168 lines
23 KiB
XML
168 lines
23 KiB
XML
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<mods:title id="003F61F9C685FD9BE0E0ADBB058CAE4A">Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist</mods:title>
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<mods:namePart id="9F694EED27036BF1ADFF0E7ECE462508">Lobban, Christopher S.</mods:namePart>
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<mods:affiliation id="CF063855A3ED17D14EB028A1A7361CEE">Division of Natural Sciences, University of Guam, Mangilao, GU 96923, USA clobban @ triton. uog. edu</mods:affiliation>
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<mods:namePart id="E9DD37893A5DBE2D98E56906E7E6B443">Witkowski, Andrzej</mods:namePart>
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<mods:affiliation id="BCBD846F1EDA93FB750AAFF155865483">University of Szczecin, Institute of Marine and Environmental Sciences Mickiewicza 16 a, PL 70 - 383 Szczecin, Poland</mods:affiliation>
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<mods:title id="D6E361EEE4F016D5476E2BD677E74F03">Micronesica</mods:title>
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<mods:date id="609C4842D2FA3E28C9C8083E770EF652">2023</mods:date>
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<treatment id="03C287AAFFF70B0CFF357DCEFD49FBD2" ID-DOI="http://doi.org/10.5281/zenodo.12581877" ID-GBIF-Taxon="233850851" ID-Zenodo-Dep="12581877" LSID="urn:lsid:plazi:treatment:03C287AAFFF70B0CFF357DCEFD49FBD2" httpUri="http://treatment.plazi.org/id/03C287AAFFF70B0CFF357DCEFD49FBD2" lastPageNumber="12" pageId="11" pageNumber="12">
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<subSubSection id="C3716537FFF70B0CFF357DCEFAE2FEC4" box="[230,1401,238,264]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8BD436BCFFF70B0CFF357DCEFAE2FEC4" blockId="11.[230,1401,238,264]" box="[230,1401,238,264]" pageId="11" pageNumber="12">
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<heading id="D09C81D0FFF70B0CFF357DCEFAE2FEC4" box="[230,1401,238,264]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
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<taxonomicName id="4C6B4D3FFFF70B0CFF357DCEFD87FEC4" authority="Grunow" authorityName="Grunow" authorityYear="1878" box="[230,540,238,264]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="incurva">
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<emphasis id="B91FEAAEFFF70B0CFF357DCEFE2EFEC4" box="[230,437,238,264]" italics="true" pageId="11" pageNumber="12">Nitzschia incurva</emphasis>
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Grunow
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</taxonomicName>
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<figureCitation id="13502A39FFF70B0CFB257DCEFAE2FEC4" box="[1270,1401,238,264]" captionStart="Figures 71–80" captionStartId="61.[290,370,929,953]" captionTargetId="figure-17@62.[231,1380,341,1810]" captionTargetPageId="62" captionText="Figures 71–80. Moreneis, Nitzschia and Olifantiella. SEM except Fig. 71. Figs 71–74. Moreneis hexagona. Fig. 71. Valve in LM (DIC). Fig. 72. Valve exterior, showing isolated pores at the Voigt faults (arrows). Fig. 73. Valve exterior detail showing central raphe endings characteristic of the genus and c-shaped areolae. Fig. 74. Valve interior. Figs 75–78. Nitzschia incurva. Figs 75, 76. Guam voucher frustules at different angles. Figs 77, 78. Yap specimens showing interior and details of areolae. Figs 79, 80. Olifantiella mascarenica frustule showing the characteristic buciniportula (arrow). Scale bars: Fig 75 = 10 µm, Figs 71, 72, 74, 76, 77 = 5 µm, Figs 73, 78 = 2 µm, Figs 79, 80 = 1 µm." figureDoi="http://doi.org/10.5281/zenodo.12581773" httpUri="https://zenodo.org/record/12581773/files/figure.png" pageId="11" pageNumber="12">Figs 75–78</figureCitation>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3716537FFF70B0CFF357C38FA84FEB2" pageId="11" pageNumber="12" type="reference_group">
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<paragraph id="8BD436BCFFF70B0CFF357C38FB95FEFE" blockId="11.[230,1038,280,306]" box="[230,1038,280,306]" pageId="11" pageNumber="12">
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Syn.:
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<taxonomicName id="4C6B4D3FFFF70B0CFEF87C38FD1FFEFE" authority="Grunow" authorityName="Grunow" authorityYear="1879" box="[299,644,280,306]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lorenziana">
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<emphasis id="B91FEAAEFFF70B0CFEF87C38FD86FEFE" box="[299,541,280,306]" italics="true" pageId="11" pageNumber="12">Nitzschia lorenziana</emphasis>
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Grunow
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</taxonomicName>
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in Cleve & Grunow (nom. illeg.)
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</paragraph>
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<paragraph id="8BD436BCFFF70B0CFF357C62FA84FEB2" blockId="11.[230,1402,322,423]" pageId="11" pageNumber="12">
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Ref. illus.:
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<bibRefCitation id="EFFA4B4DFFF70B0CFEB67C62FD92FE90" author="Navarro, J. N." box="[357,521,322,348]" pageId="11" pageNumber="12" pagination="1 - 151" refId="ref17802" refString="Navarro, J. N. 1982. Marine diatoms associated with mangrove prop roots in the Indian River, Florida, U. S. A. Bibliotheca Phycologica 61: 1 - 151." type="journal article" year="1982">Navarro 1982</bibRefCitation>
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, pl. 35, fig. 4;
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<bibRefCitation id="EFFA4B4DFFF70B0CFD787C62FCE3FE90" author="Poulin M. & Berard-Therriault, A. & Cardinal & P. B. Hamilton" box="[683,888,322,348]" pageId="11" pageNumber="12" pagination="73 - 101" refId="ref18496" refString="Poulin M., L. Berard-Therriault, A. Cardinal & P. B. Hamilton. 1990. Les diatomees (Bacillariophyta) benthiques de substrats durs des eaux marines et saumatres du Quebec. 9. Bacillariaceae. Naturaliste canadien 117: 73 - 101." type="journal article" year="1990">Poulin et al. 1990</bibRefCitation>
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, p. 87, figs 67, 68, 71, 72 (
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<collectionCode id="ED7AAE79FFF70B0CFB777C62FB7DFE90" box="[1188,1254,322,348]" pageId="11" pageNumber="12">SEM</collectionCode>
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);
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<bibRefCitation id="EFFA4B4DFFF70B0CFB2A7C62FEFEFEB2" author="Witkowski, A. & H. Lange-Bertalot & D. Metzeltin" pageId="11" pageNumber="12" refId="ref20496" refString="Witkowski, A., H. Lange-Bertalot & D. Metzeltin. 2000. Diatom Flora of Marine Coasts, I. A. R. G. Gantner Verlag, Ruggell, Germany." type="book" year="2000">Witkowski et al. 2000</bibRefCitation>
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, p. 392, pl. 210, figs 24, 25, pl. 211, fig. 3, pl. 212, figs 1–3 (all as
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<taxonomicName id="4C6B4D3FFFF70B0CFBA67C44FA8EFEB2" authorityName="Grunow" authorityYear="1879" box="[1141,1301,356,382]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lorenziana">
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<emphasis id="B91FEAAEFFF70B0CFBA67C44FA8EFEB2" box="[1141,1301,356,382]" italics="true" pageId="11" pageNumber="12">N. lorenziana</emphasis>
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</taxonomicName>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3716537FFF70B0CFF357CADFC42FE1D" pageId="11" pageNumber="12" type="description">
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<paragraph id="8BD436BCFFF70B0CFF357CADFD70FE6B" blockId="11.[230,1402,322,423]" box="[230,747,397,423]" pageId="11" pageNumber="12">Samples: GU58G-4A, GU44I-1, GU44Z-15</paragraph>
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<paragraph id="8BD436BCFFF70B0CFF357C97FC42FE1D" blockId="11.[230,1402,439,675]" box="[230,985,439,465]" pageId="11" pageNumber="12">
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Dimensions: Length 18.5 µm, width 7.4 µm; striae
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<quantity id="4C939B59FFF70B0CFC917C97FCE5FE1D" box="[834,894,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="12" unit="in" value="20.0">20 in</quantity>
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10 µm.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3716537FFF70B0CFF357CC0FECFFD6F" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="8BD436BCFFF70B0CFF357CC0FECFFD6F" blockId="11.[230,1402,439,675]" pageId="11" pageNumber="12">
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Diagnostics: Weakly sigmoid in girdle view, keel eccentric except in long-rostrate apices, striation bold in LM because of external costae between the striae; central striae biseriate towards to keel, triseriate towards the margin (
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<figureCitation id="13502A39FFF70B0CFD897F04FD2CFDF2" box="[602,695,548,574]" captionStart="Figures 71–80" captionStartId="61.[290,370,929,953]" captionTargetId="figure-17@62.[231,1380,341,1810]" captionTargetPageId="62" captionText="Figures 71–80. Moreneis, Nitzschia and Olifantiella. SEM except Fig. 71. Figs 71–74. Moreneis hexagona. Fig. 71. Valve in LM (DIC). Fig. 72. Valve exterior, showing isolated pores at the Voigt faults (arrows). Fig. 73. Valve exterior detail showing central raphe endings characteristic of the genus and c-shaped areolae. Fig. 74. Valve interior. Figs 75–78. Nitzschia incurva. Figs 75, 76. Guam voucher frustules at different angles. Figs 77, 78. Yap specimens showing interior and details of areolae. Figs 79, 80. Olifantiella mascarenica frustule showing the characteristic buciniportula (arrow). Scale bars: Fig 75 = 10 µm, Figs 71, 72, 74, 76, 77 = 5 µm, Figs 73, 78 = 2 µm, Figs 79, 80 = 1 µm." figureDoi="http://doi.org/10.5281/zenodo.12581773" httpUri="https://zenodo.org/record/12581773/files/figure.png" pageId="11" pageNumber="12">Fig. 78</figureCitation>
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), becoming rings of pores with central row of pores and diminishing towards apex (
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<figureCitation id="13502A39FFF70B0CFDF87F66FD1EFDAC" box="[555,645,582,608]" captionStart="Figures 71–80" captionStartId="61.[290,370,929,953]" captionTargetId="figure-17@62.[231,1380,341,1810]" captionTargetPageId="62" captionText="Figures 71–80. Moreneis, Nitzschia and Olifantiella. SEM except Fig. 71. Figs 71–74. Moreneis hexagona. Fig. 71. Valve in LM (DIC). Fig. 72. Valve exterior, showing isolated pores at the Voigt faults (arrows). Fig. 73. Valve exterior detail showing central raphe endings characteristic of the genus and c-shaped areolae. Fig. 74. Valve interior. Figs 75–78. Nitzschia incurva. Figs 75, 76. Guam voucher frustules at different angles. Figs 77, 78. Yap specimens showing interior and details of areolae. Figs 79, 80. Olifantiella mascarenica frustule showing the characteristic buciniportula (arrow). Scale bars: Fig 75 = 10 µm, Figs 71, 72, 74, 76, 77 = 5 µm, Figs 73, 78 = 2 µm, Figs 79, 80 = 1 µm." figureDoi="http://doi.org/10.5281/zenodo.12581773" httpUri="https://zenodo.org/record/12581773/files/figure.png" pageId="11" pageNumber="12">Fig. 77</figureCitation>
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). Eccentric keels of the two valves are on opposite sides of the frustule (nitzschioid, not hantzschioid) (
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<figureCitation id="13502A39FFF70B0CFD677F47FCA2FD4D" box="[692,825,615,641]" captionStart="Figures 71–80" captionStartId="61.[290,370,929,953]" captionTargetId="figure-17@62.[231,1380,341,1810]" captionTargetPageId="62" captionText="Figures 71–80. Moreneis, Nitzschia and Olifantiella. SEM except Fig. 71. Figs 71–74. Moreneis hexagona. Fig. 71. Valve in LM (DIC). Fig. 72. Valve exterior, showing isolated pores at the Voigt faults (arrows). Fig. 73. Valve exterior detail showing central raphe endings characteristic of the genus and c-shaped areolae. Fig. 74. Valve interior. Figs 75–78. Nitzschia incurva. Figs 75, 76. Guam voucher frustules at different angles. Figs 77, 78. Yap specimens showing interior and details of areolae. Figs 79, 80. Olifantiella mascarenica frustule showing the characteristic buciniportula (arrow). Scale bars: Fig 75 = 10 µm, Figs 71, 72, 74, 76, 77 = 5 µm, Figs 73, 78 = 2 µm, Figs 79, 80 = 1 µm." figureDoi="http://doi.org/10.5281/zenodo.12581773" httpUri="https://zenodo.org/record/12581773/files/figure.png" pageId="11" pageNumber="12">Figs 77, 78</figureCitation>
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). Keel canal is closed except for circular fenestrae (
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<figureCitation id="13502A39FFF70B0CFF3C7FA9FEDCFD6F" box="[239,327,649,675]" captionStart="Figures 71–80" captionStartId="61.[290,370,929,953]" captionTargetId="figure-17@62.[231,1380,341,1810]" captionTargetPageId="62" captionText="Figures 71–80. Moreneis, Nitzschia and Olifantiella. SEM except Fig. 71. Figs 71–74. Moreneis hexagona. Fig. 71. Valve in LM (DIC). Fig. 72. Valve exterior, showing isolated pores at the Voigt faults (arrows). Fig. 73. Valve exterior detail showing central raphe endings characteristic of the genus and c-shaped areolae. Fig. 74. Valve interior. Figs 75–78. Nitzschia incurva. Figs 75, 76. Guam voucher frustules at different angles. Figs 77, 78. Yap specimens showing interior and details of areolae. Figs 79, 80. Olifantiella mascarenica frustule showing the characteristic buciniportula (arrow). Scale bars: Fig 75 = 10 µm, Figs 71, 72, 74, 76, 77 = 5 µm, Figs 73, 78 = 2 µm, Figs 79, 80 = 1 µm." figureDoi="http://doi.org/10.5281/zenodo.12581773" httpUri="https://zenodo.org/record/12581773/files/figure.png" pageId="11" pageNumber="12">Fig. 77</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3716537FFF70B0CFF357F93FD49FBD2" pageId="11" pageNumber="12" type="discussion">
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<paragraph id="8BD436BCFFF70B0CFF357F93FD49FBD2" blockId="11.[230,1402,691,1054]" pageId="11" pageNumber="12">
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Comments:
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<bibRefCitation id="EFFA4B4DFFF70B0CFEA67F93FD8BFD01" author="Guiry, M. D." box="[373,528,691,717]" pageId="11" pageNumber="12" refId="ref14793" refString="Guiry, M. D. 2017. Nitzschia lorenziana Grunow, nom. illeg. 1880; nomenclatural note. In, Guiry, M. D. & G. M. Guiry. 2022. AlgaeBase. World-wide electronic publication, National University of Ireland, Galway. https: // www. algaebase. org" type="url" year="2017">Guiry (2017)</bibRefCitation>
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explains why
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<taxonomicName id="4C6B4D3FFFF70B0CFD6F7F93FCA2FD01" authorityName="Grunow" authorityYear="1879" box="[700,825,691,717]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lorenziana">
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<emphasis id="B91FEAAEFFF70B0CFD6F7F93FCA2FD01" box="[700,825,691,717]" italics="true" pageId="11" pageNumber="12">lorenziana</emphasis>
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</taxonomicName>
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is illegitimate and provides the earlier synonym. The ultrastructure of our specimens clearly matches
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<collectionCode id="ED7AAE79FFF70B0CFCA77FF5FC2BFD23" box="[884,944,725,751]" pageId="11" pageNumber="12">SEM</collectionCode>
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images in
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<bibRefCitation id="EFFA4B4DFFF70B0CFB937FF5FAACFD23" author="Poulin M. & Berard-Therriault, A. & Cardinal & P. B. Hamilton" box="[1088,1335,725,751]" pageId="11" pageNumber="12" pagination="73 - 101" refId="ref18496" refString="Poulin M., L. Berard-Therriault, A. Cardinal & P. B. Hamilton. 1990. Les diatomees (Bacillariophyta) benthiques de substrats durs des eaux marines et saumatres du Quebec. 9. Bacillariaceae. Naturaliste canadien 117: 73 - 101." type="journal article" year="1990">Poulin et al. (1990)</bibRefCitation>
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. It is apparently a brackish/estuarine species, with wide temperature tolerance (from Gulf of St. Lawrence to Florida +
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<collectingCountry id="F37C762CFFF70B0CFEA97E38FE5CFCFE" box="[378,455,792,818]" name="Guam" pageId="11" pageNumber="12">Guam</collectingCountry>
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). Abundant in
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<collectingCountry id="F37C762CFFF70B0CFDA57E38FD40FCFE" box="[630,731,792,818]" name="Vietnam" pageId="11" pageNumber="12">Vietnam</collectingCountry>
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and
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<collectingCountry id="F37C762CFFF70B0CFCC67E38FC1CFCFE" box="[789,903,792,818]" name="Indonesia" pageId="11" pageNumber="12">Indonesia</collectingCountry>
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(Witkowski, unpubl.). Lopez-Fuerte et al. (2010: p. 63, pl. 37, fig. 21) show
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<taxonomicName id="4C6B4D3FFFF70B0CFD527E1AFCEFFC98" authority="Grunow" authorityName="Grunow" box="[641,884,826,852]" pageId="11" pageNumber="12" rank="variety" variety="subtilis">
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var.
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<emphasis id="B91FEAAEFFF70B0CFD657E1AFC90FC98" box="[694,779,826,852]" italics="true" pageId="11" pageNumber="12">subtilis</emphasis>
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Grunow
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</taxonomicName>
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from
|
||
<collectingCountry id="F37C762CFFF70B0CFC6D7E1AFB83FC98" box="[958,1048,826,852]" name="Mexico" pageId="11" pageNumber="12">Mexico</collectingCountry>
|
||
and list the nominate variety.
|
||
<taxonomicName id="4C6B4D3FFFF70B0CFF357E7CFE2DFCBA" authorityName="Grunow" authorityYear="1878" box="[230,438,860,886]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="incurva">
|
||
<emphasis id="B91FEAAEFFF70B0CFF357E7CFE2DFCBA" box="[230,438,860,886]" italics="true" pageId="11" pageNumber="12">Nitzschia incurva</emphasis>
|
||
</taxonomicName>
|
||
somewhat resembles
|
||
<taxonomicName id="4C6B4D3FFFF70B0CFD697E7CFA88FCBA" authority="Bing Liu, S. Blanco & B. Q. Huang (2019)" authorityName="Bing Liu, S. Blanco & B. Q. Huang" authorityYear="2019" box="[698,1299,860,886]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gaoi">
|
||
<emphasis id="B91FEAAEFFF70B0CFD697E7CFC8AFCBA" box="[698,785,860,886]" italics="true" pageId="11" pageNumber="12">N. gaoi</emphasis>
|
||
Bing Liu, S. Blanco & B.Q. Huang (2019)
|
||
</taxonomicName>
|
||
, but that has small conopeal canals and uniseriate striae.
|
||
<bibRefCitation id="EFFA4B4DFFF70B0CFCC27E5DFBFDFC5B" author="Lobban, C. S. & M. P. Ashworth" box="[785,1126,893,919]" pageId="11" pageNumber="12" refId="ref16280" refString="Lobban, C. S. & M. P. Ashworth. 2022 b. Homoeocladia C. Agardh reinstated for bilaterally symmetrical conopeate Nitzschia species (Bacillariaceae, Bacillariophyta). Notulae Algarum 267, 9 pp." type="book" year="2022">Lobban & Ashworth (2022b)</bibRefCitation>
|
||
did not transfer
|
||
<taxonomicName id="4C6B4D3FFFF70B0CFAF77E5DFAE1FC5B" authorityName="Bing Liu, S. Blanco & B. Q. Huang" authorityYear="2019" box="[1316,1402,893,919]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gaoi">
|
||
<emphasis id="B91FEAAEFFF70B0CFAF77E5DFAE1FC5B" box="[1316,1402,893,919]" italics="true" pageId="11" pageNumber="12">N. gaoi</emphasis>
|
||
</taxonomicName>
|
||
or their own
|
||
<taxonomicName id="4C6B4D3FFFF70B0CFE5C7EBFFB96FC75" authority="Lobban, Ashworth, Calaor & E. C. Theriot" authorityName="Lobban, Ashworth, Calaor & E. C. Theriot" authorityYear="2019" box="[399,1037,927,953]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="maiae">
|
||
<emphasis id="B91FEAAEFFF70B0CFE5C7EBFFE64FC75" box="[399,511,927,953]" italics="true" pageId="11" pageNumber="12">N. maiae</emphasis>
|
||
Lobban, Ashworth, Calaor & E.C.Theriot
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C6B4D3FFFF70B0CFBEE7EBFFB71FC75" authorityName="C.A.Agardh" authorityYear="1827" box="[1085,1258,927,953]" class="Bacillariophyceae" family="Bacillariaceae" genus="Homoeocladia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Bacillariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="B91FEAAEFFF70B0CFBEE7EBFFB71FC75" box="[1085,1258,927,953]" italics="true" pageId="11" pageNumber="12">Homoeocladia</emphasis>
|
||
</taxonomicName>
|
||
because of maintaining a narrow definition of the morphology of
|
||
<taxonomicName id="4C6B4D3FFFF70B0CFCB97EE1FB8CFC17" authorityName="C.A.Agardh" authorityYear="1827" box="[874,1047,961,987]" class="Bacillariophyceae" family="Bacillariaceae" genus="Homoeocladia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Bacillariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="B91FEAAEFFF70B0CFCB97EE1FB8CFC17" box="[874,1047,961,987]" italics="true" pageId="11" pageNumber="12">Homoeocladia</emphasis>
|
||
</taxonomicName>
|
||
and noted some other species in the region with doubtful formation of a conopeal canal that also have T-shaped external costae (one shown in
|
||
<bibRefCitation id="EFFA4B4DFFF70B0CFE477924FDA4FBD2" author="Lobban, C. S." box="[404,575,1028,1054]" pageId="11" pageNumber="12" pagination="73 - 79" refId="ref16135" refString="Lobban, C. S. 2023 a. Nitzschia venerata, sp. nov., a simulacrum for the conopeate, tube-dwelling diatom Nitzschia martiana, has scuta not conopea: a case of convergent evolution? Botanica Marina 66: 73 - 79." type="journal article" year="2023">Lobban 2023a</bibRefCitation>
|
||
, figs 4F, G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |