treatments-xml/data/03/C2/87/03C28789FFF3FF8194F8FEAD550B1FCE.xml
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<document id="F073EC53B19E2885732C18DCC5C4A1FE" ID-DOI="10.21248/contrib.entomol.57.2.367-396" ID-ISSN="0005-805X" ID-Zenodo-Dep="5364164" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630242723631" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2007" docId="03C28789FFF3FF8194F8FEAD550B1FCE" docLanguage="en" docName="ContribEntomol.2007.57.2.367-396.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 57 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1710" docStyle="DocumentStyle:DA3631FDA31ED51D1CF99ECA6F54F23F.3:ContribEntomol.2005-2012.journal_article" docStyleId="DA3631FDA31ED51D1CF99ECA6F54F23F" docStyleName="ContribEntomol.2005-2012.journal_article" docStyleVersion="3" docTitle="Pronomaea procerula ASSING" docType="key" docVersion="3" lastPageNumber="394" masterDocId="FFFBFFF1FFEAFF9A944DFFD3565D1C3A" masterDocTitle="A revision of the species of Pronomaea E of the Western Palaearctic region, including Middle Asia (Coleoptera: Staphylinidae: Aleocharinae: Pronomaeini)" masterLastPageNumber="396" masterPageNumber="367" pageNumber="392" updateTime="1699196480509" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="73A0442661FFA697655D494E17D65626">A revision of the species of Pronomaea E of the Western Palaearctic region, including Middle Asia (Coleoptera: Staphylinidae: Aleocharinae: Pronomaeini)</mods:title>
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<mods:name id="5977DCDD1C721234B526A20113A7A1BC" type="personal">
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<mods:namePart id="89BCA68A2D62E9C14CF96DA318010846">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="224319F25C2A752A4D21082CBCB9E525">text</mods:typeOfResource>
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<mods:title id="44431E13A6067DA7F9D472F443C97C68">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="4746E626AE5748734B6FDC4F91DAF7D8">
<mods:date id="7AE9431A16836463076F9F8E30881727">2007</mods:date>
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<mods:number id="417BB3C56E1B7ECD1B28190DF52BA6FD">57</mods:number>
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<mods:identifier id="D4D7CF0D61E046E90A04C0AF41B94D78" type="DOI">10.21248/contrib.entomol.57.2.367-396</mods:identifier>
<mods:identifier id="0FBB224E2E59C172B4745E085C68F15E" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="ED284E923A950D1FB374ECF3005BC88F" type="Zenodo-Dep">5364164</mods:identifier>
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<treatment id="03C28789FFF3FF8194F8FEAD550B1FCE" LSID="urn:lsid:plazi:treatment:03C28789FFF3FF8194F8FEAD550B1FCE" httpUri="http://treatment.plazi.org/id/03C28789FFF3FF8194F8FEAD550B1FCE" lastPageId="27" lastPageNumber="394" pageId="25" pageNumber="392">
<subSubSection id="C3716514FFF3FF8394F8FEAD54E01DAC" box="[181,701,381,406]" pageId="25" pageNumber="392" type="nomenclature">
<paragraph id="8BD4369FFFF3FF8394F8FEAD54E01DAC" blockId="25.[181,701,381,406]" box="[181,701,381,406]" pageId="25" pageNumber="392">
<heading id="D09C81F3FFF3FF8394F8FEAD54E01DAC" box="[181,701,381,406]" centered="true" fontSize="8" level="2" pageId="25" pageNumber="392" reason="2">
<emphasis id="B91FEA8DFFF3FF8394F8FEAD54031DAC" bold="true" box="[181,606,381,406]" pageId="25" pageNumber="392">
<taxonomicName id="4C6B4D1CFFF3FF8394F8FEAD57A41DAB" ID-CoL="4MTB3" authority="ASSING" authorityName="ASSING" box="[181,505,381,405]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="392" phylum="Arthropoda" rank="species" species="procerula">
<emphasis id="B91FEA8DFFF3FF8394F8FEAD57F41DAF" bold="true" box="[181,425,381,405]" italics="true" pageId="25" pageNumber="392">Pronomaea procerula</emphasis>
ASSING
</taxonomicName>
, in press
</emphasis>
(
<figureCitation id="13502A1AFFF3FF839622FEAD54E91DAC" box="[623,692,382,406]" captionStart="Map 5" captionStartId="24.[191,235,1184,1205]" captionTargetBox="[189,1172,467,1175]" captionTargetId="figure-284@24.[189,1172,467,1175]" captionTargetPageId="24" captionText="Map 5: Distributions of Pronomaea flavirostris SEMENOV (filled circles) and P. procerula ASSING (open circles), based on examined and literature records." figureDoi="http://doi.org/10.5281/zenodo.5364186" httpUri="https://zenodo.org/record/5364186/files/figure.png" pageId="25" pageNumber="392">Map 5</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3716514FFF3FF839427FE7F57111DE4" pageId="25" pageNumber="392" type="materials_examined">
<paragraph id="8BD4369FFFF3FF839427FE7F57111DE4" blockId="25.[106,1097,427,478]" pageId="25" pageNumber="392">
<emphasis id="B91FEA8DFFF3FF839427FE7F57D71D86" bold="true" box="[106,394,428,444]" pageId="25" pageNumber="392">Additional material examined:</emphasis>
<materialsCitation id="3B033CC2FFF3FF8395DEFE7F571A1DE4" collectingDate="1975-06-22" collectionCode="MHNG" collectorName="N Sanandajd &amp; Senglet" country="Iran" latitude="35.466667" location="Kordestan" longLatPrecision="1193" longitude="47.016666" pageId="25" pageNumber="392" specimenCount="2" stateProvince="Kordestan">
<emphasis id="B91FEA8DFFF3FF8395DEFE7F57E21D86" bold="true" box="[403,447,428,444]" pageId="25" pageNumber="392">
<collectingCountry id="F37C760FFFF3FF8395DEFE7F57E71D86" box="[403,442,428,444]" name="Iran" pageId="25" pageNumber="392">Iran</collectingCountry>
:
</emphasis>
<specimenCount id="9D6DFD16FFF3FF83958AFE7F57A31DFB" box="[455,510,428,449]" pageId="25" pageNumber="392" type="generic">2 exs.</specimenCount>
,
<collectingRegion id="49AFF87DFFF3FF839645FE7854341DFB" box="[520,617,427,449]" country="Iran" name="Kordestan" pageId="25" pageNumber="392">Kordestan</collectingRegion>
,
<collectorName id="269E5349FFF3FF83963EFE7F54AD1DFB" box="[627,752,427,449]" pageId="25" pageNumber="392">N Sanandajd</collectorName>
,
<geoCoordinate id="EE5F5058FFF3FF8396B7FE7855161D85" box="[762,843,427,449]" degrees="35" direction="north" minutes="28" orientation="latitude" pageId="25" pageNumber="392" precision="925" value="35.466667">35°28 'N</geoCoordinate>
,
<geoCoordinate id="EE5F5058FFF3FF83971AFE7F55F91DFB" box="[855,932,428,449]" degrees="47" direction="east" minutes="01" orientation="longitude" pageId="25" pageNumber="392" precision="925" value="47.016666">47°01 'E</geoCoordinate>
,
<date id="FFD5105FFFF3FF8397E3FE7852401DFB" box="[942,1053,427,449]" pageId="25" pageNumber="392" value="1975-06-22">
<collectingDate id="EF91E9B7FFF3FF8397E3FE7852401DFB" box="[942,1053,427,449]" pageId="25" pageNumber="392" value="1975-06-22">22.VI.1975</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFF3FF839426FE1B56F01DE4" box="[107,173,456,478]" pageId="25" pageNumber="392">Senglet</collectorName>
(
<collectionCode id="ED7AAE5AFFF3FF8394F7FE1B57561DE7" box="[186,267,456,477]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="25" pageNumber="392" type="Museum">MHNG</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716514FFF3FF839426FE27542E184B" pageId="25" pageNumber="392" type="diagnosis">
<paragraph id="8BD4369FFFF3FF839426FE2756841E36" blockId="25.[107,217,500,524]" box="[107,217,500,524]" pageId="25" pageNumber="392">
<emphasis id="B91FEA8DFFF3FF839426FE2756841E36" bold="true" box="[107,217,500,524]" pageId="25" pageNumber="392">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BD4369FFFF3FF839426FDC955061E09" blockId="25.[107,859,538,563]" box="[107,859,538,563]" pageId="25" pageNumber="392">Highly distinctive species; for a detailed description see ASSING (in press).</paragraph>
<paragraph id="8BD4369FFFF3FF839426FD9256F71EC8" blockId="25.[106,1097,577,754]" pageId="25" pageNumber="392">
Body slender. Antennae and legs conspicuously long and slender, preapical antennomeres not transverse. Pronotum of distinctive shape, almost cordiform: maximal width in anterior half, strongly tapering posteriad; width of posterior margin 0.75-0.80 times maximal pronotal width; lateral margins smoothly sinuate in posterior half. Elytra only with moderately dense puncturation. For illustrations of the habitus, the forebody, and the antenna see figures
<quantity id="4C939B7AFFF3FF83973EFD6855921EE9" box="[883,975,699,723]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" metricValueMax="6.35" metricValueMin="5.842" pageId="25" pageNumber="392" unit="in" value="24.0" valueMax="25.0" valueMin="23.0">23-25 in</quantity>
ASSING (in press).
</paragraph>
<paragraph id="8BD4369FFFF3FF839423FCD055F01F6C" blockId="25.[107,1096,768,854]" pageId="25" pageNumber="392">
<specimenCount id="9D6DFD16FFF3FF839423FCD056261F29" box="[110,123,771,787]" pageId="25" pageNumber="392" type="male"></specimenCount>
: aedeagus
<quantity id="4C939B7AFFF3FF8394A4FCD357221F22" box="[233,383,768,792]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.699999999999999" metricValueMin="4.5" pageId="25" pageNumber="392" unit="mm" value="0.45999999999999996" valueMax="0.47" valueMin="0.45">0.45-0.47 mm</quantity>
long); shape of median lobe somewhat resembling that of
<taxonomicName id="4C6B4D1CFFF3FF839789FCD2521C1F22" box="[964,1089,768,792]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="392" phylum="Arthropoda" rank="species" species="flavirostris">
<emphasis id="B91FEA8DFFF3FF839789FCD2521C1F22" box="[964,1089,768,792]" italics="true" pageId="25" pageNumber="392">P. flavirostris</emphasis>
</taxonomicName>
, but more slender, ventral process more strongly bent (in relation to base), and internal sac with remarkably long flagellum; for illustrations see figures
<quantity id="4C939B7AFFF3FF8396DAFCED54AD1F6C" box="[663,752,830,854]" metricMagnitude="-1" metricUnit="m" metricValue="7.239" metricValueMax="7.62" metricValueMin="6.858" pageId="25" pageNumber="392" unit="in" value="28.5" valueMax="30.0" valueMin="27.0">27-30 in</quantity>
ASSING (in press).
</paragraph>
<paragraph id="8BD4369FFFF3FF839422FCB657C01FA1" blockId="25.[107,1096,868,923]" pageId="25" pageNumber="392">
<specimenCount id="9D6DFD16FFF3FF839422FCB656241F4D" box="[111,121,869,887]" pageId="25" pageNumber="392" type="female"></specimenCount>
: spermatheca of similar shape as that of
<taxonomicName id="4C6B4D1CFFF3FF839663FCB754C21F4C" box="[558,671,868,886]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="392" phylum="Arthropoda" rank="species" species="anatolicaanatolica">
<emphasis id="B91FEA8DFFF3FF839663FCB754C21F4C" box="[558,671,868,886]" italics="true" pageId="25" pageNumber="392">P. anatolicaanatolica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D1CFFF3FF83969AFCB755061F4C" box="[727,859,868,886]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="392" phylum="Arthropoda" rank="species" species="khnzoriani">
<emphasis id="B91FEA8DFFF3FF83969AFCB755061F4C" box="[727,859,868,886]" italics="true" pageId="25" pageNumber="392">P. khnzoriani</emphasis>
</taxonomicName>
; for an illustration see figure
<quantity id="4C939B7AFFF3FF8394E1FC5056BF1FA0" box="[172,226,898,923]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="25" pageNumber="392" unit="in" value="34.0">34 in</quantity>
ASSING (in press).
</paragraph>
<paragraph id="8BD4369FFFF3FF839426FC6B57651FEA" blockId="25.[107,312,952,976]" box="[107,312,952,976]" pageId="25" pageNumber="392">
<emphasis id="B91FEA8DFFF3FF839426FC6B57651FEA" bold="true" box="[107,312,952,976]" pageId="25" pageNumber="392">Comparative notes:</emphasis>
</paragraph>
<paragraph id="8BD4369FFFF3FF839427FC0D542E184B" blockId="25.[106,1096,990,1137]" pageId="25" pageNumber="392">The species is readily distinguished from all other representatives of the genus distributed in the Western Palaearctic region and Middle Asia by the distinctly longer and more slender antennae and legs, the conspicuous shape of the pronotum (cordiform shape, sinuate lateral margins), the sparser puncturation of the pronotum and the elytra, as well as by the morphology of the aedeagus, especially the remarkably long flagellum.</paragraph>
</subSubSection>
<subSubSection id="C3716514FFF3FF839426FB5C55C91930" pageId="25" pageNumber="392" type="distribution">
<paragraph id="8BD4369FFFF3FF839426FB5C57CB189B" blockId="25.[107,406,1167,1185]" box="[107,406,1167,1185]" pageId="25" pageNumber="392">
<emphasis id="B91FEA8DFFF3FF839426FB5C57CB189B" bold="true" box="[107,406,1167,1185]" pageId="25" pageNumber="392">Distribution and bionomics:</emphasis>
</paragraph>
<paragraph id="8BD4369FFFF3FF839427FB6755C91930" blockId="25.[106,1096,1204,1290]" pageId="25" pageNumber="392">
The known distribution is confined to western
<collectingCountry id="F37C760FFFF3FF83961FFB66542118F6" box="[594,636,1205,1228]" name="Iran" pageId="25" pageNumber="392">Iran</collectingCountry>
(
<figureCitation id="13502A1AFFF3FF8396C7FB67548C18F6" box="[650,721,1204,1228]" captionStart="Map 5" captionStartId="24.[191,235,1184,1205]" captionTargetBox="[189,1172,467,1175]" captionTargetId="figure-284@24.[189,1172,467,1175]" captionTargetPageId="24" captionText="Map 5: Distributions of Pronomaea flavirostris SEMENOV (filled circles) and P. procerula ASSING (open circles), based on examined and literature records." figureDoi="http://doi.org/10.5281/zenodo.5364186" httpUri="https://zenodo.org/record/5364186/files/figure.png" pageId="25" pageNumber="392">Map 5</figureCitation>
); for additional localities see ASSING (in press). It is here reported from
<collectingCountry id="F37C760FFFF3FF839589FB00546218D1" box="[452,575,1235,1259]" name="Afghanistan" pageId="25" pageNumber="392">Afghanistan</collectingCountry>
for the first time. The species has been collected on river banks at altitudes of 1940 and
<quantity id="4C939B7AFFF3FF839590FB2154731932" box="[477,558,1266,1288]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="25" pageNumber="392" unit="m" value="2040.0">2040 m</quantity>
((ASSING in press) in June and July.
</paragraph>
</subSubSection>
<subSubSection id="C3716514FFF3FF8194FBFAED550B1FCE" lastPageId="27" lastPageNumber="394" pageId="25" pageNumber="392" type="key">
<paragraph id="8BD4369FFFF3FF8394FBFAED56B91955" blockId="25.[181,1097,1342,1391]" pageId="25" pageNumber="392">
<emphasis id="B91FEA8DFFF3FF8394FBFAED56B91955" bold="true" pageId="25" pageNumber="392">
Key to the
<taxonomicName id="4C6B4D1CFFF3FF839560FAEC545F196C" box="[301,514,1342,1366]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="392" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91FEA8DFFF3FF839560FAEC57F0196B" bold="true" box="[301,429,1343,1361]" italics="true" pageId="25" pageNumber="392">Pronomaea</emphasis>
species
</taxonomicName>
of the Western Palaearctic region, including Middle Asia
</emphasis>
</paragraph>
<key id="FFFA61AEFFF3FF819420FA5852191F99" lastPageId="27" lastPageNumber="394" pageId="25" pageNumber="392">
<keyStep id="309F2E3AFFF3FF809420FA5852CC1D06" lastPageId="26" lastPageNumber="393" pageId="25" pageNumber="392">
<paragraph id="8BD4369FFFF3FF839420FA5852191A6F" blockId="25.[109,1096,1418,1626]" pageId="25" pageNumber="392">
<keyLead id="309A95AAFFF3FF839420FA5852191A6F" pageId="25" pageNumber="392">
1. Antennae and legs conspicuously long and slender; antennomere X not transverse. Pronotum cordiform, distinctly narrowed posteriad; width at posterior margin 0.75-0.80 times maximal pronotal width; lateral margins smoothly sinuate in posterior half (dorsal view), posterior angles pronounced. Elytral puncturation less dense.
<specimenCount id="9D6DFD16FFF3FF83977DFA3A556019C3" box="[816,829,1513,1529]" pageId="25" pageNumber="392" type="male"></specimenCount>
: median lobe of aedeagus shaped as in figures
<quantity id="4C939B7AFFF3FF8395CEF9D657801A27" box="[387,477,1541,1565]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" metricValueMax="6.35" metricValueMin="5.842" pageId="25" pageNumber="392" unit="in" value="24.0" valueMax="25.0" valueMin="23.0">23-25 in</quantity>
ASSING (in press), with very long flagellum.
<specimenCount id="9D6DFD16FFF3FF8397E1F9D555EB1A22" box="[940,950,1542,1560]" pageId="25" pageNumber="392" type="female"></specimenCount>
: spermatheca minute and with very short duct (figure
<quantity id="4C939B7AFFF3FF839619F9F754D61A06" box="[596,651,1572,1596]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="25" pageNumber="392" unit="in" value="34.0">34 in</quantity>
ASSING (in press)).
<collectingCountry id="F37C760FFFF3FF83971BF9F755DD1A01" box="[854,896,1572,1595]" name="Iran" pageId="25" pageNumber="392">Iran</collectingCountry>
(
<figureCitation id="13502A1AFFF3FF8397C3F9F755891A06" box="[910,980,1572,1596]" captionStart="Map 5" captionStartId="24.[191,235,1184,1205]" captionTargetBox="[189,1172,467,1175]" captionTargetId="figure-284@24.[189,1172,467,1175]" captionTargetPageId="24" captionText="Map 5: Distributions of Pronomaea flavirostris SEMENOV (filled circles) and P. procerula ASSING (open circles), based on examined and literature records." figureDoi="http://doi.org/10.5281/zenodo.5364186" httpUri="https://zenodo.org/record/5364186/files/figure.png" pageId="25" pageNumber="392">Map 5</figureCitation>
). .............. ........................................................................................................
<taxonomicName id="4C6B4D1CFFF3FF839736F99052191A6F" ID-CoL="4MTB3" authority="ASSING" authorityName="ASSING" box="[891,1092,1602,1626]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="392" phylum="Arthropoda" rank="species" species="procerula">
<emphasis id="B91FEA8DFFF3FF839736F99055A91A60" bold="true" box="[891,1012,1602,1626]" italics="true" pageId="25" pageNumber="392">P. procerula</emphasis>
ASSING
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF0FF8094F3FF6352CC1D06" blockId="26.[190,1179,175,322]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FF6352CC1D06" pageId="26" pageNumber="393">
- Antennae and legs shorter; antennomere X transverse, usually approximatey 1.5 times as wide as long. Pronotum weakly tapering posteriad; width at posterior margin usually approximately 0.90-0.95 times maximal pronotal width; lateral margin not or only indistinctly sinuate in posterior half; posterior angles obtuse. Elytral puncturation denser.
<specimenCount id="9D6DFD16FFF0FF80906EFEDD526D1D24" box="[1059,1072,270,286]" pageId="26" pageNumber="393" type="male"></specimenCount>
: aedeagus of different shape and with shorter flagellum. ............................................................... B
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF0FF8094F3FE8E52CE1DC8" pageId="26" pageNumber="393">
<paragraph id="8BD4369FFFF0FF8094F3FE8E52CC1D95" blockId="26.[190,1180,349,435]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FE8E52CC1D95" pageId="26" pageNumber="393">
2. Eyes less prominent and smaller, shorter than postocular region in dorsal view (
<figureCitation id="13502A1AFFF0FF80900FFE8E52D21D4F" box="[1090,1167,349,373]" captionStart="Figs 35-43" captionStartId="17.[107,144,1545,1567]" captionTargetBox="[121,1060,455,1532]" captionTargetId="figure-177@17.[106,1089,449,1543]" captionTargetPageId="17" captionText="Figs 35-43: Pronomaea araxicola REITTER (35-40) and P. spalacis sp. n. (41-43): forebody of paratype of P. orientalisorientalis (35); median lobe of aedeagus in lateral view (36-39, 41-42) of male from &quot;Araxesthal&quot; (36), holotype of P. libanotica (37), holotype of P. orientalis (38), male from Antakya (39), and paratypes of P. spalacisspalacis (41-42); spermatheca (40, 43). Scale bars: 35: 0.5 mm; 36-43: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364178" httpUri="https://zenodo.org/record/5364178/files/figure.png" pageId="26" pageNumber="393">Fig. 35</figureCitation>
). Coloration of body on average paler, head and pronotum rufous to brown.
<specimenCount id="9D6DFD16FFF0FF80906CFEAC52731DB5" box="[1057,1070,383,399]" pageId="26" pageNumber="393" type="male"></specimenCount>
: aedeagus without projection at base of ventral process (lateral view). .......................................... C
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF0FF8094F3FE1252CE1DC8" blockId="26.[190,1179,449,504]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FE1252CE1DC8" pageId="26" pageNumber="393">- Eyes more prominent and larger, at least as long as postocular region in dorsal view. Coloration of body on average darker; head and pronotum usually dark brown to blackish. ... D</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF0FF8094F3FDC752CB1ED1" pageId="26" pageNumber="393">
<paragraph id="8BD4369FFFF0FF8094F3FDC752C51EB8" blockId="26.[190,1180,531,642]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FDC752C51EB8" pageId="26" pageNumber="393">
3.
<materialsCitation id="3B033CC2FFF0FF809541FDC552C51EB8" country="Cyprus" location="Widespread" municipality="Caucasus region" pageId="26" pageNumber="393" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D6DFD16FFF0FF809541FDC557441E1C" box="[268,281,534,550]" pageId="26" pageNumber="393" type="male"></specimenCount>
: median lobe of aedeagus at base of ventral process (i. e. near crista apicalis) not bulging; apex of ventral process obliquely truncate; flagellum shorter, darker, and stouter (
<figureCitation id="13502A1AFFF0FF80903CFDE157171E52" captionStart="Figs 35-43" captionStartId="17.[107,144,1545,1567]" captionTargetBox="[121,1060,455,1532]" captionTargetId="figure-177@17.[106,1089,449,1543]" captionTargetPageId="17" captionText="Figs 35-43: Pronomaea araxicola REITTER (35-40) and P. spalacis sp. n. (41-43): forebody of paratype of P. orientalisorientalis (35); median lobe of aedeagus in lateral view (36-39, 41-42) of male from &quot;Araxesthal&quot; (36), holotype of P. libanotica (37), holotype of P. orientalis (38), male from Antakya (39), and paratypes of P. spalacisspalacis (41-42); spermatheca (40, 43). Scale bars: 35: 0.5 mm; 36-43: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364178" httpUri="https://zenodo.org/record/5364178/files/figure.png" pageId="26" pageNumber="393">Figs 36-39</figureCitation>
).
<location id="8EB46044FFF0FF809511FD83578B1E52" LSID="urn:lsid:plazi:treatment:03C28789FFF3FF8194F8FEAD550B1FCE:8EB46044FFF0FF809511FD83578B1E52" box="[348,470,592,616]" country="Cyprus" municipality="Caucasus region" name="Widespread" pageId="26" pageNumber="393">Widespread</location>
from
<collectingMunicipality id="6BB0ACE5FFF0FF80965BFD8354E01E52" box="[534,701,592,616]" pageId="26" pageNumber="393">Caucasus region</collectingMunicipality>
to
<collectingCountry id="F37C760FFFF0FF809692FD8355711E52" box="[735,812,592,616]" name="Cyprus" pageId="26" pageNumber="393">Cyprus</collectingCountry>
,
<collectingCountry id="F37C760FFFF0FF80977AFD8355C91E52" box="[823,916,592,616]" name="Lebanon" pageId="26" pageNumber="393">Lebanon</collectingCountry>
, and
<collectingCountry id="F37C760FFFF0FF809786FD8255A81E52" box="[971,1013,593,616]" name="Iran" pageId="26" pageNumber="393">Iran</collectingCountry>
(
<figureCitation id="13502A1AFFF0FF80904EFD8352161E52" box="[1027,1099,592,616]" captionStart="Map 3" captionStartId="18.[193,237,1569,1590]" captionTargetBox="[193,1176,829,1562]" captionTargetId="figure-392@18.[193,1176,829,1562]" captionTargetPageId="18" captionText="Map 3: Distributions of Pronomaea araxicola REITTERREITTER (filled circles) and P. spalacis sp. n. (open circles), based on examined and literature records. The unconfirmed record of P. araxicola from Bolu (Turkey) is omitted." figureDoi="http://doi.org/10.5281/zenodo.5364180" httpUri="https://zenodo.org/record/5364180/files/figure.png" pageId="26" pageNumber="393">Map 3</figureCitation>
). ........ .......................................................................................................
<taxonomicName id="4C6B4D1CFFF0FF80978EFDA352C51EB8" ID-CoL="789PV" authority="REITTER" authorityName="REITTER" box="[963,1176,624,642]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="393" phylum="Arthropoda" rank="species" species="araxicola">
<emphasis id="B91FEA8DFFF0FF80978EFDA352671EB8" bold="true" box="[963,1082,624,642]" italics="true" pageId="26" pageNumber="393">P. araxicola</emphasis>
REITTER
</taxonomicName>
</materialsCitation>
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF0FF8094F3FD7252CB1ED1" blockId="26.[190,1180,661,748]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FD7252CB1ED1" pageId="26" pageNumber="393">
-
<specimenCount id="9D6DFD16FFF0FF809541FD4B57441E92" box="[268,281,664,680]" pageId="26" pageNumber="393" type="male"></specimenCount>
: median lobe of aedeagus at base of ventral process (i. e. near crista apicalis) somewhat bulging; apex of ventral process rounded; flagellum longer and more slender (
<figureCitation id="13502A1AFFF0FF809050FD6752D21EF6" box="[1053,1167,692,716]" captionStart="Figs 35-43" captionStartId="17.[107,144,1545,1567]" captionTargetBox="[121,1060,455,1532]" captionTargetId="figure-177@17.[106,1089,449,1543]" captionTargetPageId="17" captionText="Figs 35-43: Pronomaea araxicola REITTER (35-40) and P. spalacis sp. n. (41-43): forebody of paratype of P. orientalisorientalis (35); median lobe of aedeagus in lateral view (36-39, 41-42) of male from &quot;Araxesthal&quot; (36), holotype of P. libanotica (37), holotype of P. orientalis (38), male from Antakya (39), and paratypes of P. spalacisspalacis (41-42); spermatheca (40, 43). Scale bars: 35: 0.5 mm; 36-43: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364178" httpUri="https://zenodo.org/record/5364178/files/figure.png" pageId="26" pageNumber="393">Figs 41-42</figureCitation>
).
<collectingCountry id="F37C760FFFF0FF809547FD00571E1ED1" box="[266,323,723,747]" name="Israel" pageId="26" pageNumber="393">Israel</collectingCountry>
. (
<figureCitation id="13502A1AFFF0FF80951EFD0057C51ED1" box="[339,408,723,747]" captionStart="Map 3" captionStartId="18.[193,237,1569,1590]" captionTargetBox="[193,1176,829,1562]" captionTargetId="figure-392@18.[193,1176,829,1562]" captionTargetPageId="18" captionText="Map 3: Distributions of Pronomaea araxicola REITTERREITTER (filled circles) and P. spalacis sp. n. (open circles), based on examined and literature records. The unconfirmed record of P. araxicola from Bolu (Turkey) is omitted." figureDoi="http://doi.org/10.5281/zenodo.5364180" httpUri="https://zenodo.org/record/5364180/files/figure.png" pageId="26" pageNumber="393">Map 3</figureCitation>
). .....................................................................................
<taxonomicName id="4C6B4D1CFFF0FF8097B9FD0052071ED6" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[1012,1114,722,748]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="393" phylum="Arthropoda" rank="species" species="spalacisspalacis" status="sp. nov.">
<emphasis id="B91FEA8DFFF0FF8097B9FD0052071ED6" bold="true" box="[1012,1114,722,748]" italics="true" pageId="26" pageNumber="393">P. spalacisspalacis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57F6FFF0FF80902DFD0A52CB1ED1" box="[1120,1174,729,747]" pageId="26" pageNumber="393" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF0FF8094F3FCD452CF1FE3" pageId="26" pageNumber="393">
<paragraph id="8BD4369FFFF0FF8094F3FCD452CD1F4C" blockId="26.[190,1179,774,886]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FCD452CD1F4C" pageId="26" pageNumber="393">
4.
<specimenCount id="9D6DFD16FFF0FF809541FCDA57441F23" box="[268,281,777,793]" pageId="26" pageNumber="393" type="male"></specimenCount>
: median lobe of aedeagus at base of ventral process (near crista apicalis) without projection or transverse carina (best seen in lateral view).
<specimenCount id="9D6DFD16FFF0FF80976FFCF555711F02" box="[802,812,806,824]" pageId="26" pageNumber="393" type="female"></specimenCount>
: spermatheca shaped as in
<figureCitation id="13502A1AFFF0FF809001FCF652CA1F07" box="[1100,1175,805,829]" captionStart="Figs 44-51" captionStartId="21.[112,149,1338,1360]" captionTargetBox="[116,1075,184,1323]" captionTargetId="figure-76@21.[111,1092,174,1336]" captionTargetPageId="21" captionText="Figs 44-51: Pronomaea anatolica FAGEL (44-47), P. khnzoriani SEMENOV (48-50), and P. flavirostris SEMENOV (51): median lobe of aedeagus in lateral view (44-46, 48-51) of holotype (44) and of males from Muğla (45-46), Iran (48-49), Amasya (50), and Tajikistan (51); spermatheca (47). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364182" httpUri="https://zenodo.org/record/5364182/files/figure.png" pageId="26" pageNumber="393">Fig. 47</figureCitation>
, minute and with extremely short duct. Eastern Mediterranean, Caucasus region, Middle Asia. ............................................................................................................................. E
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF0FF8094F3FC4752CF1FE3" blockId="26.[190,1179,904,990]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FC4752CF1FE3" pageId="26" pageNumber="393">
-
<specimenCount id="9D6DFD16FFF0FF809541FC5857441FA1" box="[268,281,907,923]" pageId="26" pageNumber="393" type="male"></specimenCount>
: median lobe of aedeagus at base of ventral process (near crista apicalis) with more or less pronounced projection or carinae.
<specimenCount id="9D6DFD16FFF0FF8096D0FC7B54FA1F80" box="[669,679,936,954]" pageId="26" pageNumber="393" type="female"></specimenCount>
: spermatheca less minute; duct distinctly longer than capsule. ................................................................................................................ G
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF0FF8094F2FC2952D21855" pageId="26" pageNumber="393">
<paragraph id="8BD4369FFFF0FF8094F2FC2952C51811" blockId="26.[191,1179,1017,1072]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F2FC2952C51811" pageId="26" pageNumber="393">
5.
<materialsCitation id="3B033CC2FFF0FF809541FC2F52C51811" country="Tajikistan" location="Tajikistan" pageId="26" pageNumber="393" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D6DFD16FFF0FF809541FC2F57441836" box="[268,281,1020,1036]" pageId="26" pageNumber="393" type="male"></specimenCount>
: median lobe of aedeagus shaped as in
<figureCitation id="13502A1AFFF0FF8096FDFC2A54A7182B" box="[688,762,1017,1041]" captionStart="Figs 44-51" captionStartId="21.[112,149,1338,1360]" captionTargetBox="[116,1075,184,1323]" captionTargetId="figure-76@21.[111,1092,174,1336]" captionTargetPageId="21" captionText="Figs 44-51: Pronomaea anatolica FAGEL (44-47), P. khnzoriani SEMENOV (48-50), and P. flavirostris SEMENOV (51): median lobe of aedeagus in lateral view (44-46, 48-51) of holotype (44) and of males from Muğla (45-46), Iran (48-49), Amasya (50), and Tajikistan (51); spermatheca (47). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364182" httpUri="https://zenodo.org/record/5364182/files/figure.png" pageId="26" pageNumber="393">Fig. 51</figureCitation>
, ventral process almost straight in lateral view.
<collectingCountry id="F37C760FFFF0FF80950CFBCB57F5180A" box="[321,424,1048,1072]" name="Tajikistan" pageId="26" pageNumber="393">Tajikistan</collectingCountry>
,
<collectingCountry id="F37C760FFFF0FF8095FEFBCB5473180A" box="[435,558,1048,1072]" name="Afghanistan" pageId="26" pageNumber="393">Afghanistan</collectingCountry>
(
<figureCitation id="13502A1AFFF0FF809671FBCB54DE180A" box="[572,643,1048,1072]" captionStart="Map 5" captionStartId="24.[191,235,1184,1205]" captionTargetBox="[189,1172,467,1175]" captionTargetId="figure-284@24.[189,1172,467,1175]" captionTargetPageId="24" captionText="Map 5: Distributions of Pronomaea flavirostris SEMENOV (filled circles) and P. procerula ASSING (open circles), based on examined and literature records." figureDoi="http://doi.org/10.5281/zenodo.5364186" httpUri="https://zenodo.org/record/5364186/files/figure.png" pageId="26" pageNumber="393">Map 5</figureCitation>
). .......................................
<taxonomicName id="4C6B4D1CFFF0FF8097E7FBCB52C51811" ID-CoL="77XNY" authority="SEMENOV" authorityName="SEMENOV" box="[938,1176,1047,1071]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="393" phylum="Arthropoda" rank="species" species="flavirostris">
<emphasis id="B91FEA8DFFF0FF8097E7FBCB52721815" bold="true" box="[938,1071,1047,1071]" italics="true" pageId="26" pageNumber="393">P. flavirostris</emphasis>
SEMENOV
</taxonomicName>
</materialsCitation>
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF0FF8094F3FB9952D21855" blockId="26.[190,1173,1086,1135]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FB9952D21855" pageId="26" pageNumber="393">
-
<specimenCount id="9D6DFD16FFF0FF809541FB925744186B" box="[268,281,1089,1105]" pageId="26" pageNumber="393" type="male"></specimenCount>
: ventral process of aedeagus strongly sinuate in lateral view. Distribution different. ..... ..................................................................................................................................... F
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF0FF8094F3FB4252C5191F" pageId="26" pageNumber="393">
<paragraph id="8BD4369FFFF0FF8094F3FB4252C518DA" blockId="26.[190,1180,1168,1248]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FB4252C518DA" pageId="26" pageNumber="393">
6.
<materialsCitation id="3B033CC2FFF0FF809541FB4052C518DA" country="Turkey, Armenia" location="Caucasus region" pageId="26" pageNumber="393" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D6DFD16FFF0FF809541FB4057441899" box="[268,281,1171,1187]" pageId="26" pageNumber="393" type="male"></specimenCount>
: apical internal structure of aedeagus very slender and evenly bent (
<figureCitation id="13502A1AFFF0FF8097F1FB4352711892" box="[956,1068,1168,1192]" captionStart="Figs 44-51" captionStartId="21.[112,149,1338,1360]" captionTargetBox="[116,1075,184,1323]" captionTargetId="figure-76@21.[111,1092,174,1336]" captionTargetPageId="21" captionText="Figs 44-51: Pronomaea anatolica FAGEL (44-47), P. khnzoriani SEMENOV (48-50), and P. flavirostris SEMENOV (51): median lobe of aedeagus in lateral view (44-46, 48-51) of holotype (44) and of males from Muğla (45-46), Iran (48-49), Amasya (50), and Tajikistan (51); spermatheca (47). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364182" httpUri="https://zenodo.org/record/5364182/files/figure.png" pageId="26" pageNumber="393">Figs 48-50</figureCitation>
).
<location id="8EB46044FFF0FF809070FB42571718FD" LSID="urn:lsid:plazi:treatment:03C28789FFF3FF8194F8FEAD550B1FCE:8EB46044FFF0FF809070FB42571718FD" country="Turkey, Armenia" name="Caucasus region" pageId="26" pageNumber="393">Caucasus region</location>
to northern
<collectingCountry id="F37C760FFFF0FF809580FB7C542618FD" box="[461,635,1199,1223]" isEnumeration="true" pageId="26" pageNumber="393">Turkey, Armenia</collectingCountry>
,
<collectingCountry id="F37C760FFFF0FF8096C8FB6354EF18FD" box="[645,690,1200,1223]" name="Iran" pageId="26" pageNumber="393">Iran</collectingCountry>
, and
<collectingCountry id="F37C760FFFF0FF8096A4FB7C552518FD" box="[745,888,1199,1223]" name="Turkmenistan" pageId="26" pageNumber="393">Turkmenistan</collectingCountry>
(
<figureCitation id="13502A1AFFF0FF8097CBFB7C559318FD" box="[902,974,1199,1223]" captionStart="Map 4" captionStartId="22.[195,239,844,865]" captionTargetBox="[195,1178,174,836]" captionTargetId="figure-325@22.[195,1178,174,836]" captionTargetPageId="22" captionText="Map 4: Distributions of Pronomaea anatolica FAGEL (filled circles) and P. khnzoriani SEMENOV (open circles), based on examined and literature records." figureDoi="http://doi.org/10.5281/zenodo.5364184" httpUri="https://zenodo.org/record/5364184/files/figure.png" pageId="26" pageNumber="393">Map 4</figureCitation>
). ........................... ..................................................................................................
<taxonomicName id="4C6B4D1CFFF0FF8097E8FB1D52C518DA" ID-CoL="789NX" authority="SEMENOV" authorityName="SEMENOV" box="[933,1176,1230,1248]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="393" phylum="Arthropoda" rank="species" species="khnzoriani">
<emphasis id="B91FEA8DFFF0FF8097E8FB1D527218DA" bold="true" box="[933,1071,1230,1248]" italics="true" pageId="26" pageNumber="393">P. khnzoriani</emphasis>
SEMENOV
</taxonomicName>
</materialsCitation>
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF0FF8094F3FAD352C5191F" blockId="26.[190,1179,1268,1323]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FAD352C5191F" pageId="26" pageNumber="393">
-
<specimenCount id="9D6DFD16FFF0FF809541FB245744193D" box="[268,281,1271,1287]" pageId="26" pageNumber="393" type="male"></specimenCount>
: apical internal structure of aedeagus of different shape (
<figureCitation id="13502A1AFFF0FF80972CFB27558E1936" box="[865,979,1268,1292]" captionStart="Figs 44-51" captionStartId="21.[112,149,1338,1360]" captionTargetBox="[116,1075,184,1323]" captionTargetId="figure-76@21.[111,1092,174,1336]" captionTargetPageId="21" captionText="Figs 44-51: Pronomaea anatolica FAGEL (44-47), P. khnzoriani SEMENOV (48-50), and P. flavirostris SEMENOV (51): median lobe of aedeagus in lateral view (44-46, 48-51) of holotype (44) and of males from Muğla (45-46), Iran (48-49), Amasya (50), and Tajikistan (51); spermatheca (47). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364182" httpUri="https://zenodo.org/record/5364182/files/figure.png" pageId="26" pageNumber="393">Figs 44-51</figureCitation>
). SE-Bulgaria, SW- and S-Anatolia (
<figureCitation id="13502A1AFFF0FF8095FDFAC057A51911" box="[432,504,1299,1323]" captionStart="Map 4" captionStartId="22.[195,239,844,865]" captionTargetBox="[195,1178,174,836]" captionTargetId="figure-325@22.[195,1178,174,836]" captionTargetPageId="22" captionText="Map 4: Distributions of Pronomaea anatolica FAGEL (filled circles) and P. khnzoriani SEMENOV (open circles), based on examined and literature records." figureDoi="http://doi.org/10.5281/zenodo.5364184" httpUri="https://zenodo.org/record/5364184/files/figure.png" pageId="26" pageNumber="393">Map 4</figureCitation>
). ...................................................................
<taxonomicName id="4C6B4D1CFFF0FF809791FAC052C5191F" ID-CoL="77Y27" authority="FAGEL" authorityName="Fagel" authorityYear="1969" box="[988,1176,1299,1317]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="393" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B91FEA8DFFF0FF809791FAC05209191F" bold="true" box="[988,1108,1299,1317]" italics="true" pageId="26" pageNumber="393">P. anatolica</emphasis>
FAGEL
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF0FF8094F3FA9452C91A4E" pageId="26" pageNumber="393">
<paragraph id="8BD4369FFFF0FF8094F3FA9452CA19C8" blockId="26.[190,1180,1350,1528]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3FA9452CA19C8" pageId="26" pageNumber="393">
7. Pronotum larger in relation to head, usually at least 1.3 times as wide as head; pronotal pubescence shorter and usually more or less depressed. Antennae usually uniformly blackish, with the basal antennomeres at most only slightly paler than apical antennomeres.
<specimenCount id="9D6DFD16FFF0FF8090C9FA5552CC19AC" box="[1156,1169,1414,1430]" pageId="26" pageNumber="393" type="male"></specimenCount>
: median lobe of aedeagus shaped as in
<figureCitation id="13502A1AFFF0FF8096C3FA7154B91980" box="[654,740,1442,1466]" captionStart="Figs 1-8" captionStartId="3.[110,147,1330,1352]" captionTargetBox="[112,1085,177,1307]" captionTargetId="figure-176@3.[110,1093,174,1326]" captionTargetPageId="3" captionText="Figs 1-8: Pronomaea rostrata ERICHSON: median lobe of aedeagus in lateral view (1-6) of males from Germany (1), southern Spain (2), Ukraine (3), Poland (4), Corsica (5), and Tunisia (6); apical part of median lobe of aedeagus in ventral view (7); spermatheca (8). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364166" httpUri="https://zenodo.org/record/5364166/files/figure.png" pageId="26" pageNumber="393">Figs 1-7</figureCitation>
.
<specimenCount id="9D6DFD16FFF0FF8096B9FA7054A3198F" box="[756,766,1443,1461]" pageId="26" pageNumber="393" type="female"></specimenCount>
: spermatheca with longer duct (
<figureCitation id="13502A1AFFF0FF80901DFA7152D31980" box="[1104,1166,1442,1466]" captionStart="Figs 1-8" captionStartId="3.[110,147,1330,1352]" captionTargetBox="[112,1085,177,1307]" captionTargetId="figure-176@3.[110,1093,174,1326]" captionTargetPageId="3" captionText="Figs 1-8: Pronomaea rostrata ERICHSON: median lobe of aedeagus in lateral view (1-6) of males from Germany (1), southern Spain (2), Ukraine (3), Poland (4), Corsica (5), and Tunisia (6); apical part of median lobe of aedeagus in ventral view (7); spermatheca (8). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364166" httpUri="https://zenodo.org/record/5364166/files/figure.png" pageId="26" pageNumber="393">Fig. 8</figureCitation>
). Distribution Atlanto-Mediterranean, from NW-Africa and the Iberian Peninsula to
<collectingCountry id="F37C760FFFF0FF80901CFA1252C119EC" box="[1105,1180,1473,1494]" name="Belarus" pageId="26" pageNumber="393">Belarus</collectingCountry>
(
<figureCitation id="13502A1AFFF0FF80955FFA33570919C2" box="[274,340,1504,1528]" captionStart="Map 1" captionStartId="4.[191,235,1572,1593]" captionTargetBox="[191,1174,756,1564]" captionTargetId="figure-389@4.[191,1174,756,1565]" captionTargetPageId="4" captionText="Map 1: Distributions of Pronomaea rostrata ERICHSON (filled circles: examined records; open circles: selected plausible literature records by FAGEL (1969) and FAUVEL (1902)), P. sardoasardoa sp. n. (open squares), and P. sicula sp. n. (filled squares)." figureDoi="http://doi.org/10.5281/zenodo.5364168" httpUri="https://zenodo.org/record/5364168/files/figure.png" pageId="26" pageNumber="393">Map 1</figureCitation>
). .......................................................................................
<taxonomicName id="4C6B4D1CFFF0FF8097F3FA3352CA19C8" ID-CoL="4MTB7" authority="ERICHSON" authorityName="ERICHSON" box="[958,1175,1504,1522]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="393" phylum="Arthropoda" rank="species" species="rostrata">
<emphasis id="B91FEA8DFFF0FF8097F3FA33527B19C8" bold="true" box="[958,1062,1504,1522]" italics="true" pageId="26" pageNumber="393">P. rostrata</emphasis>
ERICHSON
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF0FF8094F3F9D552C91A4E" blockId="26.[190,1180,1542,1652]" pageId="26" pageNumber="393">
<keyLead id="309A95AAFFF0FF8094F3F9D552C91A4E" pageId="26" pageNumber="393">
- Pronotum smaller in relation to head, usually at most 1.3 times as wide as head; pronotal pubescence longer and usually suberect. Basal antennomeres often of paler coloration than apical antennomeres.
<specimenCount id="9D6DFD16FFF0FF8095A7F99557AA1A6C" box="[490,503,1606,1622]" pageId="26" pageNumber="393" type="male"></specimenCount>
: median lobe of aedeagus of different shape.
<specimenCount id="9D6DFD16FFF0FF809781F997558B1A6C" box="[972,982,1604,1622]" pageId="26" pageNumber="393" type="female"></specimenCount>
: spermatheca with shorter duct. ................................................................................................................ H
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF1FF819426FF6352651D1D" pageId="27" pageNumber="394">
<paragraph id="8BD4369FFFF1FF819426FF63521E1CDC" blockId="27.[107,1096,175,230]" pageId="27" pageNumber="394">
<keyLead id="309A95AAFFF1FF819426FF63521E1CDC" pageId="27" pageNumber="394">
8.
<materialsCitation id="3B033CC2FFF1FF8194F4FF6152621CDC" location="Crete" pageId="27" pageNumber="394" specimenCount="1" specimenCount-male="1" stateProvince="Kriti">
<specimenCount id="9D6DFD16FFF1FF8194F4FF61569B1CF8" box="[185,198,178,194]" pageId="27" pageNumber="394" type="male"></specimenCount>
: median lobe of aedeagus at base of ventral process with fine carina very close to crista apicalis (
<figureCitation id="13502A1AFFF1FF819540FF1D57DC1CDC" box="[269,385,206,230]" captionStart="Figs 26-34" captionStartId="15.[110,147,1355,1377]" captionTargetBox="[110,1086,182,1337]" captionTargetId="figure-108@15.[110,1093,174,1352]" captionTargetPageId="15" captionText="Figs 26-34: Pronomaea wunderlei sp. n. (26-28), P. sicula sp. n. (29-31), and P. sardoa sp. n. (32-34): median lobe of aedeagus in lateral view (26-27, 29-30, 32-33) of holotypes (26, 29, 32) and paratypes (27, 30, 33); apical part of median lobe of aedeagus in ventral view (28, 31, 34). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364176" httpUri="https://zenodo.org/record/5364176/files/figure.png" pageId="27" pageNumber="394">Figs 26-28</figureCitation>
).
<collectingRegion id="49AFF87DFFF1FF8195DFFF1D57971CDC" box="[402,458,206,230]" country="Greece" name="Kriti" pageId="27" pageNumber="394">Crete</collectingRegion>
(
<figureCitation id="13502A1AFFF1FF819594FF1D547C1CDC" box="[473,545,206,230]" captionStart="Map 2" captionStartId="10.[197,241,1570,1591]" captionTargetBox="[195,1178,841,1563]" captionTargetId="figure-419@10.[195,1178,841,1563]" captionTargetPageId="10" captionText="Map 2: Distributions of Pronomaea picea HEERHEER (filled circles), P. corsicana sp. n. (open circles), and P. wunderleiwunderlei sp. n. (squares), based on examined records. The records of doubtful identity from Malta are omitted." figureDoi="http://doi.org/10.5281/zenodo.5364172" httpUri="https://zenodo.org/record/5364172/files/figure.png" pageId="27" pageNumber="394">Map 2</figureCitation>
). .................................................
<taxonomicName id="4C6B4D1CFFF1FF8197CAFF1D525B1CDA" ID-CoL="4MTBL" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[903,1030,206,224]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="394" phylum="Arthropoda" rank="species" species="wunderlei" status="sp. nov.">
<emphasis id="B91FEA8DFFF1FF8197CAFF1D525B1CDA" bold="true" box="[903,1030,206,224]" italics="true" pageId="27" pageNumber="394">P. wunderlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57F6FFF1FF819040FF0752621CDC" box="[1037,1087,212,230]" pageId="27" pageNumber="394" rank="species">sp. n</taxonomicNameLabel>
</materialsCitation>
.
</keyLead>
</paragraph>
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<keyLead id="309A95AAFFF1FF819426FF2C52651D1D" pageId="27" pageNumber="394">
-
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: median lobe of aedeagus at base of ventral process with more prominent projection in different position. Distribution different. .....................................................................I
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF1FF819426FE94521F1D86" pageId="27" pageNumber="394">
<paragraph id="8BD4369FFFF1FF819426FE9452191D4D" blockId="27.[107,1096,326,381]" pageId="27" pageNumber="394">
<keyLead id="309A95AAFFF1FF819426FE9452191D4D" pageId="27" pageNumber="394">
9.
<materialsCitation id="3B033CC2FFF1FF8194F4FE9A55131D47" location="Sardinia and Sicily" pageId="27" pageNumber="394" specimenCount="1" specimenCount-male="1" stateProvince="Sardinia and Sicily">
<specimenCount id="9D6DFD16FFF1FF8194F4FE9A569B1D63" box="[185,198,329,345]" pageId="27" pageNumber="394" type="male"></specimenCount>
: median lobe of aedeagus at base of ventral process with less pronounced and distinctly angular projection (
<figureCitation id="13502A1AFFF1FF819532FEB657AF1D47" box="[383,498,357,381]" captionStart="Figs 26-34" captionStartId="15.[110,147,1355,1377]" captionTargetBox="[110,1086,182,1337]" captionTargetId="figure-108@15.[110,1093,174,1352]" captionTargetPageId="15" captionText="Figs 26-34: Pronomaea wunderlei sp. n. (26-28), P. sicula sp. n. (29-31), and P. sardoa sp. n. (32-34): median lobe of aedeagus in lateral view (26-27, 29-30, 32-33) of holotypes (26, 29, 32) and paratypes (27, 30, 33); apical part of median lobe of aedeagus in ventral view (28, 31, 34). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364176" httpUri="https://zenodo.org/record/5364176/files/figure.png" pageId="27" pageNumber="394">Figs 30-34</figureCitation>
). Species from
<collectingRegion id="49AFF87DFFF1FF8196C0FEB655131D47" box="[653,846,357,381]" isEnumeration="true" pageId="27" pageNumber="394">Sardinia and Sicily</collectingRegion>
</materialsCitation>
. .............................. 10
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF1FF819426FE44521F1D86" blockId="27.[107,1096,395,450]" pageId="27" pageNumber="394">
<keyLead id="309A95AAFFF1FF819426FE44521F1D86" pageId="27" pageNumber="394">
-
<specimenCount id="9D6DFD16FFF1FF8194F4FE5D569B1DA4" box="[185,198,398,414]" pageId="27" pageNumber="394" type="male"></specimenCount>
: median lobe of aedeagus at base of ventral process with more massive and usually not distinctly angular projection. Distribution different. .................................................. 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF1FF819420FE0D521E1EAD" pageId="27" pageNumber="394">
<paragraph id="8BD4369FFFF1FF819420FE0D521E1E09" blockId="27.[109,1096,477,563]" pageId="27" pageNumber="394">
<keyLead id="309A95AAFFF1FF819420FE0D521E1E09" pageId="27" pageNumber="394">
10.
<materialsCitation id="3B033CC2FFF1FF8194F4FE3352621E09" location="Sardinia" pageId="27" pageNumber="394" specimenCount="1" specimenCount-male="1" stateProvince="Sardegna">
<specimenCount id="9D6DFD16FFF1FF8194F4FE33569B1DCA" box="[185,198,480,496]" pageId="27" pageNumber="394" type="male"></specimenCount>
: median lobe of aedeagus smaller, approximately
<quantity id="4C939B7AFFF1FF8196E2FE0E55501DCF" box="[687,781,477,501]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="27" pageNumber="394" unit="mm" value="0.38">0.38 mm</quantity>
long; flagellum shorter; angular projection at base of ventral process more prominent (
<figureCitation id="13502A1AFFF1FF8196ADFE2F550E1E2E" box="[736,851,508,532]" captionStart="Figs 26-34" captionStartId="15.[110,147,1355,1377]" captionTargetBox="[110,1086,182,1337]" captionTargetId="figure-108@15.[110,1093,174,1352]" captionTargetPageId="15" captionText="Figs 26-34: Pronomaea wunderlei sp. n. (26-28), P. sicula sp. n. (29-31), and P. sardoa sp. n. (32-34): median lobe of aedeagus in lateral view (26-27, 29-30, 32-33) of holotypes (26, 29, 32) and paratypes (27, 30, 33); apical part of median lobe of aedeagus in ventral view (28, 31, 34). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364176" httpUri="https://zenodo.org/record/5364176/files/figure.png" pageId="27" pageNumber="394">Figs 32-34</figureCitation>
).
<collectingRegion id="49AFF87DFFF1FF819728FE2F55E41E2E" box="[869,953,508,532]" country="Italy" name="Sardegna" pageId="27" pageNumber="394">Sardinia</collectingRegion>
(
<figureCitation id="13502A1AFFF1FF81978AFE2F52511E2E" box="[967,1036,508,532]" captionStart="Map 1" captionStartId="4.[191,235,1572,1593]" captionTargetBox="[191,1174,756,1564]" captionTargetId="figure-389@4.[191,1174,756,1565]" captionTargetPageId="4" captionText="Map 1: Distributions of Pronomaea rostrata ERICHSON (filled circles: examined records; open circles: selected plausible literature records by FAGEL (1969) and FAUVEL (1902)), P. sardoasardoa sp. n. (open squares), and P. sicula sp. n. (filled squares)." figureDoi="http://doi.org/10.5281/zenodo.5364168" httpUri="https://zenodo.org/record/5364168/files/figure.png" pageId="27" pageNumber="394">Map 1</figureCitation>
). ..... ...............................................................................................................
<taxonomicName id="4C6B4D1CFFF1FF8197E6FDC8525B1E17" ID-CoL="6W9P2" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[939,1030,539,557]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="394" phylum="Arthropoda" rank="species" species="sardoa" status="sp. nov.">
<emphasis id="B91FEA8DFFF1FF8197E6FDC8525B1E17" bold="true" box="[939,1030,539,557]" italics="true" pageId="27" pageNumber="394">P. sardoa</emphasis>
</taxonomicName>
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</materialsCitation>
.
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF1FF819426FD9F521E1EAD" blockId="27.[107,1096,577,663]" pageId="27" pageNumber="394">
<keyLead id="309A95AAFFF1FF819426FD9F521E1EAD" pageId="27" pageNumber="394">
-
<materialsCitation id="3B033CC2FFF1FF8194F4FD9752621EAD" location="Sicily" pageId="27" pageNumber="394" specimenCount="1" specimenCount-male="1" stateProvince="Sicilia">
<specimenCount id="9D6DFD16FFF1FF8194F4FD97569B1E6E" box="[185,198,580,596]" pageId="27" pageNumber="394" type="male"></specimenCount>
: median lobe of aedeagus larger, approximately
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long; flagellum longer; angular projection at base of ventral process less prominent (
<figureCitation id="13502A1AFFF1FF819683FD8C551D1E42" box="[718,832,607,632]" captionStart="Figs 26-34" captionStartId="15.[110,147,1355,1377]" captionTargetBox="[110,1086,182,1337]" captionTargetId="figure-108@15.[110,1093,174,1352]" captionTargetPageId="15" captionText="Figs 26-34: Pronomaea wunderlei sp. n. (26-28), P. sicula sp. n. (29-31), and P. sardoa sp. n. (32-34): median lobe of aedeagus in lateral view (26-27, 29-30, 32-33) of holotypes (26, 29, 32) and paratypes (27, 30, 33); apical part of median lobe of aedeagus in ventral view (28, 31, 34). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364176" httpUri="https://zenodo.org/record/5364176/files/figure.png" pageId="27" pageNumber="394">Figs 29-31</figureCitation>
).
<collectingRegion id="49AFF87DFFF1FF81971EFD8C55D71E4D" box="[851,906,607,631]" country="Italy" name="Sicilia" pageId="27" pageNumber="394">Sicily</collectingRegion>
(
<figureCitation id="13502A1AFFF1FF8197D5FDB355811E42" box="[920,988,608,632]" captionStart="Map 1" captionStartId="4.[191,235,1572,1593]" captionTargetBox="[191,1174,756,1564]" captionTargetId="figure-389@4.[191,1174,756,1565]" captionTargetPageId="4" captionText="Map 1: Distributions of Pronomaea rostrata ERICHSON (filled circles: examined records; open circles: selected plausible literature records by FAGEL (1969) and FAUVEL (1902)), P. sardoasardoa sp. n. (open squares), and P. sicula sp. n. (filled squares)." figureDoi="http://doi.org/10.5281/zenodo.5364168" httpUri="https://zenodo.org/record/5364168/files/figure.png" pageId="27" pageNumber="394">Map 1</figureCitation>
). ............ .................................................................................................................
<taxonomicName id="4C6B4D1CFFF1FF8197F9FDAC525B1EAB" ID-CoL="4MTBB" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[948,1030,639,657]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="394" phylum="Arthropoda" rank="species" species="sicula" status="sp. nov.">
<emphasis id="B91FEA8DFFF1FF8197F9FDAC525B1EAB" bold="true" box="[948,1030,639,657]" italics="true" pageId="27" pageNumber="394">P. sicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57F6FFF1FF819040FD5652621EAD" box="[1037,1087,645,663]" pageId="27" pageNumber="394" rank="species">sp. n</taxonomicNameLabel>
</materialsCitation>
.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E3AFFF1FF819420FD6152191F99" pageId="27" pageNumber="394">
<paragraph id="8BD4369FFFF1FF819420FD61521E1F1C" blockId="27.[109,1096,690,806]" pageId="27" pageNumber="394">
<keyLead id="309A95AAFFF1FF819420FD61521E1F1C" pageId="27" pageNumber="394">
11. Antenna usually of uniformly blackish coloration, basal antennomeres at most only slightly paler than apical antennomeres.
<specimenCount id="9D6DFD16FFF1FF8195B6FD0054551ED9" box="[507,520,723,739]" pageId="27" pageNumber="394" type="male"></specimenCount>
: projection at base of ventral process of aedeagus rounded to nearly angular; apex of ventral process more slender in lateral view (
<figureCitation id="13502A1AFFF1FF819735FD3C55B41F3D" box="[888,1001,751,775]" captionStart="Figs 18-25" captionStartId="13.[110,147,1297,1319]" captionTargetBox="[110,1081,175,1281]" captionTargetId="figure-111@13.[108,1091,174,1291]" captionTargetPageId="13" captionText="Figs 18-25: Pronomaea corsicana sp. n.: median lobe of aedeagus in lateral view (18-23) of males from various Corsican localities; median lobe of aedeagus in ventral view (24); spermatheca (25). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364174" httpUri="https://zenodo.org/record/5364174/files/figure.png" pageId="27" pageNumber="394">Figs 18-24</figureCitation>
).
<collectingCountry id="F37C760FFFF1FF8197B6FD3C52151F3D" box="[1019,1096,751,775]" name="France" pageId="27" pageNumber="394">Corsica</collectingCountry>
(
<figureCitation id="13502A1AFFF1FF8194F3FCDD57581F1C" box="[190,261,782,806]" captionStart="Map 2" captionStartId="10.[197,241,1570,1591]" captionTargetBox="[195,1178,841,1563]" captionTargetId="figure-419@10.[195,1178,841,1563]" captionTargetPageId="10" captionText="Map 2: Distributions of Pronomaea picea HEERHEER (filled circles), P. corsicana sp. n. (open circles), and P. wunderleiwunderlei sp. n. (squares), based on examined records. The records of doubtful identity from Malta are omitted." figureDoi="http://doi.org/10.5281/zenodo.5364172" httpUri="https://zenodo.org/record/5364172/files/figure.png" pageId="27" pageNumber="394">Map 2</figureCitation>
). ............................................................................................
<taxonomicName id="4C6B4D1CFFF1FF8197C2FCDD525B1F1A" ID-CoL="789NW" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[911,1030,782,800]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="394" phylum="Arthropoda" rank="species" species="corsicana" status="sp. nov.">
<emphasis id="B91FEA8DFFF1FF8197C2FCDD525B1F1A" bold="true" box="[911,1030,782,800]" italics="true" pageId="27" pageNumber="394">P. corsicana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57F6FFF1FF819040FCC7521E1F1C" box="[1037,1091,788,806]" pageId="27" pageNumber="394" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BD4369FFFF1FF819426FCE752191F99" blockId="27.[107,1096,820,936]" pageId="27" pageNumber="394">
<keyLead id="309A95AAFFF1FF819426FCE752191F99" pageId="27" pageNumber="394">
- Basal antennomeres usually somewhat paler than remainder of antenna, yellowish brown to brown.
<specimenCount id="9D6DFD16FFF1FF81956AFC8657691F5F" box="[295,308,853,869]" pageId="27" pageNumber="394" type="male"></specimenCount>
: projection at base of ventral process more or less truncate apically; apex of ventral process less slender (
<figureCitation id="13502A1AFFF1FF81958EFCA2547E1FB3" box="[451,547,881,905]" captionStart="Figs 9-16" captionStartId="9.[109,146,1577,1599]" captionTargetBox="[118,1047,461,1559]" captionTargetId="figure-168@9.[108,1089,443,1575]" captionTargetPageId="9" captionText="Figs 9-16: Pronomaea picea HEER: median lobe of aedeagus in lateral view (9-13) of males from France (9-10), Italy (11), Hungary (12), and Greece (13); apical part of median lobe of aedeagus in ventral view (14-15); spermatheca (16). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364170" httpUri="https://zenodo.org/record/5364170/files/figure.png" pageId="27" pageNumber="394">Figs 9-15</figureCitation>
). Widespread Ponto-Mediterranean species, distributed from the Balkans to western
<collectingCountry id="F37C760FFFF1FF819591FC43547D1F9D" box="[476,544,912,935]" name="France" pageId="27" pageNumber="394">France</collectingCountry>
(
<figureCitation id="13502A1AFFF1FF819663FC43542A1F92" box="[558,631,912,936]" captionStart="Map 2" captionStartId="10.[197,241,1570,1591]" captionTargetBox="[195,1178,841,1563]" captionTargetId="figure-419@10.[195,1178,841,1563]" captionTargetPageId="10" captionText="Map 2: Distributions of Pronomaea picea HEERHEER (filled circles), P. corsicana sp. n. (open circles), and P. wunderleiwunderlei sp. n. (squares), based on examined records. The records of doubtful identity from Malta are omitted." figureDoi="http://doi.org/10.5281/zenodo.5364172" httpUri="https://zenodo.org/record/5364172/files/figure.png" pageId="27" pageNumber="394">Map 2</figureCitation>
). ...........................................
<taxonomicName id="4C6B4D1CFFF1FF8197FAFC4352191F99" ID-CoL="4MTB2" authority="HEER" authorityName="HEERHEER" authorityYear="1841" box="[951,1092,912,936]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="394" phylum="Arthropoda" rank="species" species="picea">
<emphasis id="B91FEA8DFFF1FF8197FAFC43525E1F92" bold="true" box="[951,1027,912,936]" italics="true" pageId="27" pageNumber="394">P. picea</emphasis>
HEER
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8BD4369FFFF1FF8194FAFC0F550B1FCE" blockId="27.[183,854,988,1013]" box="[183,854,988,1013]" pageId="27" pageNumber="394">
<emphasis id="B91FEA8DFFF1FF8194FAFC0F550B1FCE" bold="true" box="[183,854,988,1013]" pageId="27" pageNumber="394">
Catalogue of the
<taxonomicName id="4C6B4D1CFFF1FF81953EFC0E541A1FCE" box="[371,583,988,1012]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="394" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91FEA8DFFF1FF81953EFC0E57AE1FD5" bold="true" box="[371,499,989,1007]" italics="true" pageId="27" pageNumber="394">Pronomaea</emphasis>
species
</taxonomicName>
of the Palaearctic region
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>