340 lines
62 KiB
XML
340 lines
62 KiB
XML
<document id="5D90856D83BAF3B85BF54C2B269EE84B" ID-DOI="10.11646/zootaxa.4759.1.1" ID-GBIF-Dataset="f75b13f6-a898-42ff-93a4-2d177da2207f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3735855" ID-ZooBank="8C3F94C9-3923-43F4-918A-6A1CA92F1F49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585730665729" checkinUser="plazi" docAuthor="Huynh, Cuong" docDate="2020" docId="03C287E98602FFE3FF40FE575726FF25" docLanguage="en" docName="zootaxa.4759.1.1.pdf" docOrigin="Zootaxa 4759 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Unixenus moniqueae Huynh 2020, new species" docType="treatment" docVersion="5" lastPageNumber="23" masterDocId="FFFBFF918616FFF5FFD7FF8A547AFFDE" masterDocTitle="Three new species of penicillate millipedes from the Con Dao Islands of southeast Vietnam (Diplopoda, Lophoproctidae and Polyxenidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="21" updateTime="1698734097456" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="05D2953371CC6807843C9FFB655536B0">
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<mods:title id="5BD00126BD104F6134EEB40CEE95EE95">Three new species of penicillate millipedes from the Con Dao Islands of southeast Vietnam (Diplopoda, Lophoproctidae and Polyxenidae)</mods:title>
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<mods:name id="078E3387381C2EC5CCBBBE020EA33838" type="personal">
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<mods:roleTerm id="5BE63E877872993C3E33F5A030304AC9">Author</mods:roleTerm>
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<mods:namePart id="04567DB7730C3DC5D8E3D1E1015A1453">Huynh, Cuong</mods:namePart>
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<mods:typeOfResource id="53DB0A3DE0B26931FAAA3A8092E2428B">text</mods:typeOfResource>
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<mods:title id="FE1C06B25A9678A44D58D55CC0642DCE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5A79246C59B519C3968A42A8DF5EFC75">
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<mods:date id="4CABE9189A5499A8DD92F5F74DAAB216">2020</mods:date>
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<mods:number id="7E73F78617F7C0996DCEE3DDDD9E5D77">2020-04-01</mods:number>
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<mods:number id="29098B1BA88E28B97BEB8F214A7F0039">4759</mods:number>
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<mods:identifier id="BEF75A6A5EA6D179202A420227B940AD" type="DOI">10.11646/zootaxa.4759.1.1</mods:identifier>
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<mods:identifier id="B746B503CA8A6F6420D42F36180E36DC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="784D66E51C36FF801710F57B17F19C35" type="Zenodo-Dep">3735855</mods:identifier>
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<treatment id="03C287E98602FFE3FF40FE575726FF25" ID-DOI="http://doi.org/10.5281/zenodo.3811987" ID-GBIF-Taxon="163420473" ID-Zenodo-Dep="3811987" LSID="urn:lsid:plazi:treatment:03C287E98602FFE3FF40FE575726FF25" httpUri="http://treatment.plazi.org/id/03C287E98602FFE3FF40FE575726FF25" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<subSubSection id="C37165748602FFE1FF40FE57564BFE29" box="[151,561,477,503]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BD436FF8602FFE1FF40FE57564BFE29" blockId="20.[151,561,477,539]" box="[151,561,477,503]" pageId="20" pageNumber="21">
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<heading id="D09C81938602FFE1FF40FE57564BFE29" bold="true" box="[151,561,477,503]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<emphasis id="B91FEAED8602FFE1FF40FE57564BFE29" bold="true" box="[151,561,477,503]" pageId="20" pageNumber="21">
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<taxonomicName id="4C6B4D7C8602FFE1FF40FE5755E0FE29" ID-CoL="8283c484-c7f0-4822-a761-36fed8d83c93" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[151,410,477,503]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="moniqueae" status="sp. nov.">
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<emphasis id="B91FEAED8602FFE1FF40FE5755E0FE29" bold="true" box="[151,410,477,503]" italics="true" pageId="20" pageNumber="21">Unixenus moniqueae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22C57968602FFE1FE76FE57564BFE29" box="[417,561,477,503]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37165748602FFE1FF40FD8B556AFDC5" box="[151,272,513,539]" pageId="20" pageNumber="21" type="description">
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<paragraph id="8BD436FF8602FFE1FF40FD8B556AFDC5" blockId="20.[151,561,477,539]" box="[151,272,513,539]" pageId="20" pageNumber="21">
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<figureCitation id="13502A7A8602FFE1FF40FD8B556AFDC5" box="[151,272,513,539]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStart-4="FIGURE 21" captionStartId-0="21.[151,250,1643,1668]" captionStartId-1="22.[151,250,1424,1449]" captionStartId-2="23.[151,250,1613,1638]" captionStartId-3="24.[151,250,1018,1043]" captionStartId-4="25.[151,250,1818,1843]" captionTargetBox-0="[205,1369,906,1626]" captionTargetBox-1="[261,1336,285,1393]" captionTargetBox-2="[173,1413,181,1588]" captionTargetBox-3="[171,1417,181,993]" captionTargetBox-4="[171,1411,184,1789]" captionTargetId-0="figure@21.[205,1382,906,1626]" captionTargetId-1="figure@22.[240,1347,273,1399]" captionTargetId-2="figure@23.[173,1415,181,1588]" captionTargetId-3="figure@24.[171,1417,181,993]" captionTargetId-4="figure@25.[171,1416,181,1792]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionTargetPageId-4="25" captionText-0="FIGURE 17. Digital image of Unixenus moniqueae n. sp." captionText-1="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." captionText-2="FIGURE 19. SEM images of Unixenus moniqueae n. sp., A. Dorsal view; head, the posterior vertex trichomes (pv), Collum (co), tergite 2 (T2) and tergite 3 (T3); B. Ventral view; head, f: frons, cl: clypeolabrum, g: gnathochilarium; C. Trichobothria a, b and c; D. Labrum showing the posterior setae (s) and pointed spherical papillate (Psp) on its surface; E. Caudal bundle (cb) with trichomes attached, T9: tergite 9, b: trichome b and ct: caudal trichomes; F. Female caudal bundle structure without trichomes; ots: ornamental trichome sockets, b: barbate trichome, bts: barbate trichome sockets, c: trichome c, cts: caudal trichome sockets, nts: nest trichome sockets, tsft: trichome socket-free tissue." captionText-3="FIGURE 20. A. Unixenus moniqueae n. sp., Scanning electron microscope (SEM) image of antennomeres VI, VII and VIII with 4 sensory cones (sc); B. Arrangement of sensilla on the antennomeres VI and VII from the holotype. (c: conical sensillum, T: thick sensillum (a: anterior, i: intermediate, p: posterior), s: setiform sensillum)" captionText-4="FIGURE 21. A. Holotype Unixenus moniqueae n. sp., 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta with a ridged funicle present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta is present in the middle of the femur; D. Seta on tibia; E. Setiform seta on tarsus 2; F. Telotarsus—claw structures, a ventral view (SEM image) from a specimen and drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi-0="http://doi.org/10.5281/zenodo.3735891" figureDoi-1="http://doi.org/10.5281/zenodo.3735893" figureDoi-2="http://doi.org/10.5281/zenodo.3735895" figureDoi-3="http://doi.org/10.5281/zenodo.3735897" figureDoi-4="http://doi.org/10.5281/zenodo.3735899" httpUri-0="https://zenodo.org/record/3735891/files/figure.png" httpUri-1="https://zenodo.org/record/3735893/files/figure.png" httpUri-2="https://zenodo.org/record/3735895/files/figure.png" httpUri-3="https://zenodo.org/record/3735897/files/figure.png" httpUri-4="https://zenodo.org/record/3735899/files/figure.png" pageId="20" pageNumber="21">Fig. 17–21</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37165748602FFE1FF40FDC257A1FD72" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="8BD436FF8602FFE1FF40FDC257A1FD72" blockId="20.[151,1437,584,2015]" pageId="20" pageNumber="21">
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<emphasis id="B91FEAED8602FFE1FF40FDC25500FDBD" bold="true" box="[151,378,584,611]" pageId="20" pageNumber="21">Material examined</emphasis>
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: Type specimens. Adult
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<specimenCount id="9D6DFD768602FFE1FD45FDC356DDFDBD" box="[658,679,585,611]" pageId="20" pageNumber="21" type="male">♂</specimenCount>
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<typeStatus id="54D0885D8602FFE1FD78FDC3576BFDBD" box="[687,785,585,611]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED7AAE3A8602FFE1FCF7FDC35724FDBD" box="[800,862,585,611]" pageId="20" pageNumber="21">QMS</collectionCode>
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109028),;
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<specimenCount id="9D6DFD768602FFE1FC35FDC3504AFDBD" box="[994,1072,585,611]" pageId="20" pageNumber="21" type="adult">6 adult</specimenCount>
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<specimenCount id="9D6DFD768602FFE1FBE0FDC35036FDBD" box="[1079,1100,585,611]" pageId="20" pageNumber="21" type="female">♀</specimenCount>
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<typeStatus id="54D0885D8602FFE1FB83FDC350C5FDBD" box="[1108,1215,585,611]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
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(
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<collectionCode id="ED7AAE3A8602FFE1FB18FDC35177FDBD" box="[1231,1293,585,611]" pageId="20" pageNumber="21">QMS</collectionCode>
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109029–34) from
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<collectingCountry id="F37C766F8602FFE1FF03FDE7554CFD59" box="[212,310,621,647]" name="Vietnam" pageId="20" pageNumber="21">Vietnam</collectingCountry>
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, Con Dao archipelago, Con Son Island, Lo Voi beach, leaf litter in casuarina woodland,
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<geoCoordinate id="EE5F50388602FFE1FADAFDE751E2FD59" box="[1293,1432,621,647]" degrees="8.691097" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="8.691097">8.691097°N</geoCoordinate>
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,
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<geoCoordinate id="EE5F50388602FFE1FF40FD1A5541FD75" box="[151,315,656,683]" degrees="106.620542" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="106.620544">106.620542°E</geoCoordinate>
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, elevation
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<quantity id="4C939B1A8602FFE1FE63FD1B5598FD75" box="[436,482,657,683]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="21" unit="m" value="9.0">9 m</quantity>
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; on
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<date id="FFD5103F8602FFE1FDC6FD1B56CDFD75" box="[529,695,656,683]" pageId="20" pageNumber="21" value="2015-07-24">24th July 2015</date>
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(Collected by C. Huynh).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37165748602FFE1FF10FD3E51EFFCE5" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="8BD436FF8602FFE1FF10FD3E51EFFCE5" blockId="20.[151,1437,584,2015]" pageId="20" pageNumber="21">
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<emphasis id="B91FEAED8602FFE1FF10FD3E5538FD11" bold="true" box="[199,322,692,719]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
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Shares similar characteristics with
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<taxonomicName id="4C6B4D7C8602FFE1FD0FFD3C5717FD11" authorityName="Verhoeff" authorityYear="1924" box="[728,877,693,719]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mjoebergi">
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<emphasis id="B91FEAED8602FFE1FD0FFD3C5717FD11" box="[728,877,693,719]" italics="true" pageId="20" pageNumber="21">U. mjoebergi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6B4D7C8602FFE1FC70FD3C5002FD11" authorityName="Huynh and Veenstra" authorityYear="2018" box="[935,1144,694,719]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="intragramineus">
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<emphasis id="B91FEAED8602FFE1FC70FD3C5002FD11" box="[935,1144,694,719]" italics="true" pageId="20" pageNumber="21">U. intragramineus</emphasis>
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</taxonomicName>
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: chaetotaxy (setae on leg articles) with one seta on pre-femur and two setae on femur, single seta on tibia and tarsus 2. Claw more similar to
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<taxonomicName id="4C6B4D7C8602FFE1FF40FD745512FCC9" authorityName="Huynh and Veenstra" authorityYear="2018" box="[151,360,766,791]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="intragramineus">
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<emphasis id="B91FEAED8602FFE1FF40FD745512FCC9" box="[151,360,766,791]" italics="true" pageId="20" pageNumber="21">U. intragramineus</emphasis>
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</taxonomicName>
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in being slender, differs from
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<taxonomicName id="4C6B4D7C8602FFE1FD13FD74572DFCC9" authorityName="Verhoeff" authorityYear="1924" box="[708,855,765,791]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mjoebergi">
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<emphasis id="B91FEAED8602FFE1FD13FD74572DFCC9" box="[708,855,765,791]" italics="true" pageId="20" pageNumber="21">U. mjoebergi</emphasis>
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</taxonomicName>
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. A posterior process shorter than half length of the claw in
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<taxonomicName id="4C6B4D7C8602FFE1FF27FCA855F7FCE5" authorityName="Huynh" authorityYear="2020" box="[240,397,802,827]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="moniqueae">
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<emphasis id="B91FEAED8602FFE1FF27FCA855F7FCE5" box="[240,397,802,827]" italics="true" pageId="20" pageNumber="21">U. moniqueae</emphasis>
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</taxonomicName>
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compared to
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||
<taxonomicName id="4C6B4D7C8602FFE1FDF1FCA8568FFCE5" authorityName="Huynh and Veenstra" authorityYear="2018" box="[550,757,802,827]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="intragramineus">
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||
<emphasis id="B91FEAED8602FFE1FDF1FCA8568FFCE5" box="[550,757,802,827]" italics="true" pageId="20" pageNumber="21">U. intragramineus</emphasis>
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</taxonomicName>
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with a posterior process longer than half length of the claw.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37165748602FFE0FF10FCCE57F6FC5D" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="8BD436FF8602FFE0FF10FCCE57F6FC5D" blockId="20.[151,1437,584,2015]" lastBlockId="21.[151,1437,153,899]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
|
||
<emphasis id="B91FEAED8602FFE1FF10FCCE552BFC81" bold="true" box="[199,337,836,863]" pageId="20" pageNumber="21">Description</emphasis>
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: Colour dark brown patch in eye area, same colour as transverse band in vertex. Body milky white, light brown pleural trichomes and darker colour in caudal bundle; dark brown marks on lateroposterior rosette trichomes form a dark band along each lateral side of body. Last tergite darkest in colour; white ventrally (
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<figureCitation id="13502A7A8602FFE1FAEDFC0751F5FC79" box="[1338,1423,909,935]" captionStart="FIGURE 17" captionStartId="21.[151,250,1643,1668]" captionTargetBox="[205,1369,906,1626]" captionTargetId="figure@21.[205,1382,906,1626]" captionTargetPageId="21" captionText="FIGURE 17. Digital image of Unixenus moniqueae n. sp." figureDoi="http://doi.org/10.5281/zenodo.3735891" httpUri="https://zenodo.org/record/3735891/files/figure.png" pageId="20" pageNumber="21">Fig. 17</figureCitation>
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).
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<typeStatus id="54D0885D8602FFE1FF40FC3B5578FC15" box="[151,258,945,971]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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, body length
|
||
<quantity id="4C939B1A8602FFE1FE4AFC3B558DFC15" box="[413,503,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="21" unit="mm" value="2.2">2.2 mm</quantity>
|
||
;
|
||
<typeStatus id="54D0885D8602FFE1FDD4FC3B5614FC15" box="[515,622,945,971]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
|
||
<quantity id="4C939B1A8602FFE1FDA0FC3B5778FC15" box="[631,770,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="20" pageNumber="21" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
|
||
. Male caudal bundle slightly narrower in width and longer (
|
||
<quantity id="4C939B1A8602FFE1FF48FC5F548DFC31" box="[159,247,981,1007]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="mm" value="0.6">0.6 mm</quantity>
|
||
) compared to female (
|
||
<quantity id="4C939B1A8602FFE1FE25FC5E5636FC31" box="[498,588,980,1007]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="mm" value="0.5">0.5 mm</quantity>
|
||
).
|
||
<emphasis id="B91FEAED8602FFE1FD88FC5F56E5FC31" bold="true" box="[607,671,981,1007]" pageId="20" pageNumber="21">Head</emphasis>
|
||
: Each side with 8 ommatidia: 4 dorsal, 4 lateral (1 anterior, 2 medial and 1 posterior). Vertex with 2 posterior trichome groups with a large medial gap, each group with 2 rows. Anterior row curved slightly on an oblique angle, all trichome sockets of similar size, posterior row with varied trichome socket sizes. A narrow space between anterior and posterior rows.
|
||
<typeStatus id="54D0885D8602FFE1FC53FBCB5796FB85" box="[900,1004,1089,1115]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||
with vertex posterior trichome groups with 14+12 sockets in anterior rows and 8+6 sockets in posterior rows (
|
||
<figureCitation id="13502A7A8602FFE1FC6BFBEF505BFBA1" box="[956,1057,1125,1151]" captionStart="FIGURE 18" captionStartId="22.[151,250,1424,1449]" captionTargetBox="[261,1336,285,1393]" captionTargetId="figure@22.[240,1347,273,1399]" captionTargetPageId="22" captionText="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." figureDoi="http://doi.org/10.5281/zenodo.3735893" httpUri="https://zenodo.org/record/3735893/files/figure.png" pageId="20" pageNumber="21">Fig. 18A</figureCitation>
|
||
) (Variations are common in
|
||
<typeStatus id="54D0885D8602FFE1FAB4FBEF54A8FB7D" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
|
||
regardless of sex, from 12–18 sockets (anterior rows) and 6–10 (posterior rows)) (
|
||
<figureCitation id="13502A7A8602FFE1FBA6FB0350A3FB7D" box="[1137,1241,1161,1187]" captionStart="FIGURE 19" captionStartId="23.[151,250,1613,1638]" captionTargetBox="[173,1413,181,1588]" captionTargetId="figure@23.[173,1415,181,1588]" captionTargetPageId="23" captionText="FIGURE 19. SEM images of Unixenus moniqueae n. sp., A. Dorsal view; head, the posterior vertex trichomes (pv), Collum (co), tergite 2 (T2) and tergite 3 (T3); B. Ventral view; head, f: frons, cl: clypeolabrum, g: gnathochilarium; C. Trichobothria a, b and c; D. Labrum showing the posterior setae (s) and pointed spherical papillate (Psp) on its surface; E. Caudal bundle (cb) with trichomes attached, T9: tergite 9, b: trichome b and ct: caudal trichomes; F. Female caudal bundle structure without trichomes; ots: ornamental trichome sockets, b: barbate trichome, bts: barbate trichome sockets, c: trichome c, cts: caudal trichome sockets, nts: nest trichome sockets, tsft: trichome socket-free tissue." figureDoi="http://doi.org/10.5281/zenodo.3735895" httpUri="https://zenodo.org/record/3735895/files/figure.png" pageId="20" pageNumber="21">Fig. 19A</figureCitation>
|
||
).
|
||
<emphasis id="B91FEAED8602FFE1FB3AFB0251EEFB7D" bold="true" box="[1261,1428,1160,1187]" pageId="20" pageNumber="21">Trichobothria</emphasis>
|
||
: trichobothrium
|
||
<emphasis id="B91FEAED8602FFE1FE91FB27552EFB19" bold="true" box="[326,340,1197,1223]" italics="true" pageId="20" pageNumber="21">a</emphasis>
|
||
(posterior position), trichobothrium
|
||
<emphasis id="B91FEAED8602FFE1FD38FB265687FB19" bold="true" box="[751,765,1196,1223]" italics="true" pageId="20" pageNumber="21">b</emphasis>
|
||
(lateral position) and trichobothrium
|
||
<emphasis id="B91FEAED8602FFE1FB69FB2750B0FB19" bold="true" box="[1214,1226,1197,1223]" italics="true" pageId="20" pageNumber="21">c</emphasis>
|
||
(anterior position) typically thin sensory hairs with narrow cylindrical funicles. Trichobothria (
|
||
<emphasis id="B91FEAED8602FFE1FC26FB5B5785FB35" bold="true" box="[1009,1023,1233,1259]" italics="true" pageId="20" pageNumber="21">a</emphasis>
|
||
,
|
||
<emphasis id="B91FEAED8602FFE1FBDAFB5A5061FB35" bold="true" box="[1037,1051,1232,1259]" italics="true" pageId="20" pageNumber="21">b</emphasis>
|
||
and
|
||
<emphasis id="B91FEAED8602FFE1FB84FB5B5025FB35" bold="true" box="[1107,1119,1233,1259]" italics="true" pageId="20" pageNumber="21">c</emphasis>
|
||
) equal in socket size, forming an isosceles triangle with equal distance between
|
||
<emphasis id="B91FEAED8602FFE1FD38FB7E5771FAD1" bold="true" box="[751,779,1268,1295]" italics="true" pageId="20" pageNumber="21">ab</emphasis>
|
||
and
|
||
<emphasis id="B91FEAED8602FFE1FC95FB7E5726FAD1" bold="true" box="[834,860,1268,1295]" italics="true" pageId="20" pageNumber="21">bc</emphasis>
|
||
(
|
||
<figureCitation id="13502A7A8602FFE1FCBBFB7F57AAFAD1" box="[876,976,1269,1295]" captionStart="FIGURE 18" captionStartId="22.[151,250,1424,1449]" captionTargetBox="[261,1336,285,1393]" captionTargetId="figure@22.[240,1347,273,1399]" captionTargetPageId="22" captionText="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." figureDoi="http://doi.org/10.5281/zenodo.3735893" httpUri="https://zenodo.org/record/3735893/files/figure.png" pageId="20" pageNumber="21">Figs 18F</figureCitation>
|
||
and
|
||
<figureCitation id="13502A7A8602FFE1FBD0FB7F5043FAD1" box="[1031,1081,1269,1295]" captionStart="FIGURE 19" captionStartId="23.[151,250,1613,1638]" captionTargetBox="[173,1413,181,1588]" captionTargetId="figure@23.[173,1415,181,1588]" captionTargetPageId="23" captionText="FIGURE 19. SEM images of Unixenus moniqueae n. sp., A. Dorsal view; head, the posterior vertex trichomes (pv), Collum (co), tergite 2 (T2) and tergite 3 (T3); B. Ventral view; head, f: frons, cl: clypeolabrum, g: gnathochilarium; C. Trichobothria a, b and c; D. Labrum showing the posterior setae (s) and pointed spherical papillate (Psp) on its surface; E. Caudal bundle (cb) with trichomes attached, T9: tergite 9, b: trichome b and ct: caudal trichomes; F. Female caudal bundle structure without trichomes; ots: ornamental trichome sockets, b: barbate trichome, bts: barbate trichome sockets, c: trichome c, cts: caudal trichome sockets, nts: nest trichome sockets, tsft: trichome socket-free tissue." figureDoi="http://doi.org/10.5281/zenodo.3735895" httpUri="https://zenodo.org/record/3735895/files/figure.png" pageId="20" pageNumber="21">19C</figureCitation>
|
||
).
|
||
<emphasis id="B91FEAED8602FFE1FB9AFB7E50BAFAD1" bold="true" box="[1101,1216,1268,1295]" pageId="20" pageNumber="21">Antennae</emphasis>
|
||
: 8 antennomeres, 4 sensory cones (
|
||
<figureCitation id="13502A7A8602FFE1FE93FA9355D7FAED" box="[324,429,1305,1331]" captionStart="FIGURE 20" captionStartId="24.[151,250,1018,1043]" captionTargetBox="[171,1417,181,993]" captionTargetId="figure@24.[171,1417,181,993]" captionTargetPageId="24" captionText="FIGURE 20. A. Unixenus moniqueae n. sp., Scanning electron microscope (SEM) image of antennomeres VI, VII and VIII with 4 sensory cones (sc); B. Arrangement of sensilla on the antennomeres VI and VII from the holotype. (c: conical sensillum, T: thick sensillum (a: anterior, i: intermediate, p: posterior), s: setiform sensillum)" figureDoi="http://doi.org/10.5281/zenodo.3735897" httpUri="https://zenodo.org/record/3735897/files/figure.png" pageId="20" pageNumber="21">Fig. 20A</figureCitation>
|
||
), typical characteristics of
|
||
<taxonomicName id="4C6B4D7C8602FFE1FD0CFA935713FAED" authorityName="Lucas" authorityYear="1840" box="[731,873,1305,1331]" class="Diplopoda" family="Polyxenidae" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Polyxenidae</taxonomicName>
|
||
. Antennomere VI with 3 thick bacilliform sensilla (T), different lengths: A short bacilliform sensillum located in the anterior position (Ta), the longest intermediate long bacilliform sensillum (Ti), and a bacilliform sensillum of (Tp) medium length in posterior position. A setiform sensillum (s) between Ta and Ti and conical sensillum (c) in next to Tp. Antennomere VII with 2 thick bacilliform sensilla, Ta slightly longer than Tp; a setiform sensillum (s) between them and a conical sensillum (c) located next to Tp (
|
||
<figureCitation id="13502A7A8602FFE1FF12FA475551FA39" box="[197,299,1485,1511]" captionStart="FIGURE 20" captionStartId="24.[151,250,1018,1043]" captionTargetBox="[171,1417,181,993]" captionTargetId="figure@24.[171,1417,181,993]" captionTargetPageId="24" captionText="FIGURE 20. A. Unixenus moniqueae n. sp., Scanning electron microscope (SEM) image of antennomeres VI, VII and VIII with 4 sensory cones (sc); B. Arrangement of sensilla on the antennomeres VI and VII from the holotype. (c: conical sensillum, T: thick sensillum (a: anterior, i: intermediate, p: posterior), s: setiform sensillum)" figureDoi="http://doi.org/10.5281/zenodo.3735897" httpUri="https://zenodo.org/record/3735897/files/figure.png" pageId="20" pageNumber="21">Fig. 20B</figureCitation>
|
||
). (This pattern of sensilla on the antennomere VII is commonly seen in all
|
||
<taxonomicName id="4C6B4D7C8602FFE1FBB0FA4450ABFA39" authorityName="Jones" authorityYear="1944" box="[1127,1233,1486,1511]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAED8602FFE1FBB0FA4450ABFA39" box="[1127,1233,1486,1511]" italics="true" pageId="20" pageNumber="21">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
species).
|
||
<emphasis id="B91FEAED8602FFE1FAE8FA46548AF9D5" bold="true" pageId="20" pageNumber="21">Clypeolabrum</emphasis>
|
||
:
|
||
<typeStatus id="54D0885D8602FFE1FED7FA7B5512F9D5" box="[256,360,1521,1547]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||
with 10 setae along labrum posterior margin, these setae being half the width of labrum (setae in
|
||
<typeStatus id="54D0885D8602FFE1FF62F99F555AF9F1" box="[181,288,1557,1583]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
|
||
ranging from
|
||
<quantity id="4C939B1A8602FFE1FE16F99F565BF9F1" box="[449,545,1557,1583]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.302" metricValueMin="2.54" pageId="20" pageNumber="21" unit="in" value="11.5" valueMax="13.0" valueMin="10.0">10–13 in</quantity>
|
||
both sexes). Labral surface with spherical papillae with pointed ends, forming 3 rows of large size setae in anterior margin and reducing in size in posterior margin. Anterior margin of labrum with 4+4 lamellae on each side of median cleft (
|
||
<figureCitation id="13502A7A8602FFE1FD7BF9D7576FF9A9" box="[684,789,1629,1655]" captionStart="FIGURE 18" captionStartId="22.[151,250,1424,1449]" captionTargetBox="[261,1336,285,1393]" captionTargetId="figure@22.[240,1347,273,1399]" captionTargetPageId="22" captionText="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." figureDoi="http://doi.org/10.5281/zenodo.3735893" httpUri="https://zenodo.org/record/3735893/files/figure.png" pageId="20" pageNumber="21">Figs.18G</figureCitation>
|
||
and
|
||
<figureCitation id="13502A7A8602FFE1FC9EF9D757E7F9A9" box="[841,925,1629,1655]" captionStart="FIGURE 19" captionStartId="23.[151,250,1613,1638]" captionTargetBox="[173,1413,181,1588]" captionTargetId="figure@23.[173,1415,181,1588]" captionTargetPageId="23" captionText="FIGURE 19. SEM images of Unixenus moniqueae n. sp., A. Dorsal view; head, the posterior vertex trichomes (pv), Collum (co), tergite 2 (T2) and tergite 3 (T3); B. Ventral view; head, f: frons, cl: clypeolabrum, g: gnathochilarium; C. Trichobothria a, b and c; D. Labrum showing the posterior setae (s) and pointed spherical papillate (Psp) on its surface; E. Caudal bundle (cb) with trichomes attached, T9: tergite 9, b: trichome b and ct: caudal trichomes; F. Female caudal bundle structure without trichomes; ots: ornamental trichome sockets, b: barbate trichome, bts: barbate trichome sockets, c: trichome c, cts: caudal trichome sockets, nts: nest trichome sockets, tsft: trichome socket-free tissue." figureDoi="http://doi.org/10.5281/zenodo.3735895" httpUri="https://zenodo.org/record/3735895/files/figure.png" pageId="20" pageNumber="21">19B, D</figureCitation>
|
||
).
|
||
<emphasis id="B91FEAED8602FFE1FC67F9D65005F9A9" bold="true" box="[944,1151,1628,1655]" pageId="20" pageNumber="21">Gnathochilarium</emphasis>
|
||
: Lateral palp 2.5 times as long as medial palp. 13 conical sensilla on lateral palp, medial palp with 22, same in
|
||
<typeStatus id="54D0885D8602FFE1FB8DF90B50C6F945" box="[1114,1212,1665,1691]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54D0885D8602FFE1FB23F90B5125F945" box="[1268,1375,1665,1691]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="13502A7A8602FFE1FAB8F90B54B1F961" captionStart="FIGURE 18" captionStartId="22.[151,250,1424,1449]" captionTargetBox="[261,1336,285,1393]" captionTargetId="figure@22.[240,1347,273,1399]" captionTargetPageId="22" captionText="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." figureDoi="http://doi.org/10.5281/zenodo.3735893" httpUri="https://zenodo.org/record/3735893/files/figure.png" pageId="20" pageNumber="21">Fig. 18H</figureCitation>
|
||
).
|
||
<emphasis id="B91FEAED8602FFE1FF09F92F5550F961" bold="true" box="[222,298,1701,1727]" pageId="20" pageNumber="21">Trunk</emphasis>
|
||
: 10 tergites, 9 pleural projections, and telson excluding caudal bundle.13 pairs of legs. Collum (tergite 1) with trichome sockets arranged in 2 oval shapes in lateral position, opposite each other connected by a posterior row of trichome sockets forming a line with a gap in the middle.
|
||
<typeStatus id="54D0885D8602FFE1FCA8F967579DF8D9" box="[895,999,1773,1799]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||
collum with 49 trichome sockets both sides, lateral protuberances with 5 trichome sockets on each side (
|
||
<figureCitation id="13502A7A8602FFE1FCA3F89B57ABF8F2" box="[884,977,1809,1836]" captionStart="FIGURE 18" captionStartId="22.[151,250,1424,1449]" captionTargetBox="[261,1336,285,1393]" captionTargetId="figure@22.[240,1347,273,1399]" captionTargetPageId="22" captionText="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." figureDoi="http://doi.org/10.5281/zenodo.3735893" httpUri="https://zenodo.org/record/3735893/files/figure.png" pageId="20" pageNumber="21">Fig.18B</figureCitation>
|
||
) (Trichome sockets on collum varying in
|
||
<typeStatus id="54D0885D8602FFE1FF40F8BF5578F891" box="[151,258,1845,1871]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
|
||
ranging from 54–60 sockets and 4–5 sockets on lateral protuberances). Tergites 2–10, each with pleural projections located in anterolateral positions. Tergal trichome socket arrangements from tergites 2–9 typically with 4 undefined rows of tergal trichomes arranged in two lateroposterior clusters, on either side of midline. Anterior row often uneven, intermediate rows rarely in defined rows and posterior row of trichomes continuous and evenly distributed, forming a line with a medial gap (
|
||
<figureCitation id="13502A7A8602FFE1FD73F84F5777F801" box="[676,781,1989,2015]" captionStart="FIGURE 19" captionStartId="23.[151,250,1613,1638]" captionTargetBox="[173,1413,181,1588]" captionTargetId="figure@23.[173,1415,181,1588]" captionTargetPageId="23" captionText="FIGURE 19. SEM images of Unixenus moniqueae n. sp., A. Dorsal view; head, the posterior vertex trichomes (pv), Collum (co), tergite 2 (T2) and tergite 3 (T3); B. Ventral view; head, f: frons, cl: clypeolabrum, g: gnathochilarium; C. Trichobothria a, b and c; D. Labrum showing the posterior setae (s) and pointed spherical papillate (Psp) on its surface; E. Caudal bundle (cb) with trichomes attached, T9: tergite 9, b: trichome b and ct: caudal trichomes; F. Female caudal bundle structure without trichomes; ots: ornamental trichome sockets, b: barbate trichome, bts: barbate trichome sockets, c: trichome c, cts: caudal trichome sockets, nts: nest trichome sockets, tsft: trichome socket-free tissue." figureDoi="http://doi.org/10.5281/zenodo.3735895" httpUri="https://zenodo.org/record/3735895/files/figure.png" pageId="20" pageNumber="21">Fig. 19A</figureCitation>
|
||
). Tergite 10 is the exception, with trichome sockets being smaller and denser. Almost no space between lateral rosette trichome sockets and those in posterior row. Trichome sockets of tergite
|
||
<quantity id="4C939B1A8603FFE0FEB6FF3755F4FF09" box="[353,398,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="21" pageNumber="22" unit="in" value="2.0">2 in</quantity>
|
||
<typeStatus id="54D0885D8603FFE0FE41FF375582FF09" box="[406,504,189,215]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
with 55 on both sides (
|
||
<figureCitation id="13502A7A8603FFE0FCD1FF375714FF09" box="[774,878,189,215]" captionStart="FIGURE 18" captionStartId="22.[151,250,1424,1449]" captionTargetBox="[261,1336,285,1393]" captionTargetId="figure@22.[240,1347,273,1399]" captionTargetPageId="22" captionText="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." figureDoi="http://doi.org/10.5281/zenodo.3735893" httpUri="https://zenodo.org/record/3735893/files/figure.png" pageId="21" pageNumber="22">Fig. 18C</figureCitation>
|
||
). Tergite 10 with 53 (L) and 54 (R) sockets (
|
||
<figureCitation id="13502A7A8603FFE0FAB8FF3754B2FF25" captionStart="FIGURE 18" captionStartId="22.[151,250,1424,1449]" captionTargetBox="[261,1336,285,1393]" captionTargetId="figure@22.[240,1347,273,1399]" captionTargetPageId="22" captionText="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." figureDoi="http://doi.org/10.5281/zenodo.3735893" httpUri="https://zenodo.org/record/3735893/files/figure.png" pageId="21" pageNumber="22">Fig. 18D</figureCitation>
|
||
) (In contrast, trichome sockets on tergite
|
||
<quantity id="4C939B1A8603FFE0FD76FF6B56B7FF25" box="[673,717,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="21" pageNumber="22" unit="in" value="2.0">2 in</quantity>
|
||
<typeStatus id="54D0885D8603FFE0FD02FF6B573AFF25" box="[725,832,225,251]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
|
||
50–65, whereas tergite 10 containing between 54–68 trichome sockets).
|
||
<emphasis id="B91FEAED8603FFE0FEBEFE8F55DBFEC1" bold="true" box="[361,417,261,287]" pageId="21" pageNumber="22">Legs</emphasis>
|
||
: Leg segmentation following
|
||
<bibRefCitation id="EFFA4B0E8603FFE0FD39FE8F57E3FEC1" author="Manton, S. M." box="[750,921,260,287]" pageId="21" pageNumber="22" pagination="153 - 187" refId="ref15174" refString="Manton, S. M. (1956) The Evolution of Arthropodan Locomotory Mechanisms-Part 5: The Structure, Habits and Evolution of the Pselaphognatha (Diplopoda). Journal of the Linnean Society London, 43, 153 - 187. https: // doi. org / 10.1111 / j. 1096 - 3642.1957. tb 02516. x" type="journal article" year="1956">Manton (1956)</bibRefCitation>
|
||
. Legs 1 and 2 without trochanter, leg 1 lacking tarsus 1. Chaetotaxy (setae on leg articles): coxa 1: 1 seta, coxa 2: 2 setae, coxae 3–13: 2–6 setae; pre-femur, tibia and tarsus 2 with seta; femur with 2 setae (
|
||
<figureCitation id="13502A7A8603FFE0FD56FEC75690FEB9" box="[641,746,333,359]" captionStart="FIGURE 21" captionStartId="25.[151,250,1818,1843]" captionTargetBox="[171,1411,184,1789]" captionTargetId="figure@25.[171,1416,181,1792]" captionTargetPageId="25" captionText="FIGURE 21. A. Holotype Unixenus moniqueae n. sp., 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta with a ridged funicle present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta is present in the middle of the femur; D. Seta on tibia; E. Setiform seta on tarsus 2; F. Telotarsus—claw structures, a ventral view (SEM image) from a specimen and drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735899" httpUri="https://zenodo.org/record/3735899/files/figure.png" pageId="21" pageNumber="22">Fig. 21A</figureCitation>
|
||
). Setae on coxa, pre-femur, femur distally with with longitudinal ridges on basal funicle; each ridge extending distally into long thin spine, spines surrounding flagellum base (
|
||
<figureCitation id="13502A7A8603FFE0FF77FE1F557CFE71" box="[160,262,405,431]" captionStart="FIGURE 21" captionStartId="25.[151,250,1818,1843]" captionTargetBox="[171,1411,184,1789]" captionTargetId="figure@25.[171,1416,181,1792]" captionTargetPageId="25" captionText="FIGURE 21. A. Holotype Unixenus moniqueae n. sp., 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta with a ridged funicle present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta is present in the middle of the femur; D. Seta on tibia; E. Setiform seta on tarsus 2; F. Telotarsus—claw structures, a ventral view (SEM image) from a specimen and drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735899" httpUri="https://zenodo.org/record/3735899/files/figure.png" pageId="21" pageNumber="22">Fig. 21B</figureCitation>
|
||
). Seta in mid femur similar, but smaller (
|
||
<figureCitation id="13502A7A8603FFE0FD02FE1F5741FE71" box="[725,827,405,431]" captionStart="FIGURE 21" captionStartId="25.[151,250,1818,1843]" captionTargetBox="[171,1411,184,1789]" captionTargetId="figure@25.[171,1416,181,1792]" captionTargetPageId="25" captionText="FIGURE 21. A. Holotype Unixenus moniqueae n. sp., 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta with a ridged funicle present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta is present in the middle of the femur; D. Seta on tibia; E. Setiform seta on tarsus 2; F. Telotarsus—claw structures, a ventral view (SEM image) from a specimen and drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735899" httpUri="https://zenodo.org/record/3735899/files/figure.png" pageId="21" pageNumber="22">Fig. 21C</figureCitation>
|
||
), tibia and tarsus 2 each with setiform seta (
|
||
<figureCitation id="13502A7A8603FFE0FAFBFE1F54A7FE0D" captionStart="FIGURE 21" captionStartId="25.[151,250,1818,1843]" captionTargetBox="[171,1411,184,1789]" captionTargetId="figure@25.[171,1416,181,1792]" captionTargetPageId="25" captionText="FIGURE 21. A. Holotype Unixenus moniqueae n. sp., 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta with a ridged funicle present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta is present in the middle of the femur; D. Seta on tibia; E. Setiform seta on tarsus 2; F. Telotarsus—claw structures, a ventral view (SEM image) from a specimen and drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735899" httpUri="https://zenodo.org/record/3735899/files/figure.png" pageId="21" pageNumber="22">Figs. 21D and E</figureCitation>
|
||
). Posterior edge of last sternite with 4 setae similar those present on coxa (Number of these setae same in
|
||
<typeStatus id="54D0885D8603FFE0FF40FE575572FE29" box="[151,264,477,503]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
|
||
).
|
||
<emphasis id="B91FEAED8603FFE0FECDFE565581FE29" bold="true" box="[282,507,476,503]" pageId="21" pageNumber="22">Sex organs in male</emphasis>
|
||
: pair of penes present on coxa 2, 2 pairs of coxal glands on coxal plates of legs 8th and 9th.
|
||
<emphasis id="B91FEAED8603FFE0FF20FD8A55C0FDC5" bold="true" box="[247,442,512,539]" pageId="21" pageNumber="22">Telotarsus-Claw</emphasis>
|
||
: A slender structure bearing a posterior lateral process that is shorter by half the length of claw. A small anterior lateral process and a lamella process both present, anterior setiform process slightly enlarged at base and longer than the claw (
|
||
<figureCitation id="13502A7A8603FFE0FDCEFDC35604FDBD" box="[537,638,585,611]" captionStart="FIGURE 21" captionStartId="25.[151,250,1818,1843]" captionTargetBox="[171,1411,184,1789]" captionTargetId="figure@25.[171,1416,181,1792]" captionTargetPageId="25" captionText="FIGURE 21. A. Holotype Unixenus moniqueae n. sp., 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta with a ridged funicle present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta is present in the middle of the femur; D. Seta on tibia; E. Setiform seta on tarsus 2; F. Telotarsus—claw structures, a ventral view (SEM image) from a specimen and drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735899" httpUri="https://zenodo.org/record/3735899/files/figure.png" pageId="21" pageNumber="22">Fig. 21F</figureCitation>
|
||
).
|
||
<emphasis id="B91FEAED8603FFE0FD44FDC3569AFDBD" bold="true" box="[659,736,585,611]" pageId="21" pageNumber="22">Telson</emphasis>
|
||
: Dorsal ornamental trichome sockets arranged symmetrically on both side of telson with 5 trichomes
|
||
<emphasis id="B91FEAED8603FFE0FD8FFDE7561CFD59" bold="true" box="[600,614,621,647]" italics="true" pageId="21" pageNumber="22">a</emphasis>
|
||
sockets in
|
||
<typeStatus id="54D0885D8603FFE0FD31FDE75732FD59" box="[742,840,621,647]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
(
|
||
<typeStatus id="54D0885D8603FFE0FC80FDE757BCFD59" box="[855,966,621,647]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
|
||
with 5–6 trichomes
|
||
<emphasis id="B91FEAED8603FFE0FB78FDE750C7FD59" bold="true" box="[1199,1213,621,647]" italics="true" pageId="21" pageNumber="22">a</emphasis>
|
||
). A single trichome
|
||
<emphasis id="B91FEAED8603FFE0FF40FD1A54DFFD75" bold="true" box="[151,165,656,683]" italics="true" pageId="21" pageNumber="22">b</emphasis>
|
||
socket and 3 trichome
|
||
<emphasis id="B91FEAED8603FFE0FE7FFD1B55CEFD75" bold="true" box="[424,436,657,683]" italics="true" pageId="21" pageNumber="22">c</emphasis>
|
||
with large protruding base sockets:
|
||
<emphasis id="B91FEAED8603FFE0FC9FFD1B572EFD75" bold="true" box="[840,852,657,683]" italics="true" pageId="21" pageNumber="22">c</emphasis>
|
||
1,
|
||
<emphasis id="B91FEAED8603FFE0FCA6FD1B5707FD75" bold="true" box="[881,893,657,683]" italics="true" pageId="21" pageNumber="22">c</emphasis>
|
||
2 and
|
||
<emphasis id="B91FEAED8603FFE0FC16FD1B57B7FD75" bold="true" box="[961,973,657,683]" italics="true" pageId="21" pageNumber="22">c</emphasis>
|
||
3, forming a triangular shape each side of telson. Circular indentation
|
||
<emphasis id="B91FEAED8603FFE0FE07FD3E55A4FD11" bold="true" box="[464,478,692,719]" italics="true" pageId="21" pageNumber="22">d</emphasis>
|
||
apparent near exterior side of trichomes
|
||
<emphasis id="B91FEAED8603FFE0FC7CFD3F57CDFD11" bold="true" box="[939,951,693,719]" italics="true" pageId="21" pageNumber="22">c</emphasis>
|
||
(
|
||
<figureCitation id="13502A7A8603FFE0FC10FD3F5057FD11" box="[967,1069,693,719]" captionStart="FIGURE 18" captionStartId="22.[151,250,1424,1449]" captionTargetBox="[261,1336,285,1393]" captionTargetId="figure@22.[240,1347,273,1399]" captionTargetPageId="22" captionText="FIGURE 18. Holotype Unixenus moniqueae n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. Tergite 2 (T2); D. Tergite 10—the last tergite (T10); E. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d); F. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; G. Clypeo-labrum showing the pointed spherical papillate on its surface and the structures 4 + 4 lamellae; H. Gnathochilarium: lateral palp (LP) with 13 sensilla (a sensillum number 3 missing) and medial palp (MP) with 22 sensilla." figureDoi="http://doi.org/10.5281/zenodo.3735893" httpUri="https://zenodo.org/record/3735893/files/figure.png" pageId="21" pageNumber="22">Fig. 18E</figureCitation>
|
||
).
|
||
<emphasis id="B91FEAED8603FFE0FB96FD3E5086FD11" bold="true" box="[1089,1276,692,719]" pageId="21" pageNumber="22">Caudal bundles</emphasis>
|
||
: Male, caudal bundle with a single uniform large trichome socket of caudal trichomes. Female with two obvious distinguishing structures: a main dorsal structure similar to male, but with 2 latero-sternal bundles of smaller trichome sockets of nest trichomes (
|
||
<figureCitation id="13502A7A8603FFE0FE9DFCAB55A3FCE5" box="[330,473,801,827]" captionStart="FIGURE 19" captionStartId="23.[151,250,1613,1638]" captionTargetBox="[173,1413,181,1588]" captionTargetId="figure@23.[173,1415,181,1588]" captionTargetPageId="23" captionText="FIGURE 19. SEM images of Unixenus moniqueae n. sp., A. Dorsal view; head, the posterior vertex trichomes (pv), Collum (co), tergite 2 (T2) and tergite 3 (T3); B. Ventral view; head, f: frons, cl: clypeolabrum, g: gnathochilarium; C. Trichobothria a, b and c; D. Labrum showing the posterior setae (s) and pointed spherical papillate (Psp) on its surface; E. Caudal bundle (cb) with trichomes attached, T9: tergite 9, b: trichome b and ct: caudal trichomes; F. Female caudal bundle structure without trichomes; ots: ornamental trichome sockets, b: barbate trichome, bts: barbate trichome sockets, c: trichome c, cts: caudal trichome sockets, nts: nest trichome sockets, tsft: trichome socket-free tissue." figureDoi="http://doi.org/10.5281/zenodo.3735895" httpUri="https://zenodo.org/record/3735895/files/figure.png" pageId="21" pageNumber="22">Figs. 19E, F</figureCitation>
|
||
). Caudal trichomes 2–4 hooks. (These caudal structures are like those observed in
|
||
<taxonomicName id="4C6B4D7C8603FFE0FA56FCA856F1FC81" authority="(Huynh and Veenstra 2018 c)" baseAuthorityName="Huynh and Veenstra" baseAuthorityYear="2018" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="intragramineus">
|
||
<emphasis id="B91FEAED8603FFE0FA56FCA8553EFC81" italics="true" pageId="21" pageNumber="22">U. intragramineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFA4B0E8603FFE0FE85FCCF56F9FC81" author="Huynh, C. & Veenstra, A." box="[338,643,837,863]" pageId="21" pageNumber="22" pagination="283 - 302" refId="ref14755" refString="Huynh, C. & Veenstra, A. (2018 c) Two new species pf penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam. Zootaxa, 4402 (2), 283 - 302. https: // doi. org / 10.11646 / zootaxa. 4402.2.3" type="journal article" year="2018">Huynh and Veenstra 2018c</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
which is common to members of the genus
|
||
<taxonomicName id="4C6B4D7C8603FFE0FBAEFCCC5099FC81" authorityName="Jones" authorityYear="1944" box="[1145,1251,838,863]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAED8603FFE0FBAEFCCC5099FC81" box="[1145,1251,838,863]" italics="true" pageId="21" pageNumber="22">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
and is classified as caudal bundle type I (
|
||
<bibRefCitation id="EFFA4B0E8603FFE0FE7FFCE35700FC5D" author="Conde, B. & Nguyen Duy-Jacquemin, M." box="[424,890,873,899]" pageId="21" pageNumber="22" pagination="291 - 302" refId="ref14429" refString="Conde, B. & Nguyen Duy-Jacquemin, M. (2008) Classification actuelle des Diplopodes Penicillates (Myriapodes) avec nouvelles d e finition des taxa. Bulletin de la Societe Zoologique de France, 133 (4), 291 - 302." type="journal article" year="2008">Condé and Nguyen Duy-Jacquemin 2008</bibRefCitation>
|
||
))
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1466778603FFE0FF40F9E1569DF95A" ID-DOI="http://doi.org/10.5281/zenodo.3735891" ID-Zenodo-Dep="3735891" box="[151,743,1643,1668]" httpUri="https://zenodo.org/record/3735891/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1643,1668]" targetBox="[205,1369,906,1626]" targetPageId="21">
|
||
<paragraph id="8BD436FF8603FFE0FF40F9E1569DF95A" blockId="21.[151,743,1643,1668]" box="[151,743,1643,1668]" pageId="21" pageNumber="22">
|
||
<emphasis id="B91FEAED8603FFE0FF40F9E1555AF95A" bold="true" box="[151,288,1643,1668]" pageId="21" pageNumber="22">FIGURE 17.</emphasis>
|
||
Digital image of
|
||
<taxonomicName id="4C6B4D7C8603FFE0FE05F9E756DDF95A" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[466,679,1645,1668]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="moniqueae" status="sp. nov.">
|
||
<emphasis id="B91FEAED8603FFE0FE05F9E756DDF95A" box="[466,679,1645,1668]" italics="true" pageId="21" pageNumber="22">Unixenus moniqueae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57968603FFE0FD79F9E6569DF95A" box="[686,743,1644,1668]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37165748603FFE0FF10F93950C9F833" pageId="21" pageNumber="22" type="discussion">
|
||
<paragraph id="8BD436FF8603FFE0FF10F93950C9F833" blockId="21.[151,1436,1714,2029]" pageId="21" pageNumber="22">
|
||
<emphasis id="B91FEAED8603FFE0FF10F939554EF913" bold="true" box="[199,308,1715,1741]" pageId="21" pageNumber="22">Remarks</emphasis>
|
||
:
|
||
<taxonomicName id="4C6B4D7C8603FFE0FE95F9395657F912" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[322,557,1715,1740]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="moniqueae" status="sp. nov.">
|
||
<emphasis id="B91FEAED8603FFE0FE95F9395657F912" box="[322,557,1715,1740]" italics="true" pageId="21" pageNumber="22">Unixenus moniqueae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91FEAED8603FFE0FDE5F939560AF913" bold="true" box="[562,624,1715,1741]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A22C57968603FFE0FDE5F939560AF913" box="[562,624,1715,1741]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was first identified as
|
||
<taxonomicName id="4C6B4D7C8603FFE0FCBDF93957AEF912" authorityName="Jones" authorityYear="1944" box="[874,980,1715,1740]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAED8603FFE0FCBDF93957AEF912" box="[874,980,1715,1740]" italics="true" pageId="21" pageNumber="22">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
sp. by Monique
|
||
<bibRefCitation id="EFFA4B0E8603FFE0FB5BF939551AF92F" author="Nguyen Duy-Jacquemin, M. & Conde, B." pageId="21" pageNumber="22" pagination="43 - 81" refId="ref15366" refString="Nguyen Duy-Jacquemin, M. & Conde, B. (1967) Morphologie et geonemie du genre Monographis. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 64, 43 - 81." type="journal article" year="1967">Nguyen Duy-Jacquemin and Condé (1967)</bibRefCitation>
|
||
based on two adult specimens (a male and a female), which were collected from Con Son Island,
|
||
<collectingCountry id="F37C766F8603FFE0FF40F9705480F8CA" box="[151,250,1786,1812]" name="Vietnam" pageId="21" pageNumber="22">Vietnam</collectingCountry>
|
||
. These specimens were part of the Indochinese collections of Dr C. Dawydoff (
|
||
<bibRefCitation id="EFFA4B0E8603FFE0FB4BF971514FF8CB" author="Attems, C." box="[1180,1333,1787,1813]" pageId="21" pageNumber="22" pagination="187 - 353" refId="ref14175" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'histoire naturelle, Nouvelle Serie, 6, 187 - 353." type="journal article" year="1938">Attems 1938</bibRefCitation>
|
||
). Formal naming of these specimens was deferred because of uncertainty around the specific diagnosis of
|
||
<taxonomicName id="4C6B4D7C8603FFE0FB33F8955134F8E6" authorityName="Jones" authorityYear="1944" box="[1252,1358,1823,1848]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAED8603FFE0FB33F8955134F8E6" box="[1252,1358,1823,1848]" italics="true" pageId="21" pageNumber="22">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
at that time, despite this species having similar characteristics to
|
||
<taxonomicName id="4C6B4D7C8603FFE0FCE1F8C957B6F882" authorityName="Verhoeff" authorityYear="1924" box="[822,972,1858,1884]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mjoebergi">
|
||
<emphasis id="B91FEAED8603FFE0FCE1F8C957B6F882" box="[822,972,1858,1884]" italics="true" pageId="21" pageNumber="22">U. mjoebergi</emphasis>
|
||
</taxonomicName>
|
||
and other evidence that supported them belonging to the same genus (
|
||
<bibRefCitation id="EFFA4B0E8603FFE0FE33F8ED57C5F85F" author="Nguyen Duy-Jacquemin, M. & Conde, B." box="[484,959,1894,1921]" pageId="21" pageNumber="22" pagination="43 - 81" refId="ref15366" refString="Nguyen Duy-Jacquemin, M. & Conde, B. (1967) Morphologie et geonemie du genre Monographis. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 64, 43 - 81." type="journal article" year="1967">Nguyen Duy-Jacquemin and Condé, 1967</bibRefCitation>
|
||
). The identification of
|
||
<taxonomicName id="4C6B4D7C8603FFE0FB69F8ED5152F85E" authorityName="Jones" authorityYear="1944" box="[1214,1320,1895,1920]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAED8603FFE0FB69F8ED5152F85E" box="[1214,1320,1895,1920]" italics="true" pageId="21" pageNumber="22">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
to species proved difficult due to similarities in the morphology of the main characteristics used for identification: the sensilla on antennomere VI, pointed spherical papillae of the labrum, 13 sensilla on the lateral palp of gnathochilarium, the arrangement of the tergal trichome sockets, chaetotaxy and the setiform sensillum on tarsus 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37165748600FFE3FF10FF135726FF25" pageId="22" pageNumber="23" type="etymology">
|
||
<paragraph id="8BD436FF8600FFE3FF10FF135726FF25" blockId="22.[151,1436,153,251]" pageId="22" pageNumber="23">
|
||
<emphasis id="B91FEAED8600FFE3FF10FF135532FF6D" bold="true" box="[199,328,153,179]" pageId="22" pageNumber="23">Etymology</emphasis>
|
||
: The species is named in honour of Dr. Monique Nguyen Duy-Jacquemin who made a significant contribution to the study of penicillate millipedes since the 1960s. She was the first to identify this species collected from Con Son Island, Con Dao Islands in 1967 as
|
||
<taxonomicName id="4C6B4D7C8600FFE3FD1CFF68574FFF25" authorityName="Jones" authorityYear="1944" box="[715,821,226,251]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAED8600FFE3FD1CFF68574FFF25" box="[715,821,226,251]" italics="true" pageId="22" pageNumber="23">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |