="plazi">
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<mods:title id="2117B6F6585FE609FF7DA58718E147B3">Venezuelan geckos (Gekkonidae, Phyllodactylidae, Sphaerodactylidae) in the collection of the Universidad de Concepción in Chile, with description of the type series of Gonatodes ligiae and Gonatodes petersi (Sphaerodactylidae)</mods:title>
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<mods:namePart id="33ED9EDECAC9E74DCF316AB91682D4B4">Barrio-Amorós, Cesar L.</mods:namePart>
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<mods:namePart id="F6DCFD55404D37290D7DFB7896F2B34C">Ortíz, Juan C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4CC1DA6D68DD14810C9C16230F943B21">text</mods:typeOfResource>
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<mods:title id="8BF3777BEBDCE2546FFAD283269C5D7F">Zootaxa</mods:title>
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<mods:part id="608E1D617DB6C3234BF831B20B135A38">
<mods:date id="3BA12D6F4F3479899DACD28A2916CD07">2016</mods:date>
<mods:detail id="612B31335FDD10241BD4AF2B5215EBE4" type="volume">
<mods:number id="C9AA218997AA0DF5FDE33A8FA230E43A">4136</mods:number>
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<mods:classification id="4E44099615423928E8B4766C4246F8E3">journal article</mods:classification>
<mods:identifier id="D2231E71D0AE5575CC5A110134A57703" type="DOI">10.11646/zootaxa.4136.3.6</mods:identifier>
<mods:identifier id="19BCF9728979C2E09DAABED4E51757D4" type="GBIF-Dataset">cafaba1e-ca02-4145-99ac-7fc197006b21</mods:identifier>
<mods:identifier id="D29D9E07A6D71350879FBADD78979C86" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FB6E1D8D5BF4AB991E5F80F1CE944054" type="Zenodo-Dep">271917</mods:identifier>
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<treatment id="03C287EBFFE1FFA30AA32940F1F3FA17" ID-DOI="http://doi.org/10.5281/zenodo.5697926" ID-GBIF-Taxon="121545907" ID-Zenodo-Dep="5697926" LSID="urn:lsid:plazi:treatment:03C287EBFFE1FFA30AA32940F1F3FA17" httpUri="http://treatment.plazi.org/id/03C287EBFFE1FFA30AA32940F1F3FA17" lastPageId="13" lastPageNumber="550" pageId="10" pageNumber="547">
<subSubSection id="C3716576FFE1FFA40AA32940F152FEE9" pageId="10" pageNumber="547" type="nomenclature">
<paragraph id="8BD436FDFFE1FFA40AA32940F2F0FEC8" blockId="10.[151,642,296,355]" box="[151,642,296,322]" pageId="10" pageNumber="547">
<heading id="D09C8191FFE1FFA40AA32940F2F0FEC8" bold="true" box="[151,642,296,322]" fontSize="11" level="1" pageId="10" pageNumber="547" reason="1">
<emphasis id="B91FEAEFFFE1FFA40AA32940F2F0FEC8" bold="true" box="[151,642,296,322]" pageId="10" pageNumber="547">
<taxonomicName id="4C6B4D7EFFE1FFA40AA32940F20CFEC8" authority="Donoso-Barros, 1967" authorityName="Donoso-Barros" authorityYear="1967" box="[151,638,296,322]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE1FFA40AA32940F11DFEC8" bold="true" box="[151,367,296,322]" italics="true" pageId="10" pageNumber="547">Gonatodes petersi</emphasis>
<bibRefCitation id="EFFA4B0CFFE1FFA40B422940F20CFEC8" author="Donoso-Barros" box="[374,638,296,322]" pageId="10" pageNumber="547" refString="Donoso-Barros, R. (1967) Diagnosis de dos nuevas especies del genero Gonatodes de Venezuela. Noticiario Mensual del Museo Nacional de Historia Natural, 129, fourth page (not numbered)." type="book chapter" year="1967">Donoso-Barros, 1967</bibRefCitation>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AA32922F152FEE9" blockId="10.[151,642,296,355]" box="[151,288,330,355]" pageId="10" pageNumber="547">
(
<figureCitation id="13502A78FFE1FFA40AAB2922F093FEE9" box="[159,225,330,355]" captionStart="FIGURE 6" captionStartId="11.[151,250,1906,1928]" captionTargetBox="[226,1359,373,1883]" captionTargetId="figure@11.[226,1360,373,1885]" captionTargetPageId="11" captionText="FIGURE 6. Gonatodes petersi. A: male; B: female. La Orchila, Sierra de Perijá, estado Zulia. Uncollected specimens. Photos: Fernando Rojas-Runjaic." httpUri="https://zenodo.org/record/271923/files/figure.png" pageId="10" pageNumber="547">Fig. 6</figureCitation>
A–B)
</paragraph>
</subSubSection>
<subSubSection id="C3716576FFE1FFA40AA329FBF56EFE79" pageId="10" pageNumber="547" type="description">
<paragraph id="8BD436FDFFE1FFA40AA329FBF56EFE79" blockId="10.[151,1436,402,499]" pageId="10" pageNumber="547">
<bibRefCitation id="EFFA4B0CFFE1FFA40AA329FBF1E1FE21" author="Donoso-Barros" box="[151,403,402,427]" pageId="10" pageNumber="547" refString="Donoso-Barros, R. (1967) Diagnosis de dos nuevas especies del genero Gonatodes de Venezuela. Noticiario Mensual del Museo Nacional de Historia Natural, 129, fourth page (not numbered)." type="book chapter" year="1967">Donoso-Barros (1967)</bibRefCitation>
briefly defines
<taxonomicName id="4C6B4D7EFFE1FFA4087029FBF362FE20" box="[580,784,403,427]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE1FFA4087029FBF362FE20" box="[580,784,403,427]" italics="true" pageId="10" pageNumber="547">Gonatodes petersi</emphasis>
</taxonomicName>
. The translation of the original paragraph in Spanish is: A
<taxonomicName id="4C6B4D7EFFE1FFA40AA329DEF162FE44" box="[151,272,438,462]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="genus">
<emphasis id="B91FEAEFFFE1FFA40AA329DEF162FE44" box="[151,272,438,462]" italics="true" pageId="10" pageNumber="547">Gonatodes</emphasis>
</taxonomicName>
of the
<taxonomicName id="4C6B4D7EFFE1FFA40B6829DFF1DDFE44" box="[348,431,439,462]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="B91FEAEFFFE1FFA40B6829DFF1DDFE44" box="[348,431,439,462]" italics="true" pageId="10" pageNumber="547">vittatus</emphasis>
</taxonomicName>
Group, but larger, with rounded snout. The vertebral stripe with three “swellings” through its length. 23 lamellae under fourth toe”. We provide detailed descriptions of the
<typeStatus id="54D0885FFFE1FFA40E1629B2F4F6FE79" box="[1058,1156,474,499]" pageId="10" pageNumber="547" type="holotype">holotype</typeStatus>
and allotype.
</paragraph>
</subSubSection>
<subSubSection id="C3716576FFE1FFA40AA32A4BF33EFC99" pageId="10" pageNumber="547" type="reference_group">
<paragraph id="8BD436FDFFE1FFA40AA32A4BF229FDB1" blockId="10.[151,844,546,787]" box="[151,603,546,571]" pageId="10" pageNumber="547">
<taxonomicName id="4C6B4D7EFFE1FFA40AA32A4BF229FDB1" authority="Donoso-Barros, 1967" authorityName="Donoso-Barros" authorityYear="1967" box="[151,603,546,571]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE1FFA40AA32A4BF111FDB0" box="[151,355,547,571]" italics="true" pageId="10" pageNumber="547">Gonatodes petersi</emphasis>
<bibRefCitation id="EFFA4B0CFFE1FFA40B5E2A4BF229FDB1" author="Donoso-Barros" box="[362,603,546,571]" pageId="10" pageNumber="547" refString="Donoso-Barros, R. (1967) Diagnosis de dos nuevas especies del genero Gonatodes de Venezuela. Noticiario Mensual del Museo Nacional de Historia Natural, 129, fourth page (not numbered)." type="book chapter" year="1967">Donoso-Barros, 1967</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AA32A2EF21CFDD5" blockId="10.[151,844,546,787]" box="[151,622,582,607]" pageId="10" pageNumber="547">
<taxonomicName id="4C6B4D7EFFE1FFA40AA32A2EF218FDD5" authority="Aleman, 1953: 291" authorityName="Aleman" authorityPageNumber="291" authorityYear="1953" box="[151,618,582,607]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="B91FEAEFFFE1FFA40AA32A2EF162FDD4" box="[151,272,582,606]" italics="true" pageId="10" pageNumber="547">Gonatodes</emphasis>
cf.
<emphasis id="B91FEAEFFFE1FFA40B0F2A2FF1F3FDD5" box="[315,385,583,607]" italics="true" pageId="10" pageNumber="547">fuscus</emphasis>
: Alemán, 1953: 291
</taxonomicName>
.
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AA32A03F2EEFD2D" blockId="10.[151,844,546,787]" pageId="10" pageNumber="547">
<treatmentCitationGroup id="AB7B11D3FFE1FFA40AA32A03F2EEFD2D" pageId="10" pageNumber="547">
<taxonomicName id="4C6B4D7EFFE1FFA40AA32A03F215FD09" authority="Donoso-Barros, 1967" authorityName="Donoso-Barros" authorityYear="1967" box="[151,615,618,643]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE1FFA40AA32A03F111FD08" box="[151,355,619,643]" italics="true" pageId="10" pageNumber="547">Gonatodes petersi</emphasis>
:
<bibRefCitation id="EFFA4B0CFFE1FFA40B462A03F215FD09" author="Donoso-Barros" box="[370,615,618,643]" pageId="10" pageNumber="547" refString="Donoso-Barros, R. (1967) Diagnosis de dos nuevas especies del genero Gonatodes de Venezuela. Noticiario Mensual del Museo Nacional de Historia Natural, 129, fourth page (not numbered)." type="book chapter" year="1967">Donoso-Barros, 1967</bibRefCitation>
</taxonomicName>
: not numbered page
<taxonomicName id="4C6B4D7EFFE1FFA40AA32AE6F2EAFD2D" authority="Rivero-Blanco, 1968: 115" authorityName="Rivero-Blanco" authorityPageNumber="115" authorityYear="1968" box="[151,664,654,679]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE1FFA40AA32AE6F111FD2C" box="[151,355,654,678]" italics="true" pageId="10" pageNumber="547">Gonatodes petersi</emphasis>
:
<treatmentCitation id="0ACA10ECFFE1FFA40B462AE6F2EAFD2D" author="Rivero-Blanco" box="[370,664,654,679]" page="115" pageId="10" pageNumber="547" year="1968">
<bibRefCitation id="EFFA4B0CFFE1FFA40B462AE6F213FD2D" author="Rivero-Blanco" box="[370,609,654,679]" pageId="10" pageNumber="547" refString="Rivero-Blanco, C. (1968) Un genero y dos especies de tuqueques (Sauria: Sphaerodactylinae) citados por primera vez para Venezuela, con notas sobre la distribucion de otras especies poco conocidas. Memoria de la Sociedad de Ciencias Naturales La Salle, 27, 104 - 119." type="journal article" year="1968">Rivero-Blanco, 1968</bibRefCitation>
: 115
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AA32ADBF2E4FD65" blockId="10.[151,844,546,787]" pageId="10" pageNumber="547">
<treatmentCitationGroup id="AB7B11D3FFE1FFA40AA32ADBF2E4FD65" pageId="10" pageNumber="547">
<taxonomicName id="4C6B4D7EFFE1FFA40AA32ADBF375FD41" authority="Peters &amp; Donoso-Barros, 1970: 133" authorityName="Peters &amp; Donoso-Barros" authorityPageNumber="133" authorityYear="1970" box="[151,775,691,715]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE1FFA40AA32ADBF111FD40" box="[151,355,691,715]" italics="true" pageId="10" pageNumber="547">Gonatodes petersi</emphasis>
:
<treatmentCitation id="0ACA10ECFFE1FFA40B462ADBF375FD41" author="Peters" box="[370,775,691,715]" page="133" pageId="10" pageNumber="547" year="1970">
<bibRefCitation id="EFFA4B0CFFE1FFA40B462ADBF2BDFD41" author="Peters" box="[370,719,691,715]" pageId="10" pageNumber="547" refString="Peters, J. A. &amp; Donoso-Barros, R. (1970) Catalogue of the Neotropical Squamata. Part II. Lizards and Amphisbaenians. United States national Museum Bulletin, 297, 293 pp." type="book" year="1970">Peters &amp; Donoso-Barros, 1970</bibRefCitation>
: 133
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="4C6B4D7EFFE1FFA40AA32ABEF2E4FD65" authority="Rivero-Blanco, 1979: 121" authorityName="Rivero-Blanco" authorityPageNumber="121" authorityYear="1979" box="[151,662,726,751]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE1FFA40AA32ABEF111FD64" box="[151,355,726,750]" italics="true" pageId="10" pageNumber="547">Gonatodes petersi</emphasis>
: Rivero-Blanco, 1979: 121
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AA32A93F33EFC99" blockId="10.[151,844,546,787]" box="[151,844,762,787]" pageId="10" pageNumber="547">
<treatmentCitationGroup id="AB7B11D3FFE1FFA40AA32A93F33EFC99" box="[151,844,762,787]" pageId="10" pageNumber="547">
<taxonomicName id="4C6B4D7EFFE1FFA40AA32A93F33AFC99" authority="Rojas-Runjaic &amp; Infante-Rivero 2009: 83" authorityName="Rojas-Runjaic &amp; Infante-Rivero" authorityPageNumber="83" authorityYear="2009" box="[151,840,762,787]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE1FFA40AA32A93F111FC98" box="[151,355,763,787]" italics="true" pageId="10" pageNumber="547">Gonatodes petersi</emphasis>
: Rojas-Runjaic &amp;
<treatmentCitation id="0ACA10ECFFE1FFA408022A92F33AFC99" author="Infante-Rivero" box="[566,840,762,787]" page="83" pageId="10" pageNumber="547" year="2009">
<bibRefCitation id="EFFA4B0CFFE1FFA408022A92F36CFC99" author="Infante-Rivero" box="[566,798,762,787]" pageId="10" pageNumber="547" refString="Rojas - Runjaic, F. J. M. &amp; Infante-Rivero, E. (2009) Redescripcion de Gonatodes petersi Donoso-Barros, 1967 (Squamata: Gekkonidae), un tuqueque endemico de la vertiente venezolana de la Sierra de Perija. Memoria de la Fundacion La Salle de Ciencias Naturales, 170, 81 - 98." type="journal article" year="2009">Infante-Rivero 2009</bibRefCitation>
: 83
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3716576FFE1FFA40AA32B2AF2DFFC4D" pageId="10" pageNumber="547" type="materials_examined">
<paragraph id="8BD436FDFFE1FFA40AA32B2AF48CFCF5" blockId="10.[151,1437,834,2011]" pageId="10" pageNumber="547">
<emphasis id="B91FEAEFFFE1FFA40AA32B2AF179FCD1" bold="true" box="[151,267,834,859]" pageId="10" pageNumber="547">
<typeStatus id="54D0885FFFE1FFA40AA32B2AF174FCD1" box="[151,262,834,859]" pageId="10" pageNumber="547" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED7AAE38FFE1FFA40B272B2BF116FCD1" box="[275,356,835,859]" country="Italy" httpUri="http://grbio.org/cool/70mh-t5r1" name="Universita di Cagliari" pageId="10" pageNumber="547">MZUC</collectionCode>
8836 (field number in
<collectingCountry id="F37C766DFFE1FFA4085B2B2AF290FCD1" box="[623,738,834,859]" name="Venezuela" pageId="10" pageNumber="547">Venezuela</collectingCountry>
276; number in R. Donoso-Barros collection RDB 1511), an adult male, collected by RDB at Sierra de Perijá, estado Zulia,
<collectingCountry id="F37C766DFFE1FFA4096C2B0EF3B9FCF5" box="[856,971,870,895]" name="Venezuela" pageId="10" pageNumber="547">Venezuela</collectingCountry>
<date id="FFD5103DFFE1FFA409E62B0EF415FCF5" box="[978,1127,870,895]" pageId="10" pageNumber="547" value="1964-05-26">26 May 1964</date>
(
<figureCitation id="13502A78FFE1FFA40E422B0EF4CBFCF5" box="[1142,1209,870,895]" captionStart="FIGURE 7" captionStartId="12.[151,250,1969,1991]" captionTargetBox="[285,1335,225,1908]" captionTargetId="figure@12.[248,1335,193,1927]" captionTargetPageId="12" captionText="FIGURE 7. Gonatodes petersi. A: holotype, dorsal view. B: ventral view. C: Alotype dorsal view. D: alotype, ventral view. Not to scale. Photos by Juan Pablo Donoso." httpUri="https://zenodo.org/record/271924/files/figure.png" pageId="10" pageNumber="547">Fig. 7</figureCitation>
A–B).
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AF32BE2F2DFFC4D" blockId="10.[151,1437,834,2011]" pageId="10" pageNumber="547">
<emphasis id="B91FEAEFFFE1FFA40AF32BE2F141FC29" bold="true" box="[199,307,906,931]" pageId="10" pageNumber="547">Allotype.</emphasis>
<collectionCode id="ED7AAE38FFE1FFA40B0B2BE3F1E2FC29" box="[319,400,907,931]" country="Italy" httpUri="http://grbio.org/cool/70mh-t5r1" name="Universita di Cagliari" pageId="10" pageNumber="547">MZUC</collectionCode>
12105 (field number in
<collectingCountry id="F37C766DFFE1FFA4088C2BE2F359FC29" box="[696,811,906,931]" name="Venezuela" pageId="10" pageNumber="547">Venezuela</collectingCountry>
V277; number in R. Donoso-Barros collection RDB 1582), with same data as
<typeStatus id="54D0885FFFE1FFA40B802BC6F264FC4D" box="[436,534,942,967]" pageId="10" pageNumber="547" type="holotype">holotype</typeStatus>
(
<figureCitation id="13502A78FFE1FFA408112BC6F215FC4D" box="[549,615,942,967]" captionStart="FIGURE 7" captionStartId="12.[151,250,1969,1991]" captionTargetBox="[285,1335,225,1908]" captionTargetId="figure@12.[248,1335,193,1927]" captionTargetPageId="12" captionText="FIGURE 7. Gonatodes petersi. A: holotype, dorsal view. B: ventral view. C: Alotype dorsal view. D: alotype, ventral view. Not to scale. Photos by Juan Pablo Donoso." httpUri="https://zenodo.org/record/271924/files/figure.png" pageId="10" pageNumber="547">Fig. 7</figureCitation>
C–D).
</paragraph>
</subSubSection>
<subSubSection id="C3716576FFE1FFA30AF32BBAF13DFB62" lastPageId="13" lastPageNumber="550" pageId="10" pageNumber="547" type="description">
<paragraph id="8BD436FDFFE1FFA40AF32BBAF116FBF1" blockId="10.[151,1437,834,2011]" pageId="10" pageNumber="547">
<emphasis id="B91FEAEFFFE1FFA40AF32BBAF195FC61" bold="true" box="[199,487,978,1003]" pageId="10" pageNumber="547">
Description of
<typeStatus id="54D0885FFFE1FFA40B4E2BBAF195FC61" box="[378,487,978,1003]" pageId="10" pageNumber="547" type="holotype">Holotype</typeStatus>
</emphasis>
. An adult male, with snout-vent length of
<quantity id="4C939B18FFE1FFA409E42BBAF44BFC60" box="[976,1081,978,1003]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="10" pageNumber="547" unit="mm" value="35.3">35.3 mm</quantity>
. Head approximately 1.4 times longer than wide (HL:
<quantity id="4C939B18FFE1FFA40BAE2B9FF270FB84" box="[410,514,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="10" pageNumber="547" unit="mm" value="10.8">10.8 mm</quantity>
; HW:
<quantity id="4C939B18FFE1FFA4087F2B9EF2D7FB85" box="[587,677,1014,1039]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="547" unit="mm" value="6.2">6.2 mm</quantity>
), 30.5% of SVL. Snout
<quantity id="4C939B18FFE1FFA409802B9FF46BFB84" box="[948,1049,1015,1039]" metricMagnitude="-3" metricUnit="m" metricValue="3.83" pageId="10" pageNumber="547" unit="mm" value="3.83">3.83 mm</quantity>
long, round, moderately elongate, subacuminate dorsally, sloping toward top of head. Neck narrower than head and body. Body nearly cylindrical but wider than high; axilla-groin distance
<quantity id="4C939B18FFE1FFA408722C56F2DCFBDC" box="[582,686,1086,1111]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="10" pageNumber="547" unit="mm" value="15.8">15.8 mm</quantity>
. Limbs well developed, fourth toe length
<quantity id="4C939B18FFE1FFA40EB32C56F493FBDC" box="[1159,1249,1086,1111]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="10" pageNumber="547" unit="mm" value="4.6">4.6 mm</quantity>
. Tail completely missing (
<figureCitation id="13502A78FFE1FFA40AC92C0AF133FBF1" box="[253,321,1122,1147]" captionStart="FIGURE 7" captionStartId="12.[151,250,1969,1991]" captionTargetBox="[285,1335,225,1908]" captionTargetId="figure@12.[248,1335,193,1927]" captionTargetPageId="12" captionText="FIGURE 7. Gonatodes petersi. A: holotype, dorsal view. B: ventral view. C: Alotype dorsal view. D: alotype, ventral view. Not to scale. Photos by Juan Pablo Donoso." httpUri="https://zenodo.org/record/271924/files/figure.png" pageId="10" pageNumber="547">Fig. 7</figureCitation>
A).
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AF32CEEF349F98C" blockId="10.[151,1437,834,2011]" pageId="10" pageNumber="547">
Rostral moderately large, visible from above, rounded, similar to that observed in other
<taxonomicName id="4C6B4D7EFFE1FFA40E872CEEF55EFB14" box="[1203,1324,1158,1182]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="genus">
<emphasis id="B91FEAEFFFE1FFA40E872CEEF55EFB14" box="[1203,1324,1158,1182]" italics="true" pageId="10" pageNumber="547">Gonatodes</emphasis>
</taxonomicName>
except in those species with acuminate snout (e.g.
<taxonomicName id="4C6B4D7EFFE1FFA408532CC3F2AFFB49" box="[615,733,1195,1219]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="daudini">
<emphasis id="B91FEAEFFFE1FFA408532CC3F2AFFB49" box="[615,733,1195,1219]" italics="true" pageId="10" pageNumber="547">G. daudini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D7EFFE1FFA409222CC3F3D5FB49" box="[790,935,1195,1219]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="547" phylum="Chordata" rank="species" species="naufragus">
<emphasis id="B91FEAEFFFE1FFA409222CC3F3D5FB49" box="[790,935,1195,1219]" italics="true" pageId="10" pageNumber="547">G. naufragus</emphasis>
</taxonomicName>
); irregularly shaped, much wider than long, but with a small longitudinal groove that divides the scale almost completely except for a short section towards the lip. Three postrostrals, the lateral two (supranasals) distinctly larger than the median postrostral; median postrostral slightly larger than adjacent posterior scales on snout. Nostril bordered proximally by rostral, from above by supra rostral, posteriorly by three (left) and three (right) postnasals. Postnasals slightly larger than adjacent loreals. Scales on top of snout rounded, juxtaposed, relatively uniform in size. Canthus rostralis gently rounded. Loreal region with scales slightly more elongate than those on snout, largest on row adjacent to supralabials; those anterior to the eyelids more conical; 10/9 (right/left side) loreals in a straight line between postnasals and anterior margin of orbit. Top and posterior portion of the head with granular scales smaller than those on top of snout. Scales decrease slightly in size from the postrostrals toward interocular area.
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AE62E7AF176F931" blockId="10.[151,1437,834,2011]" pageId="10" pageNumber="547">
Supraciliary flap missing, with only 1/1 (left/right) barely elongate supraciliary spine on each side. Pupil round, eye diameter
<quantity id="4C939B18FFE1FFA40B482E5FF1A6F9C4" box="[380,468,1591,1615]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="547" unit="mm" value="2.3">2.3 mm</quantity>
. Scales on temporal region similar to those on top of head. Ear-opening small,
<quantity id="4C939B18FFE1FFA40F742E5EF5EAF9C4" box="[1344,1432,1590,1615]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="547" unit="mm" value="0.6">0.6 mm</quantity>
, elongate, posterior to, and at the same level of, angle of mouth. Supralabials 6/6 (right/left) to center of eye, distinctly enlarged anteriorly and decreasing in size posteriorly, followed by much smaller scales along lip to rictus of mouth.
</paragraph>
<paragraph id="8BD436FDFFE1FFA40AF32EAEF094F8E5" blockId="10.[151,1437,834,2011]" pageId="10" pageNumber="547">Mental large, distinctly wider anteriorly than posteriorly, with regular posterior margin. Postmentals number two, hexagonal, larger than adjacent posterior scales. Scales on chin flat, smooth, hexagonal to rounded, juxtaposed, larger anteriorly and medially, decreasing in size posteriorly. Infralabials 4/4 to center of eye, decreasing in size posteriorly. There are small scales (4 left side/3 right side) on the commissure to the corner of the mouth.</paragraph>
<paragraph id="8BD436FDFFE1FFA40AF32F12F4FEF83D" blockId="10.[151,1437,834,2011]" pageId="10" pageNumber="547">Scales on nape and sides of neck very small, similar in size, and granular, slightly conical. Scales on throat small, slightly granular, with a rather abrupt transitional zone between the throat and neck.</paragraph>
<paragraph id="8BD436FDFFE1FFA50AF32FAAF3DDFECA" blockId="10.[151,1437,834,2011]" lastBlockId="11.[151,1436,151,320]" lastPageId="11" lastPageNumber="548" pageId="10" pageNumber="547">
Dorsal scales granular, juxtaposed, similar in size all over. Ventrals larger than dorsals, roughly hexagonal, flat, smooth, imbricate, in oblique rows also forming rather regular longitudinal rows on the belly, with 53 scales along the midventral line between the anterior part of the arm insertions and the vent (not including small granules lining the edge of the cloaca). Part of the left flank is damaged and some skin missing; is not possible to count the scales around midbody. Scales on precloacal plate similar to ventrals, except for those bordering the vent, which are disposed in one row of very small scales. Escutcheon absent (
<figureCitation id="13502A78FFE0FFA5097D294FF3FFFECA" box="[841,909,295,320]" captionStart="FIGURE 7" captionStartId="12.[151,250,1969,1991]" captionTargetBox="[285,1335,225,1908]" captionTargetId="figure@12.[248,1335,193,1927]" captionTargetPageId="12" captionText="FIGURE 7. Gonatodes petersi. A: holotype, dorsal view. B: ventral view. C: Alotype dorsal view. D: alotype, ventral view. Not to scale. Photos by Juan Pablo Donoso." httpUri="https://zenodo.org/record/271924/files/figure.png" pageId="11" pageNumber="548">Fig. 7</figureCitation>
B).
</paragraph>
<caption id="DF146675FFE0FFA50AA32F1AF1E7F82D" httpUri="https://zenodo.org/record/271923/files/figure.png" pageId="11" pageNumber="548" targetBox="[226,1359,373,1883]" targetPageId="11">
<paragraph id="8BD436FDFFE0FFA50AA32F1AF1E7F82D" blockId="11.[151,1436,1906,1959]" pageId="11" pageNumber="548">
<emphasis id="B91FEAEFFFE0FFA50AA32F1AF160F80D" bold="true" box="[151,274,1906,1928]" pageId="11" pageNumber="548">FIGURE 6.</emphasis>
<taxonomicName id="4C6B4D7EFFE0FFA50B2D2F1AF1A0F802" box="[281,466,1906,1928]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="548" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE0FFA50B2D2F1AF1A0F802" box="[281,466,1906,1928]" italics="true" pageId="11" pageNumber="548">Gonatodes petersi</emphasis>
</taxonomicName>
. A: male; B: female. La Orchila, Sierra de Perijá, estado Zulia. Uncollected specimens. Photos: Fernando Rojas-Runjaic.
</paragraph>
</caption>
<caption id="DF146675FFE7FFA20AA32FD9F254F86F" httpUri="https://zenodo.org/record/271924/files/figure.png" pageId="12" pageNumber="549" targetBox="[285,1335,225,1908]" targetPageId="12">
<paragraph id="8BD436FDFFE7FFA20AA32FD9F254F86F" blockId="12.[151,1436,1969,2021]" pageId="12" pageNumber="549">
<emphasis id="B91FEAEFFFE7FFA20AA32FD9F160F84C" bold="true" box="[151,274,1969,1991]" pageId="12" pageNumber="549">FIGURE 7.</emphasis>
<taxonomicName id="4C6B4D7EFFE7FFA20B2C2FD9F1A2F84D" box="[280,464,1969,1991]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="549" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91FEAEFFFE7FFA20B2C2FD9F1A2F84D" box="[280,464,1969,1991]" italics="true" pageId="12" pageNumber="549">Gonatodes petersi</emphasis>
</taxonomicName>
. A: holotype, dorsal view. B: ventral view. C: Alotype dorsal view. D: alotype, ventral view. Not to scale. Photos by Juan Pablo Donoso.
</paragraph>
</caption>
<paragraph id="8BD436FDFFE6FFA30AF328FFF28AFE7F" blockId="13.[151,1437,151,1437]" pageId="13" pageNumber="550">
Scales on anterodorsal surface of forelimbs flat, smooth, rhomboidal, imbricate, and largest on arm; on posterodorsal and ventral surface of forelimbs scales are granular, rounded, much smaller than anterodorsal part. Scales on anterodorsal surface of thighs and ventral surface of hind limbs flat, smooth, irregularly rounded, imbricate and bigger than those on posterodorsal surface of thighs and dorsal surface of lower legs, which are juxtaposed, rhomboidal to triangular or rounded to the base of the ankle (some skin on the dorsal surface of the right thigh is missing; part of the left shank is damaged; the posterior portion of the dorsum is skinless. Lamellae under first (
<emphasis id="B91FEAEFFFE6FFA30B292907F15AFE02" bold="true" box="[285,296,367,392]" pageId="13" pageNumber="550">I</emphasis>
) through fifth (
<emphasis id="B91FEAEFFFE6FFA30BE82907F182FE02" bold="true" box="[476,496,367,392]" pageId="13" pageNumber="550">V</emphasis>
) finger (right/left hand; proximal lamellae in parentheses):
<emphasis id="B91FEAEFFFE6FFA30EA12907F4D2FE02" bold="true" box="[1173,1184,367,392]" pageId="13" pageNumber="550">I</emphasis>
: 12/11,
<emphasis id="B91FEAEFFFE6FFA30ECA2907F566FE02" bold="true" box="[1278,1300,367,392]" pageId="13" pageNumber="550">II</emphasis>
: 14/15,
<emphasis id="B91FEAEFFFE6FFA30F402907F5E7FE02" bold="true" box="[1396,1429,367,392]" pageId="13" pageNumber="550">III</emphasis>
: 17/17,
<emphasis id="B91FEAEFFFE6FFA30AD229FCF171FE27" bold="true" box="[230,259,404,429]" pageId="13" pageNumber="550">IV</emphasis>
: 16/17,
<emphasis id="B91FEAEFFFE6FFA30B5729FCF107FE27" bold="true" box="[355,373,404,429]" pageId="13" pageNumber="550">V</emphasis>
: 14/14. Lamellae under first (
<emphasis id="B91FEAEFFFE6FFA308F829FCF2A5FE27" bold="true" box="[716,727,404,429]" pageId="13" pageNumber="550">I</emphasis>
) through fifth (
<emphasis id="B91FEAEFFFE6FFA309BF29FCF3EDFE27" bold="true" box="[907,927,404,429]" pageId="13" pageNumber="550">V</emphasis>
) toe (right/left foot):
<emphasis id="B91FEAEFFFE6FFA30EA229FCF4D3FE27" bold="true" box="[1174,1185,404,429]" pageId="13" pageNumber="550">I</emphasis>
: 11/11,
<emphasis id="B91FEAEFFFE6FFA30ECA29FCF566FE27" bold="true" box="[1278,1300,404,429]" pageId="13" pageNumber="550">II</emphasis>
: 14/14,
<emphasis id="B91FEAEFFFE6FFA30F4029FCF5E7FE27" bold="true" box="[1396,1429,404,429]" pageId="13" pageNumber="550">III</emphasis>
: 18/18,
<emphasis id="B91FEAEFFFE6FFA30AD129DFF170FE5A" bold="true" box="[229,258,439,464]" pageId="13" pageNumber="550">IV</emphasis>
: 21/
<date id="FFD5103DFFE6FFA30B0329D0F1E4FE5A" box="[311,406,439,464]" pageId="13" pageNumber="550" value="1921-05-19">
21,
<emphasis id="B91FEAEFFFE6FFA30B6B29DFF101FE5A" bold="true" box="[351,371,439,464]" pageId="13" pageNumber="550">V</emphasis>
19
</date>
/20. Fingers and toes with two lateral rows of scales distally. Claws exposed, non-retractile, between two basal scales (one dorsal and one ventral).
</paragraph>
<paragraph id="8BD436FDFFE6FFA30AF32997F345FC2F" blockId="13.[151,1437,151,1437]" pageId="13" pageNumber="550">
<emphasis id="B91FEAEFFFE6FFA30AF32997F1BAFD92" bold="true" box="[199,456,511,536]" pageId="13" pageNumber="550">Color in preservative</emphasis>
. Dorsal color dark brown on head and anterior part of body, being paler towards the middle of the body (posterior part of skin on body missing). There is a white vertebral stripe which is quite regular, but differs in width from 4-5 scales on narrow parts to 8-9 scales on wider parts, becoming less contrasting towards the posterior part of the dorsum. There are a few small white spots symmetrically disposed on exterior side of head and neck, and three transversal short stripes crossing the vertebral stripe. Legs and hind limbs pale brown with diffuse dark mottling. Flanks dirty pale brown. Right side of head dark brown on top, with two white small spots, one immediately behind eye, and other on the lateral occipital area; a white horizontal stripe (regular on the superior margin, irregular on the inferior one, runs from behind eye to the pre-humeral region; a black irregular stripe forming part of the gular reticulation lies immediately below, including the tympanic area. Supralabials and imfralabials crossed vertically by white and dark brown stripes. Pupil black. Gular area white with irregular black reticulation, extending through chest, where reticulation becomes diffuse and the belly is immaculate white with dirty pale brown on the flanks. Palms and soles ivory white.
</paragraph>
<paragraph id="8BD436FDFFE6FFA30AF32BC7F13DFB62" blockId="13.[151,1437,151,1437]" pageId="13" pageNumber="550">
<emphasis id="B91FEAEFFFE6FFA30AF32BC7F147FC42" bold="true" box="[199,309,943,968]" pageId="13" pageNumber="550">Variation</emphasis>
. The allotype is an adult (MZUC 12105) of
<quantity id="4C939B18FFE6FFA309732BD8F3CFFC4D" box="[839,957,944,968]" metricMagnitude="-2" metricUnit="m" metricValue="3.921" pageId="13" pageNumber="550" unit="mm" value="39.21">39.21 mm</quantity>
SVL, with entire (but almost completely regenerated) tail. Dorsal coloration includes a dirty white head and neck sulcated by two dark brown irregular longitudinal stripes starting behind nostrils, passing interior to the eyelids and ending on nape; there is a white wide area between the two dark irregular stripes. There are three postocular longitudinal diffuse dark stripes. Dorsum greyish brown with dark brown diffuse markings. A dirty white diffuse vertebral stripe runs from nape to sacrum. There are two symmetrical prehumeral vertical dirty white stripes. Tail is regenerated and conserves only
<quantity id="4C939B18FFE6FFA30F722C0CF5EFFBF6" box="[1350,1437,1124,1149]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="550" unit="mm" value="2.5">2.5 mm</quantity>
of base: the regenerated tail is dorsally pale brown and becomes dirty white towards the tip (
<figureCitation id="13502A78FFE6FFA30E962CEFF497FB2A" box="[1186,1253,1159,1184]" captionStart="FIGURE 7" captionStartId="12.[151,250,1969,1991]" captionTargetBox="[285,1335,225,1908]" captionTargetId="figure@12.[248,1335,193,1927]" captionTargetPageId="12" captionText="FIGURE 7. Gonatodes petersi. A: holotype, dorsal view. B: ventral view. C: Alotype dorsal view. D: alotype, ventral view. Not to scale. Photos by Juan Pablo Donoso." httpUri="https://zenodo.org/record/271924/files/figure.png" pageId="13" pageNumber="550">Fig. 7</figureCitation>
C). Gular region dirty white with a diffuse dark irregular reticulation. Chest and belly immaculate dirty white, as is the ventral side of tail (
<figureCitation id="13502A78FFE6FFA30ADC2CA7F15EFB62" box="[232,300,1231,1256]" captionStart="FIGURE 7" captionStartId="12.[151,250,1969,1991]" captionTargetBox="[285,1335,225,1908]" captionTargetId="figure@12.[248,1335,193,1927]" captionTargetPageId="12" captionText="FIGURE 7. Gonatodes petersi. A: holotype, dorsal view. B: ventral view. C: Alotype dorsal view. D: alotype, ventral view. Not to scale. Photos by Juan Pablo Donoso." httpUri="https://zenodo.org/record/271924/files/figure.png" pageId="13" pageNumber="550">Fig. 7</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="C3716576FFE6FFA30AF32C9CF1F3FA17" pageId="13" pageNumber="550" type="discussion">
<paragraph id="8BD436FDFFE6FFA30AF32C9CF1F3FA17" blockId="13.[151,1437,151,1437]" pageId="13" pageNumber="550">
<emphasis id="B91FEAEFFFE6FFA30AF32C9CF146FA87" bold="true" box="[199,308,1268,1293]" pageId="13" pageNumber="550">Remarks</emphasis>
. The
<typeStatus id="54D0885FFFE6FFA30B422C9DF1D4FA87" box="[374,422,1269,1293]" pageId="13" pageNumber="550">type</typeStatus>
locality
<bibRefCitation id="EFFA4B0CFFE6FFA3083E2C9DF375FA87" author="Donoso-Barros" box="[522,775,1268,1293]" pageId="13" pageNumber="550" refString="Donoso-Barros, R. (1967) Diagnosis de dos nuevas especies del genero Gonatodes de Venezuela. Noticiario Mensual del Museo Nacional de Historia Natural, 129, fourth page (not numbered)." type="book chapter" year="1967">Donoso-Barros (1967)</bibRefCitation>
gives is too vague, Sierra de Perijá, as it refers to a whole mountain chain of
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long for
<quantity id="4C939B18FFE6FFA3081A2D70F204FABA" box="[558,630,1303,1328]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="550" unit="km" value="30.0">30 km</quantity>
wide. Rivero-Blanco (1979) restricts the
<typeStatus id="54D0885FFFE6FFA30E7B2D71F40DFABB" box="[1103,1151,1305,1329]" pageId="13" pageNumber="550">type</typeStatus>
locality to
<quantity id="4C939B18FFE6FFA30F352D70F53BFABA" box="[1281,1353,1303,1328]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="550" unit="km" value="10.0">10 km</quantity>
east of Misión el Tokuko. Rojas-Runjaic and Infante (2009) redescribed this species based on specimens from the whole range. Additionally,
<bibRefCitation id="EFFA4B0CFFE6FFA30BB02D37F2CDFAF2" box="[388,703,1375,1400]" pageId="13" pageNumber="550" refString="Rojas-Runjaic, FJM., Becerra, A &amp; Infante, EE. (2008) Aportes al conocimiento de la biologia reproductiva del tuqueque de Perija Gonatodes petersi Donoso-Barros, 1967 (Reptilia, Gekkonidae, Sphaerodactylinae). Saber, 20, 117 - 123." type="journal article">
Rojas-Runjaic
<emphasis id="B91FEAEFFFE6FFA3081A2D09F218FAF2" box="[558,618,1376,1400]" italics="true" pageId="13" pageNumber="550">et al.</emphasis>
(2008)
</bibRefCitation>
provide information about his natural history and reproductive biology respectively.
</paragraph>
</subSubSection>
</treatment>
</document>
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