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<document id="979BDDF2C69C41E38CDC4317652BB8AD" ID-CLB-Dataset="24885" ID-DOI="10.11646/zootaxa.4695.6.9" ID-GBIF-Dataset="e0078a85-2a99-4e9b-8add-91c9029db2b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3542761" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573811501692" checkinUser="plazi" docAuthor="Delgado, Leonardo &amp; Mora-Aguilar, Eder F." docDate="2019" docId="03C287E25F334931D4B386D4FA4DA073" docLanguage="en" docName="zootaxa.4695.6.9.pdf" docOrigin="Zootaxa 4695 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus dicranius Bates 1887" docType="key" docVersion="9" lastPageNumber="591" masterDocId="FFFBFF9A5F364934D424864EFFD1A420" masterDocTitle="A new Mexican species of Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae), with a revised key to the species of the O. dicranius species complex" masterLastPageNumber="592" masterPageNumber="586" pageNumber="591" updateTime="1698765991886" updateUser="ExternalLinkService">
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<mods:title id="0487A2195D032B22DE97D327E7AD3604">A new Mexican species of Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae), with a revised key to the species of the O. dicranius species complex</mods:title>
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<mods:namePart id="62D6D540E631380683DC4794CDE1C9C3">Delgado, Leonardo</mods:namePart>
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<mods:namePart id="990FBD5A4C8F5FF13C7E68E30BE4B22B">Mora-Aguilar, Eder F.</mods:namePart>
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<mods:title id="1EED0D01CCA972B686DDC4A28BC6B7AA">Zootaxa</mods:title>
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<mods:date id="FCC5235C3B85B71D3E9628B4E5DA38AD">2019</mods:date>
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<mods:number id="6946FF4F01EDB7A02C95FAD1C4F639D0">2019-11-13</mods:number>
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<mods:number id="6972399970512434E9884ADC5EC83E92">4695</mods:number>
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<mods:identifier id="B8423F15D459CC029A5848E6FA9A4994" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C287E25F334931D4B386D4FA4DA073" ID-GBIF-Taxon="160406265" LSID="urn:lsid:plazi:treatment:03C287E25F334931D4B386D4FA4DA073" httpUri="http://treatment.plazi.org/id/03C287E25F334931D4B386D4FA4DA073" lastPageNumber="591" pageId="5" pageNumber="591">
<subSubSection id="C371657F5F334931D4B386D4FC3DA4DB" pageId="5" pageNumber="591" type="nomenclature">
<paragraph id="8BD436F45F334931D4B386D4FC28A494" blockId="5.[151,1017,153,180]" box="[151,1017,153,180]" pageId="5" pageNumber="591">
<heading id="D09C81985F334931D4B386D4FC28A494" bold="true" box="[151,1017,153,180]" fontSize="11" level="1" pageId="5" pageNumber="591" reason="1">
<emphasis id="B91FEAE65F334931D4B386D4FC28A494" bold="true" box="[151,1017,153,180]" pageId="5" pageNumber="591">
Key to separate species in the
<taxonomicName id="4C6B4D775F334931D62A86D7FCFDA494" ID-CoL="7XMRK" authorityName="Bates" authorityYear="1887" box="[526,812,153,180]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="dicranius">
<emphasis id="B91FEAE65F334931D62A86D7FCFDA494" bold="true" box="[526,812,153,180]" italics="true" pageId="5" pageNumber="591">Onthophagus dicranius</emphasis>
</taxonomicName>
species complex
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD436F45F334931D4B386AFFC3DA4DB" blockId="5.[151,1004,225,251]" box="[151,1004,225,251]" pageId="5" pageNumber="591">
(partially modified from
<bibRefCitation id="EFFA4B055F334931D58A86AFFD4DA4DB" author="Howden, H. F. &amp; Gill, B. D." box="[430,668,225,251]" pageId="5" pageNumber="591" pagination="1091 - 1114" refId="ref4346" refString="Howden, H. F. &amp; Gill, B. D. (1993) Mesoamerican Onthophagus Latreille in the dicranius and mirabilis species group (Coleoptera: Scarabaeidae). The Canadian Entomologist, 125, 1091 - 1114. https: // doi. org / 10.4039 / Ent 1251091 - 6" type="journal article" year="1993">Howden &amp; Gill 1993</bibRefCitation>
and
<bibRefCitation id="EFFA4B055F334931D6F686AFFC35A4DB" author="Kohlmann, B. &amp; Solis, A." box="[722,996,225,251]" pageId="5" pageNumber="591" pagination="159 - 261" refId="ref4401" refString="Kohlmann, B. &amp; Solis, A. (2001) El genero Onthophagus (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 49, 159 - 261." type="journal article" year="2001">Kohlmann &amp; Solís 2001</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C371657F5F334931D4B38768FA4DA073" pageId="5" pageNumber="591" type="key">
<keyStep id="309F2E515F334931D4B38768FA4DA578" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B38768FA4DA51B" blockId="5.[151,1437,294,1108]" box="[151,1436,294,316]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B38768FA4DA51B" box="[151,1436,294,316]" pageId="5" pageNumber="591">
1. Elytral intervals with stout setae, in three or more rows on second interval;
<collectingCountry id="F37C76645F334931D7908768FBD1A51B" box="[948,1024,294,315]" name="Panama" pageId="5" pageNumber="591">Panama</collectingCountry>
...
<taxonomicName id="4C6B4D775F334931D0118768FA4DA51B" ID-CoL="7XMSG" authority="Howden &amp; Gill, 1987" authorityName="Howden &amp; Gill" authorityYear="1987" box="[1077,1436,294,316]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="dorsipilulus">
<emphasis id="B91FEAE65F334931D0118768FB14A51B" box="[1077,1221,294,315]" italics="true" pageId="5" pageNumber="591">O. dorsipilulus</emphasis>
<bibRefCitation id="EFFA4B055F334931D0EF8768FA4DA51B" author="Howden, H &amp; Gill, B. D." box="[1227,1436,294,316]" pageId="5" pageNumber="591" pagination="201 - 224" refId="ref4305" refString="Howden, H .. &amp; Gill, B. D. (1987) New species and new records of Panamanian and Costa Rican Scarabaeinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 41, 201 - 224." type="journal article" year="1987">Howden &amp; Gill, 1987</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD436F45F334931D4B3870AFA4DA578" blockId="5.[151,1437,294,1108]" box="[151,1436,323,345]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B3870AFA4DA578" box="[151,1436,323,345]" pageId="5" pageNumber="591">- Elytral intervals glabrous or with fine setae, in two rows on second interval....................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B3872FFA4DA5EE" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B3872FFA4DA5B3" blockId="5.[151,1437,294,1108]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B3872FFA4DA5B3" pageId="5" pageNumber="591">2 Punctures of the pronotal disc simple or weakly annulate, small, shallow, and sparse (most separated by two or more puncture diameters)........................................................................................... 3</keyLead>
</paragraph>
<paragraph id="8BD436F45F334931D4B387D2FA4DA5EE" blockId="5.[151,1437,294,1108]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B387D2FA4DA5EE" pageId="5" pageNumber="591">- Punctures of the pronotal disc with annulate and/or large, deep, and dense (most separated by less than one puncture diameter)............................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B38798FA4DA664" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B38798FA4DA628" blockId="5.[151,1437,294,1108]" pageId="5" pageNumber="591">
<materialsCitation id="3B033CA95F334931D4B38798FA4DA628" ID-GBIF-Occurrence="2451575214" country="Mexico" location="Mexico" pageId="5" pageNumber="591" specimenCount="3" specimenCount-male="3">
<keyLead id="309A95C15F334931D4B38798FA4DA628" pageId="5" pageNumber="591">
<specimenCount id="9D6DFD7D5F334931D4B38798FECCA5CC" box="[151,285,470,492]" pageId="5" pageNumber="591" type="male">3 Males</specimenCount>
with frons and vertex flat, with distinct, moderate-sized punctures (
<figureCitation id="13502A715F334931D78A8798FBC9A5CC" box="[942,1048,470,492]" captionStart="FIGURES 9–18" captionStartId="3.[151,275,1853,1880]" captionTargetBox="[192,1415,190,1785]" captionTargetId="figure@3.[151,1435,164,1833]" captionText="FIGURES 9–18. Onthophagus species. 9, ¾ profile view, male of O. subcancer; 10, ¾ profile view, female of O. subcancer; 11, lateral view of genitalia, male of O. subcancer; 12, dorsal view of genitalia, male of O. subcancer; 13, protibia of O. chimalapensis, holotype; 14, protibia of O. aperodorsatus, male paratype; 15, protibia of O. neomirabilis, male; 16, protibia of O. chimalapensis, female paratype; 17, protibia of O. asperodorsatus, female; 18, protibia of O. neomirabilis, female." figureDoi="http://doi.org/10.5281/zenodo.3613726" httpUri="https://zenodo.org/record/3613726/files/figure.png" pageId="5" pageNumber="591">Figs. 9–10</figureCitation>
); females with pronotum evenly convex (
<figureCitation id="13502A715F334931D4CD87BDFEE0A629" box="[233,305,499,521]" captionStart="FIGURES 9–18" captionStartId="3.[151,275,1853,1880]" captionTargetBox="[192,1415,190,1785]" captionTargetId="figure@3.[151,1435,164,1833]" captionText="FIGURES 9–18. Onthophagus species. 9, ¾ profile view, male of O. subcancer; 10, ¾ profile view, female of O. subcancer; 11, lateral view of genitalia, male of O. subcancer; 12, dorsal view of genitalia, male of O. subcancer; 13, protibia of O. chimalapensis, holotype; 14, protibia of O. aperodorsatus, male paratype; 15, protibia of O. neomirabilis, male; 16, protibia of O. chimalapensis, female paratype; 17, protibia of O. asperodorsatus, female; 18, protibia of O. neomirabilis, female." figureDoi="http://doi.org/10.5281/zenodo.3613726" httpUri="https://zenodo.org/record/3613726/files/figure.png" pageId="5" pageNumber="591">Fig. 10</figureCitation>
);
<collectingCountry id="F37C76645F334931D56787BDFE5DA629" box="[323,396,499,521]" name="Mexico" pageId="5" pageNumber="591">Mexico</collectingCountry>
...............................................................
<taxonomicName id="4C6B4D775F334931D0AD87BAFA4DA628" ID-CoL="7XP96" authority="Howden, 1973" authorityName="Howden" authorityYear="1973" box="[1161,1436,499,521]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="subcancer">
<emphasis id="B91FEAE65F334931D0AD87BAFAD8A628" box="[1161,1289,499,521]" italics="true" pageId="5" pageNumber="591">O. subcancer</emphasis>
Howden, 1973
</taxonomicName>
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="8BD436F45F334931D4B3845FFA4DA664" blockId="5.[151,1437,294,1108]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B3845FFA4DA664" pageId="5" pageNumber="591">
- Males with frons and vertex convex, with fine punctures (
<figureCitation id="13502A715F334931D706845FFC52A606" box="[802,899,528,551]" captionStart="FIGURES 1–8" captionStartId="2.[151,275,1940,1967]" captionTargetBox="[183,1411,197,1903]" captionTargetId="figure@2.[151,1436,164,1920]" captionText="FIGURES 1–8. Onthophagus chimalapensis. 1, Habitus, holotype; 2, habitus, female paratype; 3, lateral view of male paratype; 4, lateral view of female paratype; 5, ¾ profile view of male holotype; 6, ¾ profile view of female paratype; 7, lateral view of genitalia, male paratype; 8, dorsal view of genitalia, male paratype." figureDoi="http://doi.org/10.5281/zenodo.3613724" httpUri="https://zenodo.org/record/3613724/files/figure.png" pageId="5" pageNumber="591">Figs. 5–6</figureCitation>
); females with pronotal disc slightly concave (
<figureCitation id="13502A715F334931D176845FFA41A607" box="[1362,1424,529,551]" captionStart="FIGURES 1–8" captionStartId="2.[151,275,1940,1967]" captionTargetBox="[183,1411,197,1903]" captionTargetId="figure@2.[151,1436,164,1920]" captionText="FIGURES 1–8. Onthophagus chimalapensis. 1, Habitus, holotype; 2, habitus, female paratype; 3, lateral view of male paratype; 4, lateral view of female paratype; 5, ¾ profile view of male holotype; 6, ¾ profile view of female paratype; 7, lateral view of genitalia, male paratype; 8, dorsal view of genitalia, male paratype." figureDoi="http://doi.org/10.5281/zenodo.3613724" httpUri="https://zenodo.org/record/3613724/files/figure.png" pageId="5" pageNumber="591">Fig. 6</figureCitation>
);
<collectingCountry id="F37C76645F334931D4C68460FEFAA664" box="[226,299,558,580]" name="Mexico" pageId="5" pageNumber="591">Mexico</collectingCountry>
.................................................
<taxonomicName id="4C6B4D775F334931D7588460FAF1A664" ID-CoL="7XML7" authority="Delgado &amp; Mora-Aguilar" authorityName="Delgado &amp; Mora-Aguilar" authorityYear="2019" box="[892,1312,558,580]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="chimalapensis" status="sp. nov.">
<emphasis id="B91FEAE65F334931D7588460FBF3A663" box="[892,1058,558,579]" italics="true" pageId="5" pageNumber="591">O. chimalapensis</emphasis>
Delgado &amp; Mora-Aguilar
</taxonomicName>
,
<taxonomicNameLabel id="A22C579D5F334931D10D8460FA4DA664" box="[1321,1436,558,580]" pageId="5" pageNumber="591" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B38405FA4DA65E" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B38405FA4DA641" blockId="5.[151,1437,294,1108]" box="[151,1436,587,609]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B38405FA4DA641" box="[151,1436,587,609]" pageId="5" pageNumber="591">4 Pronotum and elytra distinctly setose with short or long setae.................................................. 5</keyLead>
</paragraph>
<paragraph id="8BD436F45F334931D4B38427FA4DA65E" blockId="5.[151,1437,294,1108]" box="[151,1436,617,639]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B38427FA4DA65E" box="[151,1436,617,639]" pageId="5" pageNumber="591">- Pronotum and elytra glabrous or with minute setae.......................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B384C8FA4DA699" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B384C8FA4DA6BB" blockId="5.[151,1437,294,1108]" box="[151,1436,646,668]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B384C8FA4DA6BB" box="[151,1436,646,668]" pageId="5" pageNumber="591">
5 Elytral intervals with simple punctures;
<collectingCountry id="F37C76645F334931D64784C8FD7DA6BC" box="[611,684,646,668]" name="Mexico" pageId="5" pageNumber="591">Mexico</collectingCountry>
.............................
<taxonomicName id="4C6B4D775F334931D03084C8FA4DA6BB" ID-CoL="7XMCX" authority="Howden &amp; Gill, 1993" authorityName="Howden &amp; Gill" authorityYear="1993" box="[1044,1436,646,668]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="asperodorsatus">
<emphasis id="B91FEAE65F334931D03084C8FB14A6BB" box="[1044,1221,646,667]" italics="true" pageId="5" pageNumber="591">O. asperodorsatus</emphasis>
<bibRefCitation id="EFFA4B055F334931D0EF84C8FA4DA6BB" author="Howden, H. F. &amp; Gill, B. D." box="[1227,1436,646,668]" pageId="5" pageNumber="591" pagination="1091 - 1114" refId="ref4346" refString="Howden, H. F. &amp; Gill, B. D. (1993) Mesoamerican Onthophagus Latreille in the dicranius and mirabilis species group (Coleoptera: Scarabaeidae). The Canadian Entomologist, 125, 1091 - 1114. https: // doi. org / 10.4039 / Ent 1251091 - 6" type="journal article" year="1993">Howden &amp; Gill, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD436F45F334931D4B384EAFA4DA699" blockId="5.[151,1437,294,1108]" box="[151,1436,675,697]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B384EAFA4DA699" box="[151,1436,675,697]" pageId="5" pageNumber="591">- Elytral intervals with annulate punctures................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B3848FFA4DA730" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B3848FFA4DA6F6" blockId="5.[151,1437,294,1108]" box="[151,1436,705,727]" pageId="5" pageNumber="591">
<materialsCitation id="3B033CA95F334931D4B3848FFA4DA6F6" ID-GBIF-Occurrence="2451575213" box="[151,1436,705,727]" country="Costa Rica" location="Costa Rica" pageId="5" pageNumber="591" specimenCount="6" specimenCount-male="6">
<keyLead id="309A95C15F334931D4B3848FFA4DA6F6" box="[151,1436,705,727]" pageId="5" pageNumber="591">
<specimenCount id="9D6DFD7D5F334931D4B3848FFEC2A6F7" box="[151,275,705,727]" pageId="5" pageNumber="591" type="male">6 Male</specimenCount>
with a short, broad, distinctly emarginate pronotal protrusion;
<collectingCountry id="F37C76645F334931D74A848FFC06A6F7" box="[878,983,705,727]" name="Costa Rica" pageId="5" pageNumber="591">Costa Rica</collectingCountry>
.
<taxonomicName id="4C6B4D775F334931D7D5848FFA4DA6F6" ID-CoL="7XMP7" authority="Kohlmann &amp; Solis, 2001" authorityName="Kohlmann &amp; Solis" authorityYear="2001" box="[1009,1436,705,727]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="cryptodicranius">
<emphasis id="B91FEAE65F334931D7D5848FFB77A6F6" box="[1009,1190,705,726]" italics="true" pageId="5" pageNumber="591">O. cryptodicranius</emphasis>
<bibRefCitation id="EFFA4B055F334931D088848FFA4DA6F6" author="Kohlmann, B. &amp; Solis, A." box="[1196,1436,705,727]" pageId="5" pageNumber="591" pagination="159 - 261" refId="ref4401" refString="Kohlmann, B. &amp; Solis, A. (2001) El genero Onthophagus (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 49, 159 - 261." type="journal article" year="2001">Kohlmann &amp; Solís, 2001</bibRefCitation>
</taxonomicName>
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</materialsCitation>
</paragraph>
<paragraph id="8BD436F45F334931D4B38490FA4DA730" blockId="5.[151,1437,294,1108]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B38490FA4DA730" pageId="5" pageNumber="591">
- Male with a moderately long, slender, slightly emarginate pronotal horn; female unknown;
<collectingCountry id="F37C76645F334931D06A8490FB67A6D4" box="[1102,1206,734,756]" name="Costa Rica" pageId="5" pageNumber="591">Costa Rica</collectingCountry>
...........................................................................................
<taxonomicName id="4C6B4D775F334931D04484B5FA4DA730" ID-CoL="7XNPL" authority="Kohlmann &amp; Solis, 2001" authorityName="Kohlmann &amp; Solis" authorityYear="2001" box="[1120,1436,763,785]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="nubilus">
<emphasis id="B91FEAE65F334931D04484B5FB76A730" box="[1120,1191,763,784]" italics="true" pageId="5" pageNumber="591">nubilus</emphasis>
<bibRefCitation id="EFFA4B055F334931D08884B5FA4DA730" author="Kohlmann, B. &amp; Solis, A." box="[1196,1436,763,785]" pageId="5" pageNumber="591" pagination="159 - 261" refId="ref4401" refString="Kohlmann, B. &amp; Solis, A. (2001) El genero Onthophagus (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 49, 159 - 261." type="journal article" year="2001">Kohlmann &amp; Solís, 2001</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B38557FA4DA76B" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B38557FA4DA70E" blockId="5.[151,1437,294,1108]" box="[151,1436,793,815]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B38557FA4DA70E" box="[151,1436,793,815]" pageId="5" pageNumber="591">
7 Profemora with central region yellow, the rest brown;
<collectingCountry id="F37C76645F334931D6CE8557FC84A70F" box="[746,853,793,815]" name="Costa Rica" pageId="5" pageNumber="591">Costa Rica</collectingCountry>
,
<collectingCountry id="F37C76645F334931D77B8557FC7AA70E" box="[863,939,793,814]" name="Panama" pageId="5" pageNumber="591">Panama</collectingCountry>
.....................
<taxonomicName id="4C6B4D775F334931D0898557FA4DA70E" ID-CoL="7XMRK" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[1197,1436,793,814]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="dicranius">
<emphasis id="B91FEAE65F334931D0898557FAF4A70E" box="[1197,1317,793,814]" italics="true" pageId="5" pageNumber="591">O. dicranius</emphasis>
Bates, 1887
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD436F45F334931D4B38578FA4DA76B" blockId="5.[151,1437,294,1108]" box="[151,1436,822,844]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B38578FA4DA76B" box="[151,1436,822,844]" pageId="5" pageNumber="591">- Profemora entirely brown.............................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B3851DFA4DA7A6" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B3851DFA4DA748" blockId="5.[151,1437,294,1108]" box="[151,1436,851,873]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B3851DFA4DA748" box="[151,1436,851,873]" pageId="5" pageNumber="591">
8 Elytra short, moderately rounded; brachypterous;
<collectingCountry id="F37C76645F334931D6E4851DFCF8A749" box="[704,809,851,873]" name="Costa Rica" pageId="5" pageNumber="591">Costa Rica</collectingCountry>
..................
<taxonomicName id="4C6B4D775F334931D035851AFA4DA748" ID-CoL="7XN5Y" authority="Kohlmann &amp; Solis, 2001" authorityName="Kohlmann &amp; Solis" authorityYear="2001" box="[1041,1436,851,873]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="inediapterus">
<emphasis id="B91FEAE65F334931D035851AFB77A748" box="[1041,1190,851,873]" italics="true" pageId="5" pageNumber="591">O. inediapterus</emphasis>
<bibRefCitation id="EFFA4B055F334931D088851DFA4DA748" author="Kohlmann, B. &amp; Solis, A." box="[1196,1436,851,873]" pageId="5" pageNumber="591" pagination="159 - 261" refId="ref4401" refString="Kohlmann, B. &amp; Solis, A. (2001) El genero Onthophagus (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 49, 159 - 261." type="journal article" year="2001">Kohlmann &amp; Solís, 2001</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD436F45F334931D4B3853FFA4DA7A6" blockId="5.[151,1437,294,1108]" box="[151,1436,881,903]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B3853FFA4DA7A6" box="[151,1436,881,903]" pageId="5" pageNumber="591">- Elytra and wings well developed......................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B385C0FA4DA7FE" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B385C0FA4DA7E0" blockId="5.[151,1437,294,1108]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B385C0FA4DA7E0" pageId="5" pageNumber="591">
9 Pygidium with deep and simple punctures, apex shiny; male with two laterally compressed and triangular pronotal projections;
<collectingCountry id="F37C76645F334931D4C685E5FEF3A7E1" box="[226,290,939,961]" name="Belize" pageId="5" pageNumber="591">Belize</collectingCountry>
,
<collectingCountry id="F37C76645F334931D50F85E5FEA5A7E1" box="[299,372,939,961]" name="Mexico" pageId="5" pageNumber="591">Mexico</collectingCountry>
...........................................................
<taxonomicName id="4C6B4D775F334931D06085E2FA4DA7E0" ID-CoL="7XNV9" authority="Howden &amp; Gill, 1993" authorityName="Howden &amp; Gill" authorityYear="1993" box="[1092,1436,939,961]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="petenensis">
<emphasis id="B91FEAE65F334931D06085E2FB14A7E1" box="[1092,1221,940,961]" italics="true" pageId="5" pageNumber="591">O. petenensis</emphasis>
<bibRefCitation id="EFFA4B055F334931D0EF85E5FA4DA7E0" author="Howden, H. F. &amp; Gill, B. D." box="[1227,1436,939,961]" pageId="5" pageNumber="591" pagination="1091 - 1114" refId="ref4346" refString="Howden, H. F. &amp; Gill, B. D. (1993) Mesoamerican Onthophagus Latreille in the dicranius and mirabilis species group (Coleoptera: Scarabaeidae). The Canadian Entomologist, 125, 1091 - 1114. https: // doi. org / 10.4039 / Ent 1251091 - 6" type="journal article" year="1993">Howden &amp; Gill, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD436F45F334931D4B38587FA4DA7FE" blockId="5.[151,1437,294,1108]" box="[151,1436,969,991]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B38587FA4DA7FE" box="[151,1436,969,991]" pageId="5" pageNumber="591">- Pygidium with shallow, annulate punctures, surface completely dull; males with a slender, long pronotal horn........... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E515F334931D4B385A8FA4DA073" pageId="5" pageNumber="591">
<paragraph id="8BD436F45F334931D4B385A8FA4DA038" blockId="5.[151,1437,294,1108]" pageId="5" pageNumber="591">
<materialsCitation id="3B033CA95F334931D4B385A8FA4DA038" ID-GBIF-Occurrence="2451575215" country="Costa Rica" location="Costa Rica" pageId="5" pageNumber="591" specimenCount="10" specimenCount-male="10">
<keyLead id="309A95C15F334931D4B385A8FA4DA038" pageId="5" pageNumber="591">
<specimenCount id="9D6DFD7D5F334931D4B385A8FEC2A7DC" box="[151,275,998,1020]" pageId="5" pageNumber="591" type="male">10 Male</specimenCount>
with a slender, feebly emarginate clypeal horn; female with a weak, frontoclypeal carina;
<collectingCountry id="F37C76645F334931D05285A8FB0FA7DC" box="[1142,1246,998,1020]" name="Costa Rica" pageId="5" pageNumber="591">Costa Rica</collectingCountry>
.......................................................................................
<taxonomicName id="4C6B4D775F334931D076824AFA4DA038" ID-CoL="7XP22" authority="Howden &amp; Gill, 1993" authorityName="Howden &amp; Gill" authorityYear="1993" box="[1106,1436,1027,1049]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="quetzalis">
<emphasis id="B91FEAE65F334931D076824AFB14A038" box="[1106,1221,1027,1049]" italics="true" pageId="5" pageNumber="591">O. quetzalis</emphasis>
<bibRefCitation id="EFFA4B055F334931D0EF824DFA4DA038" author="Howden, H. F. &amp; Gill, B. D." box="[1227,1436,1027,1049]" pageId="5" pageNumber="591" pagination="1091 - 1114" refId="ref4346" refString="Howden, H. F. &amp; Gill, B. D. (1993) Mesoamerican Onthophagus Latreille in the dicranius and mirabilis species group (Coleoptera: Scarabaeidae). The Canadian Entomologist, 125, 1091 - 1114. https: // doi. org / 10.4039 / Ent 1251091 - 6" type="journal article" year="1993">Howden &amp; Gill, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="8BD436F45F334931D4B3826FFA4DA073" blockId="5.[151,1437,294,1108]" pageId="5" pageNumber="591">
<keyLead id="309A95C15F334931D4B3826FFA4DA073" pageId="5" pageNumber="591">
- Male with a distally widened, bifurcate clypeal horn; female with a distinct frontoclypeal carina;
<collectingCountry id="F37C76645F334931D0A2826FFB03A016" box="[1158,1234,1057,1078]" name="Panama" pageId="5" pageNumber="591">Panama</collectingCountry>
..............................................................................................
<taxonomicName id="4C6B4D775F334931D0B38270FA4DA073" ID-CoL="7XN9P" authority="Genier, 2017" authorityName="Genier" authorityYear="2017" box="[1175,1436,1086,1108]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="kohlmanni">
<emphasis id="B91FEAE65F334931D0B38270FACBA073" box="[1175,1306,1086,1107]" italics="true" pageId="5" pageNumber="591">O. kohlmanni</emphasis>
<bibRefCitation id="EFFA4B055F334931D1048270FA4DA073" author="Genier, F." box="[1312,1436,1086,1108]" pageId="5" pageNumber="591" pagination="574 - 580" refId="ref4215" refString="Genier, F. (2017) A new Guatemalan cloud forest endemic Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae). The Canadian Entomologist, 149, 574 - 580. https: // doi. org / 10.4039 / tce. 2017.32" type="journal article" year="2017">Génier, 2017</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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