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<document id="140419EACE830A4778B9D52E1F6884EC" ID-DOI="10.15298/rusentj.31.3.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="13178078" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722593664899" checkinUser="felipe" docAuthor="Shcherbakov, D. E." docDate="2022" docId="03C287CFFFDB7E6E136BFE99FA996C5D" docLanguage="en" docName="RussEntomolJ.31.3.230-233.pdf" docOrigin="Russian Entomological Journal 31 (3)" docSource="http://dx.doi.org/10.15298/rusentj.31.3.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Ilimpeika humerosa Shcherbakov 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="231" masterDocId="FFFBFFB7FFDA7E6F1010FFA2FFBA6B2A" masterDocTitle="New genus of Ignotalidae (Cicadomorpha) with notes on other Homoptera from the Permian and Triassic of the Tunguska Basin" masterLastPageNumber="233" masterPageNumber="230" pageNumber="231" updateTime="1722633464945" updateUser="ExternalLinkService">
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<mods:title id="F0CC7FADE3E2FB6F2C19DBE43916B0B6">New genus of Ignotalidae (Cicadomorpha) with notes on other Homoptera from the Permian and Triassic of the Tunguska Basin</mods:title>
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<mods:namePart id="55873AC37DD21420B615EC68985AA693">Shcherbakov, D. E.</mods:namePart>
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<mods:title id="806906E801B5275ED1372EE37C529ED5">Russian Entomological Journal</mods:title>
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<mods:date id="D4BBF6A9A3E1A9BEA746E305CA6DB9D1">2022</mods:date>
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<mods:number id="C25349A425C1F79A8CFCC0096BA637A1">2022-09-30</mods:number>
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<mods:number id="C361CBE36DD129739AF71F1A5037CCD2">31</mods:number>
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<mods:identifier id="75B14F30789F3F433DB69E1F3622DB0A" type="DOI">10.15298/rusentj.31.3.03</mods:identifier>
<mods:identifier id="DE430B2CE799F2EB5EFC111C45E5A0F8" type="ISSN">0132-8069</mods:identifier>
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<treatment id="03C287CFFFDB7E6E136BFE99FA996C5D" LSID="urn:lsid:plazi:treatment:03C287CFFFDB7E6E136BFE99FA996C5D" httpUri="http://treatment.plazi.org/id/03C287CFFFDB7E6E136BFE99FA996C5D" lastPageNumber="231" pageId="1" pageNumber="231">
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<paragraph id="8BD436D9FFDB7E6E136BFE99FA926A79" blockId="1.[812,1399,315,1179]" box="[891,1320,315,339]" pageId="1" pageNumber="231">
<heading id="D09C81B5FFDB7E6E136BFE99FA926A79" box="[891,1320,315,339]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="231" reason="8">
<taxonomicName id="4C6B4D5AFFDB7E6E136BFE99FB5C6A79" ID-CoL="9WHX3" authority="Shcherbakov" authorityName="Shcherbakov" authorityYear="2022" box="[891,1254,315,339]" class="Insecta" family="Ignotalidae" genus="Ilimpeika" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="231" phylum="Arthropoda" rank="species" species="humerosa" status="sp. nov.">
<emphasis id="B91FEACBFFDB7E6E136BFE99FBEA6A79" box="[891,1104,315,339]" italics="true" pageId="1" pageNumber="231">Ilimpeika humerosa</emphasis>
Shcherbakov
</taxonomicName>
,
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<taxonomicNameLabel id="A22C57B0FFDB7E6E14E2FE99FA926A79" box="[1266,1320,315,339]" pageId="1" pageNumber="231" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3716552FFDB7E6E1433FEFBFB3A6A5B" box="[1059,1152,345,369]" pageId="1" pageNumber="231" type="description">
<paragraph id="8BD436D9FFDB7E6E1433FEFBFB3A6A5B" blockId="1.[812,1399,315,1179]" box="[1059,1152,345,369]" pageId="1" pageNumber="231">
<figureCitation id="13502A5CFFDB7E6E1433FEFBFB3A6A5B" box="[1059,1152,345,369]" captionStart="Figs 14" captionStartId="1.[226,263,1945,1964]" captionTargetBox="[188,1398,1229,1892]" captionTargetPageId="1" captionText="Figs 14. Ilimpeika humerosa gen. et sp.n.: 12 — holotype tegmen: 1 — photograph (mirrored); 2 — venation (dash-dotted, boundary of precostal area); 34 — paratype hind wing: 3 — photograph (mirrored); 4 — venation. Scale bars: 10 mm. Рис. 14. Ilimpeika humerosa gen. et sp.n.: 12 — голотип, переднее крыло: 1 — фото (отражено Зеркально); 2 — жилкование (Штрих-пунктиром граница прекостального полЯ); 34 — паратип, Заднее крыло: 3 — фото (отражено Зеркально); 4 — жилкование. МасШтаб: 10 мм." figureDoi="http://doi.org/10.5281/zenodo.13178080" httpUri="https://zenodo.org/record/13178080/files/figure.png" pageId="1" pageNumber="231">Figs 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716552FFDB7E6E1342FEDCFAFD697B" pageId="1" pageNumber="231" type="materials_examined">
<paragraph id="8BD436D9FFDB7E6E1342FEDCFAFD697B" blockId="1.[812,1399,315,1179]" pageId="1" pageNumber="231">
MATERIAL.
<typeStatus id="54D0887BFFDB7E6E13DFFEDCFBA56ABB" box="[975,1055,382,401]" pageId="1" pageNumber="231" type="holotype">Holotype</typeStatus>
<collectionCode id="ED7AAE1CFFDB7E6E1438FEDCFBF16ABB" box="[1064,1099,382,401]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="231">PIN</collectionCode>
2099/2, incomplete right tegmen (clavus missing) on reddish burnt carbonaceous siltstone;
<typeStatus id="54D0887BFFDB7E6E153CFE34FACC6A83" box="[1324,1398,406,425]" pageId="1" pageNumber="231" type="paratype">paratype</typeStatus>
<collectionCode id="ED7AAE1CFFDB7E6E133CFE0CFCF56AEB" box="[812,847,430,449]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="231">PIN</collectionCode>
2099/1, left hind wing (anal area missing) on orange burnt siltstone;
<collectingRegion id="49AFF83BFFDB7E6E1391FE64FBA76AF3" box="[897,1053,454,473]" country="Russia" name="Krasnoyarsk" pageId="1" pageNumber="231">Krasnoyarsk Krai</collectingRegion>
, Evenkiysky District, right bank of the Ilimpeya River
<quantity id="4C939B3CFFDB7E6E13AAFE7CFC436ADB" box="[954,1017,478,497]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="231" unit="km" value="3.3">3.3 km</quantity>
downstream of the Ukshun(n)ukan River, collected by N.I. Emelyanov (All-Union Aerogeological Trust) in 1962; locality Ilimpeya, or Red Cliff (Красный утёс) [
<bibRefCitation id="EFFA4B28FFDB7E6E1522FDACFCE16913" author="Meyen S. V." pageId="1" pageNumber="231" pagination="1 - 220" refId="ref2816" refString="Meyen S. V. 1966. [Cordaites from the Upper Paleozoic of northern Eurasia (morphology, epidermal structure, taxonomy and stratigraphic significance)] // Tr. Geol. Inst. AN SSSR. Vol. 150. P. 1 - 220 [in Russian]." type="journal article" year="1966">Meyen, 1966</bibRefCitation>
], or Ukshunukan-2 (outcrop 716 of G.N.Sadovnikov);
<geoCoordinate id="EE5F501EFFDB7E6E1538FD87FAC96912" box="[1320,1395,549,568]" degrees="62.25" direction="north" orientation="latitude" pageId="1" pageNumber="231" precision="555" value="62.25">62.25°N</geoCoordinate>
,
<geoCoordinate id="EE5F501EFFDB7E6E133CFD9FFC3A697A" box="[812,896,573,592]" degrees="105.29" direction="east" orientation="longitude" pageId="1" pageNumber="231" precision="555" value="105.29">105.29°E</geoCoordinate>
; Degali Formation [
<bibRefCitation id="EFFA4B28FFDB7E6E1421FD9CFB11697B" author="Meyen S. V." box="[1073,1195,574,593]" pageId="1" pageNumber="231" pagination="1 - 220" refId="ref2816" refString="Meyen S. V. 1966. [Cordaites from the Upper Paleozoic of northern Eurasia (morphology, epidermal structure, taxonomy and stratigraphic significance)] // Tr. Geol. Inst. AN SSSR. Vol. 150. P. 1 - 220 [in Russian]." type="journal article" year="1966">Meyen, 1966</bibRefCitation>
], Upper Permian.
</paragraph>
</subSubSection>
<subSubSection id="C3716552FFDB7E6E1342FDF4FABF6881" pageId="1" pageNumber="231" type="description">
<paragraph id="8BD436D9FFDB7E6E1342FDF4FABF6881" blockId="1.[812,1399,315,1179]" pageId="1" pageNumber="231">
DESCRIPTION. Tegmen
<quantity id="4C939B3CFFDB7E6E145AFDF4FB366941" box="[1098,1164,598,619]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="231" unit="mm" value="47.0">47 mm</quantity>
long and
<quantity id="4C939B3CFFDB7E6E14F8FDF4FA906941" box="[1256,1322,598,619]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="231" unit="mm" value="19.0">19 mm</quantity>
wide as preserved (estimated full length ca.
<quantity id="4C939B3CFFDB7E6E146BFDD2FB7A69AF" box="[1147,1216,624,646]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="231" unit="mm" value="60.0">60 mm</quantity>
), elongate. Prenodal Sc branches weak, not branched profusely and not turned longitudinal towards wing margin. RA apparently with at least 4 branches, RP with about 6, M and CuA with about 10 branches each. Distinct crossveins only in postnodal part (except for one nodal
<emphasis id="B91FEACBFFDB7E6E13EDFD54FBA56821" box="[1021,1055,758,779]" italics="true" pageId="1" pageNumber="231">r-m</emphasis>
), numerous, slightly inclined distally, more close-set in CuA area. Hind wing
<quantity id="4C939B3CFFDB7E6E14AFFCB2FAB8680C" box="[1215,1282,784,806]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="231" unit="mm" value="35.0">35 mm</quantity>
long and
<quantity id="4C939B3CFFDB7E6E154FFCB2FCEB686A" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="231" unit="mm" value="22.0">22 mm</quantity>
wide as preserved, much shorter than tegmen. Costal margin strongly arched at base, with about ten strong coupling hooks. R stem short and oblique, RA with 3 main branches, RP with 6 branches, M with 3 or 4, CuA with bundle of at least 13 branches. Colour pattern on wings not preserved.
</paragraph>
</subSubSection>
<subSubSection id="C3716552FFDB7E6E1342FC12FC7F6FAB" pageId="1" pageNumber="231" type="discussion">
<paragraph id="8BD436D9FFDB7E6E1342FC12FC7F6FAB" blockId="1.[812,1399,315,1179]" pageId="1" pageNumber="231">
REMARKS. The locality was listed among those with the Korvunchana flora of possible Early Triassic age (and the formation was mentioned as the Limptekon Formation) [
<bibRefCitation id="EFFA4B28FFDB7E6E1543FC47FC6E6F3F" author="Shcherbakov D. E." pageId="1" pageNumber="231" refId="ref3447" refString="Shcherbakov D. E. 2000. Permian faunas of Homoptera (Hemiptera) in relation to phytogeography and the Permo-Triassic crisis // Paleontol. J. Vol. 34. Suppl. 3. P. S 251 - S 267." type="book" year="2000">Shcherbakov, 2000</bibRefCitation>
;
<bibRefCitation id="EFFA4B28FFDB7E6E13F2FBA2FA806F3F" author="Ponomarenko A. G. &amp; Shcherbakov D. E." box="[994,1338,1024,1046]" pageId="1" pageNumber="231" refId="ref3035" refString="Ponomarenko A. G., Shcherbakov D. E. 2004. New lacewings (Neuroptera) from the terminal Permian and basal Triassic of Siberia // Paleontol. J. Vol. 38. Suppl. 2. P. S 197 - S 203." type="book" year="2004">Ponomarenko, Shcherbakov, 2004</bibRefCitation>
;
<bibRefCitation id="EFFA4B28FFDB7E6E1559FBA3FC746F1B" author="Ponomarenko A. G." pageId="1" pageNumber="231" pagination="295 - 296" refId="ref2981" refString="Ponomarenko A. G. 2006. A new beetle species of the genus Taldycupes (Taldycupedidae, Coleoptera) from the Permian of the Tunguska River basin // Paleontol. J. Vol. 40. P. 295 - 296. https: // doi. org / 10.1134 / S 0031030106030105" type="journal article" year="2006">Ponomarenko, 2006</bibRefCitation>
], but in fact it is rich in cordaites, belongs to the Degali Formation s.str. (lower subformation of the former Degali Formation s.l.) and is undoubtedly Late Permian [
<bibRefCitation id="EFFA4B28FFDB7E6E1322FBC9FC016FAB" author="Meyen S. V." box="[818,955,1131,1153]" pageId="1" pageNumber="231" pagination="1 - 220" refId="ref2816" refString="Meyen S. V. 1966. [Cordaites from the Upper Paleozoic of northern Eurasia (morphology, epidermal structure, taxonomy and stratigraphic significance)] // Tr. Geol. Inst. AN SSSR. Vol. 150. P. 1 - 220 [in Russian]." type="journal article" year="1966">Meyen, 1966</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C3716552FFDB7E6E1342FB24FA996C5D" pageId="1" pageNumber="231" type="etymology">
<paragraph id="8BD436D9FFDB7E6E1342FB24FA896FB1" blockId="1.[812,1399,315,1179]" box="[850,1331,1157,1179]" pageId="1" pageNumber="231">
ETYMOLOGY. From Latin
<emphasis id="B91FEACBFFDB7E6E147EFB27FB7B6FB0" box="[1134,1217,1157,1178]" italics="true" pageId="1" pageNumber="231">humerus</emphasis>
(shoulder).
</paragraph>
<paragraph id="8BD436D9FFDB7E6E1504F8FFFA996C5D" blockId="1.[1300,1315,1885,1911]" box="[1300,1315,1885,1911]" pageId="1" pageNumber="231">
<emphasis id="B91FEACBFFDB7E6E1504F8FFFA996C5D" bold="true" box="[1300,1315,1885,1911]" pageId="1" pageNumber="231">4</emphasis>
</paragraph>
</subSubSection>
</treatment>
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