320 lines
38 KiB
XML
320 lines
38 KiB
XML
<document id="49CEDB58CEC296DC8AC9CA52C67C4CEE" ID-DOI="10.11646/zootaxa.5396.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10441291" ID-ZooBank="6D5C03F9-7954-423B-836F-0DEF34AE37F6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703840530472" checkinUser="plazi" docAuthor="Huang, Di-Ying, Xu, Min-Min, Zhao, Qian, Gao, Jian & Nel, André" docDate="2023" docId="03C2695FFF99DB0B14D04843E377FE51" docLanguage="en" docName="zootaxa.5396.1.7.pdf" docOrigin="Zootaxa 5396 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5396.1.7/52600" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxfordgomphus trescellulae Huang & Xu & Zhao & Gao & Nel 2023, sp. nov." docType="treatment" docUuid="15C1AF17-A570-420B-8D4D-060DF148CE16" docUuidSource="ZooBank" docVersion="2" lastPageNumber="32" masterDocId="FFFB1127FF9DDB0D14474B00E268FFD8" masterDocTitle="New stem-anisopteran dragonflies in the Jurassic of China (Odonata: Epiproctophora)" masterLastPageNumber="34" masterPageNumber="26" pageNumber="30" updateTime="1703840930145" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="C675E51BA3A3C782627785D66A129F39">
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<mods:title id="95A800ECC9CADFC1474C1B131C6A0802">New stem-anisopteran dragonflies in the Jurassic of China (Odonata: Epiproctophora)</mods:title>
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</mods:titleInfo>
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<mods:name id="3B965E40FDE7B754BE7FA9D605CB9550" type="personal">
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<mods:roleTerm id="957C41BD51A16CE28BB9C018F65892DC">Author</mods:roleTerm>
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<mods:namePart id="A802F803378F76C333E4B7DBBEA06629">Huang, Di-Ying</mods:namePart>
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<mods:affiliation id="38FC4A4094155BCD96AD398F6CB2FD49">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing. Institute of Geology and Paleontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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</mods:name>
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<mods:name id="F066D54E32DCC9A3A9A2D28CF02788F2" type="personal">
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<mods:role id="826A6A50377496140F381A6B4A50B6AF">
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<mods:roleTerm id="1954A016E6E36FE0962258B6D4AB411E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FBF90E8067BEDF6D36AE6C0FB2E87B00">Xu, Min-Min</mods:namePart>
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<mods:affiliation id="CCD22EBD8A6DA5711DAA7271BA4A88A0">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing. Institute of Geology and Paleontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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</mods:name>
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<mods:name id="36698E917D818AB31073593CCE54147E" type="personal">
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<mods:role id="19ECE0162C3621807A27CB2BF1174D4A">
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<mods:roleTerm id="5CEE7290E785CE68BE2BE96E0B97E48D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="43A9CC56E46DAB590076DC7FC10E96A4">Zhao, Qian</mods:namePart>
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<mods:affiliation id="DD6470DF0C517D4A7059906F760B7DD3">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing. Institute of Geology and Paleontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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</mods:name>
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<mods:name id="68B65979968D01E546ACDFC412B7139E" type="personal">
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<mods:roleTerm id="BCDFF82024A1E2C1B4697B4BBB65AD08">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="77DEE8A24FB52CD02A3B226A53A88B0D">Gao, Jian</mods:namePart>
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<mods:affiliation id="4D4D0DDF34E63DE2062C80569E20F9EF">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing. Institute of Geology and Paleontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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</mods:name>
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<mods:name id="744B665B62861A7D669B9D7DFBBDFC6F" type="personal">
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<mods:roleTerm id="7DBDE1B732E5CB2211BB0C20F42D2F77">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="031421C334973496311CB35040318051">Nel, André</mods:namePart>
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<mods:affiliation id="C57942EBDF79EC3044916A53F48841A3">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 57 rue Cuvier, 75005 Paris, France</mods:affiliation>
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<mods:typeOfResource id="324690F38DC6B0DC87BF55DC671FC399">text</mods:typeOfResource>
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<mods:title id="F1CDF1C996430962E1F080692CF792A4">Zootaxa</mods:title>
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<mods:part id="0628211C2EF1434A4B6AA58DC4F06CF3">
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<mods:date id="083ED5107A6123F60C74C279C08C5D5B">2023</mods:date>
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<mods:number id="D295C2B738C43F55F5BC440B12C486D4">2023-12-29</mods:number>
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<mods:number id="F915DE112D5D60FBFC157016A8C6EF03">5396</mods:number>
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<mods:detail id="CA3A35A6C25954F50EB1D8C4BBD53297" type="issue">
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<mods:number id="8849283EA3A1EC3ABC007492383B2444">1</mods:number>
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<mods:start id="AA61EFB0A4C541C1F50DEA76042C6370">26</mods:start>
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<mods:end id="7D41E2478B23DE27101B1C760E894216">34</mods:end>
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<mods:location id="B9D859970ADD8C309838C2F369010246">
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<mods:url id="F5C9B62F968128BB4F932C341036F4AB">https://www.mapress.com/zt/article/download/zootaxa.5396.1.7/52600</mods:url>
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</mods:location>
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<mods:classification id="865B6A48D9DD26CA68CD2EDCA8A88120">journal article</mods:classification>
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<mods:identifier id="FC473A9C5C9278953394009BC22FDCEE" type="DOI">10.11646/zootaxa.5396.1.7</mods:identifier>
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<mods:identifier id="E4FA31981F804B37D8C8754547D7C109" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="46A3E4DC879B4FF95EAB1AD9B6D050CC" type="Zenodo-Dep">10441291</mods:identifier>
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<mods:identifier id="E4900026DB19F35809C5EE548D801B1C" type="ZooBank">6D5C03F9-7954-423B-836F-0DEF34AE37F6</mods:identifier>
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</mods:mods>
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<treatment id="03C2695FFF99DB0B14D04843E377FE51" ID-DOI="http://doi.org/10.5281/zenodo.10441303" ID-Zenodo-Dep="10441303" LSID="urn:lsid:zoobank.org:act:15C1AF17-A570-420B-8D4D-060DF148CE16" httpUri="http://treatment.plazi.org/id/03C2695FFF99DB0B14D04843E377FE51" lastPageId="6" lastPageNumber="32" pageId="4" pageNumber="30">
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<subSubSection id="C3718BC2FF99DB0914D04843E1A1FC5A" pageId="4" pageNumber="30" type="nomenclature">
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||
<paragraph id="8BD4D849FF99DB0914D04843E169FC85" blockId="4.[151,969,835,933]" box="[151,769,835,862]" pageId="4" pageNumber="30">
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<heading id="D09C6F25FF99DB0914D04843E169FC85" bold="true" box="[151,769,835,862]" fontSize="11" level="1" pageId="4" pageNumber="30" reason="1">
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<emphasis id="B91F045BFF99DB0914D04843E169FC85" bold="true" box="[151,769,835,862]" pageId="4" pageNumber="30">
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<taxonomicName id="4C6BA3CAFF99DB0914D04843E0F3FC85" authority="Huang & Nel" authorityName="Huang & Xu & Zhao & Gao & Nel" authorityYear="2023" box="[151,667,835,862]" class="Insecta" family="Juragomphidae" genus="Oxfordgomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="trescellulae" status="sp. nov.">
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<emphasis id="B91F045BFF99DB0914D04843E387FC86" bold="true" box="[151,495,835,862]" italics="true" pageId="4" pageNumber="30">Oxfordgomphus trescellulae</emphasis>
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Huang & Nel
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</taxonomicName>
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<taxonomicNameLabel id="A22CB920FF99DB0916E44843E169FC85" box="[675,769,835,861]" pageId="4" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BD4D849FF99DB0914D04867E1A1FC5A" blockId="4.[151,969,835,933]" box="[151,969,871,898]" pageId="4" pageNumber="30">
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||
<uri id="FFFAD44BFF99DB0914D04867E1A1FC5A" box="[151,969,871,898]" pageId="4" pageNumber="30">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="FFCDE29CFF99DB0915EA4867E1A1FC5A" box="[429,969,871,898]" pageId="4" pageNumber="30">15C1AF17-A570-420B-8D4D-060DF148CE16</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3718BC2FF99DB0914D0488BE374FC7D" box="[151,284,907,933]" pageId="4" pageNumber="30" type="description">
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||
<paragraph id="8BD4D849FF99DB0914D0488BE374FC7D" blockId="4.[151,969,835,933]" box="[151,284,907,933]" pageId="4" pageNumber="30">
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<figureCitation id="1350C4CCFF99DB0914D0488BE296FC7D" box="[151,254,907,933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1816,1840]" captionTargetBox="[164,1423,237,1792]" captionTargetId="figure-49@5.[164,1423,237,1792]" captionTargetPageId="5" captionText="FIGURE 3. Oxfordgomphus trescellulae Huang & Nel, gen. et sp. nov., holotype NIGP202929, photographs. a, part; b, counterpart. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10441297" httpUri="https://zenodo.org/record/10441297/files/figure.png" pageId="4" pageNumber="30">Figures 3</figureCitation>
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–4
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3718BC2FF99DB0914D048D3E616FC35" box="[151,1150,979,1005]" pageId="4" pageNumber="30" type="etymology">
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<paragraph id="8BD4D849FF99DB0914D048D3E616FC35" blockId="4.[151,1437,979,2014]" box="[151,1150,979,1005]" pageId="4" pageNumber="30">
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<emphasis id="B91F045BFF99DB0914D048D3E376FC35" bold="true" box="[151,286,979,1005]" pageId="4" pageNumber="30">Etymology.</emphasis>
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||
Named after the subdivision of the discoidal triangle into three smaller cells.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3718BC2FF99DB09148048F7E110FBA5" pageId="4" pageNumber="30" type="materials_examined">
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||
<paragraph id="8BD4D849FF99DB09148048F7E13BFBED" blockId="4.[151,1437,979,2014]" pageId="4" pageNumber="30">
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||
<emphasis id="B91F045BFF99DB09148048F7E35EFBC9" bold="true" box="[199,310,1015,1041]" pageId="4" pageNumber="30">Material.</emphasis>
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<materialsCitation id="3B03D214FF99DB09157B48F7E13BFBED" collectionCode="CAS" country="China" county="Nanjing" location="Nanjing Institute of Geology" municipality="Palaeontology" pageId="4" pageNumber="30" specimenCode="NIGP202929" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54D066EBFF99DB09157B48F7E3CCFBC9" box="[316,420,1015,1041]" pageId="4" pageNumber="30" type="holotype">Holotype</typeStatus>
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<specimenCode id="DBCD7032FF99DB0915ED48F7E057FBC9" box="[426,575,1015,1041]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="4" pageNumber="30">NIGP202929</specimenCode>
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(part and counterpart of the basal fourth of a forewing), housed in the
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<location id="8EB48E92FF99DB09110548F7E310FBED" LSID="urn:lsid:plazi:treatment:03C2695FFF99DB0B14D04843E377FE51:8EB48E92FF99DB09110548F7E310FBED" country="China" county="Nanjing" municipality="Palaeontology" name="Nanjing Institute of Geology" pageId="4" pageNumber="30">Nanjing Institute of Geology</location>
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and
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<collectingMunicipality id="6BB04233FF99DB0915E84F1BE038FBED" box="[431,592,1051,1077]" pageId="4" pageNumber="30">Palaeontology</collectingMunicipality>
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,
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<collectionCode id="ED7A408CFF99DB09161C4F1BE0FCFBED" box="[603,660,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="4" pageNumber="30" type="Museum">CAS</collectionCode>
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,
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<collectingCounty id="62B5A0C5FF99DB0916E74F1BE095FBED" box="[672,765,1051,1077]" pageId="4" pageNumber="30">Nanjing</collectingCounty>
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,
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<collectingCountry id="F37C98D9FF99DB09174F4F1BE127FBED" box="[776,847,1051,1077]" name="China" pageId="4" pageNumber="30">China</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD4D849FF99DB0914804F3FE110FBA5" blockId="4.[151,1437,979,2014]" pageId="4" pageNumber="30">
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<emphasis id="B91F045BFF99DB0914804F3FE3FBFB81" bold="true" box="[199,403,1087,1113]" pageId="4" pageNumber="30">Age and outcrop.</emphasis>
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Earliest Late Jurassic (Oxfordian); uppermost part of Yangshuzhuang Formation near Anyao Village, Chengliu Township, Jiyuan City,
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<collectingRegion id="49AF16ABFF99DB09162B4F64E14AFBA5" box="[620,802,1123,1150]" country="China" name="Henan" pageId="4" pageNumber="30">Henan Province</collectingRegion>
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,
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<collectingCountry id="F37C98D9FF99DB09176A4F63E11CFBA5" box="[813,884,1123,1149]" name="China" pageId="4" pageNumber="30">China</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3718BC2FF99DB0914804F87E0C9FB7A" box="[199,673,1159,1186]" pageId="4" pageNumber="30" type="diagnosis">
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<paragraph id="8BD4D849FF99DB0914804F87E0C9FB7A" blockId="4.[151,1437,979,2014]" box="[199,673,1159,1186]" pageId="4" pageNumber="30">
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<emphasis id="B91F045BFF99DB0914804F87E32AFB79" bold="true" box="[199,322,1159,1185]" pageId="4" pageNumber="30">Diagnosis.</emphasis>
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As for the genus by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3718BC2FF99DB0914804FABE15FF98A" pageId="4" pageNumber="30" type="description">
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<paragraph id="8BD4D849FF99DB0914804FABE15FF98A" blockId="4.[151,1437,979,2014]" pageId="4" pageNumber="30">
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<emphasis id="B91F045BFF99DB0914804FABE331FB1D" bold="true" box="[199,345,1195,1221]" pageId="4" pageNumber="30">Description.</emphasis>
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Basal fourth of a forewing, apparently hyaline, fragment
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<quantity id="4C9375ACFF99DB0910414FABE606FB1E" box="[1030,1134,1195,1222]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="4" pageNumber="30" unit="mm" value="11.6">11.6 mm</quantity>
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long, wing
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<quantity id="4C9375ACFF99DB0911474FABE733FB1E" box="[1280,1371,1195,1222]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="4" pageNumber="30" unit="mm" value="7.4">7.4 mm</quantity>
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wide at level of discoidal triangle; distance from base to arculus
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<quantity id="4C9375ACFF99DB0917134FCFE1C7FB32" box="[852,943,1231,1258]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="30" unit="mm" value="5.6">5.6 mm</quantity>
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, from base to Ax1
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<quantity id="4C9375ACFF99DB0910D04FCFE69CFB32" box="[1175,1268,1231,1258]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="30" unit="mm" value="4.3">4.3 mm</quantity>
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; Ax2 opposite distal side of discoidal triangle,
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<quantity id="4C9375ACFF99DB0916414FF3E035FAD6" box="[518,605,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="30" unit="mm" value="4.6">4.6 mm</quantity>
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distad Ax1; Ax2 obliquely directed; two secondary antenodal crossveins between Ax1 and Ax2, no antenodal basad Ax1, three distad Ax2; arculus angular; RP and MA well separated in arculus; two crossveins between RA and RP above discoidal triangle; four preserved crossveins between RP and MA; hypertriangle
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<quantity id="4C9375ACFF99DB0915364E5FE3A0FAA2" box="[369,456,1375,1402]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="30" unit="mm" value="3.4">3.4 mm</quantity>
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long,
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<quantity id="4C9375ACFF99DB0916564E5FE000FAA2" box="[529,616,1375,1402]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="30" unit="mm" value="0.6">0.6 mm</quantity>
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wide, short, free, with basal part of MA straight; short distance between arculus and basal side of discoidal triangle,
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<quantity id="4C9375ACFF99DB0916C04E83E0B6FA46" box="[647,734,1411,1438]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="30" unit="mm" value="0.6">0.6 mm</quantity>
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apart; anterior side of discoidal triangle ending at distal angle of hypertriangle; discoidal triangle equilateral, divided into three smaller cells; sides of discoidal triangle
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<quantity id="4C9375ACFF99DB0911024EA7E7F5FA1A" box="[1349,1437,1447,1474]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="30" unit="mm" value="2.9">2.9 mm</quantity>
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long; postdiscoidal area with two rows of cells just distad discoidal triangle; median and submedian spaces free; CuP distinctly curved, situated at the point where AA is posteriorly curved; subdiscoidal space basally closed by CuP rather than by a PsA, acutely triangular, and crossed by two-three irregularly curved crossveins; a rather broad elongate cell situated below subdiscoidal space in anal area.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3718BC2FF99DB0B14804D5BE377FE51" lastPageId="6" lastPageNumber="32" pageId="4" pageNumber="30" type="discussion">
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<paragraph id="8BD4D849FF99DB0814804D5BE13AFF0D" blockId="4.[151,1437,979,2014]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="31" pageId="4" pageNumber="30">
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<emphasis id="B91F045BFF99DB0914804D5BE353F9AD" bold="true" box="[199,315,1627,1653]" pageId="4" pageNumber="30">Remarks.</emphasis>
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Although fragmentary, this forewing base shows characters of the two families
|
||
<taxonomicName id="4C6BA3CAFF99DB0910904D5BE351F942" authority="Tillyard, 1935" authorityName="Tillyard" authorityYear="1935" class="Insecta" family="Liassogomphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="family">
|
||
Liassogomphidae
|
||
<bibRefCitation id="EFFAA5B8FF99DB0914D04D7FE351F942" author="Tillyard, R. J." box="[151,313,1663,1690]" pageId="4" pageNumber="30" pagination="374 - 384" refId="ref5944" refString="Tillyard, R. J. (1935) Upper Permian insects of New South Wales. 4. The order Odonata. Proceedings of the Linnean Society of New South Wales, 60, 374 - 384." type="journal article" year="1935">Tillyard, 1935</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6BA3CAFF99DB0915324D7FE04BF941" box="[373,547,1663,1689]" class="Insecta" family="Juragomphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="family">Juragomphidae</taxonomicName>
|
||
, viz. a CuP distinctly curved and distally displaced at the point where AA is posteriorly curved, subdiscoidal space basally closed by CuP rather than by a PsA, acutely triangular, and crossed by two-three irregularly curved crossveins; a rather broad elongate cell situated below the subdiscoidal space; a short hypertriangle with base of discoidal triangle very close to arculus; numerous crossveins in area between RP and MA basad fork of MA into MAa and MAb (
|
||
<bibRefCitation id="EFFAA5B8FF99DB0916804C0FE110F8F1" author="Nel, A. & Martinez-Delclos, X. & Paicheler, J. - C. & Henrotay, M." box="[711,888,1807,1833]" pageId="4" pageNumber="30" pagination="1 - 311" refId="ref5602" refString="Nel, A., Martinez-Delclos, X., Paicheler, J. - C. & Henrotay, M. (1993) Les ' Anisozygoptera' fossiles. Phylogenie et classification (Odonata). Martinia Numero Hors Serie, 3, 1 - 311." type="journal article" year="1993">
|
||
Nel
|
||
<emphasis id="B91F045BFF99DB0916BD4C10E15CF8F1" box="[762,820,1807,1833]" italics="true" pageId="4" pageNumber="30">et al.</emphasis>
|
||
1993
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFAA5B8FF99DB0917C14C10E60FF8F1" author="Etter, W. & Kuhn, O." box="[902,1127,1807,1834]" pageId="4" pageNumber="30" pagination="967 - 977" refId="ref4246" refString="Etter, W. & Kuhn, O. (2000) An articulated dragonfly (Insecta, Odonata) from the Upper Liassic Posidonia shale of Northern Switzerland. Palaeontology, 43 (5), 967 - 977. https: // doi. org / 10.1111 / 1475 - 4983.00157" type="journal article" year="2000">Etter & Kuhn 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFAA5B8FF99DB0910334C0FE74CF8F1" author="Nel, A. & Bechly, G. & Martinez-Delclos, X. & Fleck, G." box="[1140,1316,1807,1833]" pageId="4" pageNumber="30" pagination="1 - 9" refId="ref5416" refString="Nel, A., Bechly, G., Martinez-Delclos, X. & Fleck, G. (2001) A new family of Anisoptera from the Upper Jurassic of Karatau in Kazakhstan (Insecta: Odonata: Juragomphidae n. fam.). Stuttgarter Beitrage zur Naturkunde, (B), 314, 1 - 9." type="journal article" year="2001">
|
||
Nel
|
||
<emphasis id="B91F045BFF99DB0910E24C10E688F8F1" box="[1189,1248,1807,1833]" italics="true" pageId="4" pageNumber="30">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
). The new fossil shares with
|
||
<taxonomicName id="4C6BA3CAFF99DB0915224C33E0D0F895" authority="Nel et al., 2001" authorityName="Nel" authorityYear="2001" box="[357,696,1843,1870]" class="Insecta" family="Juragomphidae" genus="Juragomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F045BFF99DB0915224C33E395F895" box="[357,509,1843,1869]" italics="true" pageId="4" pageNumber="30">Juragomphus</emphasis>
|
||
<bibRefCitation id="EFFAA5B8FF99DB0916414C33E0D0F895" author="Nel, A. & Bechly, G. & Martinez-Delclos, X. & Fleck, G." box="[518,696,1843,1870]" pageId="4" pageNumber="30" pagination="1 - 9" refId="ref5416" refString="Nel, A., Bechly, G., Martinez-Delclos, X. & Fleck, G. (2001) A new family of Anisoptera from the Upper Jurassic of Karatau in Kazakhstan (Insecta: Odonata: Juragomphidae n. fam.). Stuttgarter Beitrage zur Naturkunde, (B), 314, 1 - 9." type="journal article" year="2001">
|
||
Nel
|
||
<emphasis id="B91F045BFF99DB0916704C34E018F895" box="[567,624,1843,1869]" italics="true" pageId="4" pageNumber="30">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
an equilateral discoidal triangle as a putative synapomorphy, but this character is convergently present in some other Mesozoic
|
||
<taxonomicName id="4C6BA3CAFF99DB0917224C57E18BF8A9" authorityName="Selys" authorityYear="1854" box="[869,995,1879,1905]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="30" phylum="Tracheophyta" rank="subOrder" subOrder="Anisoptera">Anisoptera</taxonomicName>
|
||
, viz. in the
|
||
<taxonomicName id="4C6BA3CAFF99DB09102A4C57E292F84D" authority="Bechly et al., 2001" authorityName="Bechly" authorityYear="2001" class="Insecta" family="Liupanshaniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="family">
|
||
Liupanshaniidae
|
||
<bibRefCitation id="EFFAA5B8FF99DB0911684C57E292F84D" author="Bechly, G. & Nel, A. & Martinez-Delclos, X. & Jarzembowski, E. A. & Coram, R. & Martill, D. & Fleck, G. & Escuillie, F. & Wisshak, M. M. & Maisch, M." pageId="4" pageNumber="30" pagination="1 - 219" refId="ref4008" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 219." type="journal article" year="2001">
|
||
Bechly
|
||
<emphasis id="B91F045BFF99DB0911CF4C58E2C5F84D" italics="true" pageId="4" pageNumber="30">et al</emphasis>
|
||
., 2001
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(Aeshnoptera
|
||
<bibRefCitation id="EFFAA5B8FF99DB0915E74C7BE05CF84D" author="Bechly, G." box="[416,564,1915,1941]" pageId="4" pageNumber="30" pagination="1 - 402" refId="ref3814" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura Special Volume, 2, 1 - 402." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
) and in the
|
||
<taxonomicName id="4C6BA3CAFF99DB0916FE4C7BE154F84D" box="[697,828,1915,1941]" class="Insecta" family="Aktassiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="family">Aktassiidae</taxonomicName>
|
||
Aktassiinae
|
||
<bibRefCitation id="EFFAA5B8FF99DB09178C4C7BE610F84D" author="Pritykina, L. N." box="[971,1144,1915,1941]" pageId="4" pageNumber="30" pagination="26 - 55" refId="ref5650" refString="Pritykina, L. N. (1968) Strekozy Karatau (Odonata). [Dragonflies from Karatau (Odonata)]. In: Panfilov, D. V. (coord.), Yurskie Nasekomye Karatau. [Jurassic insects from Karatau]. Academy of Sciences of the S. S. S. R., Nauka, Moscow, pp. 26 - 55. [in Russian]" type="book chapter" year="1968">Pritykina, 1968</bibRefCitation>
|
||
(Petalurida
|
||
<bibRefCitation id="EFFAA5B8FF99DB0911464C7BE7F0F84D" author="Bechly, G." box="[1281,1432,1915,1941]" pageId="4" pageNumber="30" pagination="1 - 402" refId="ref3814" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura Special Volume, 2, 1 - 402." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C6BA3CAFF99DB0914D04C9FE3C7F861" authority="Pritykina, 1968" authorityName="Pritykina" authorityYear="1968" box="[151,431,1951,1977]" class="Insecta" family="Aktassiidae" genus="Aktassia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F045BFF99DB0914D04C9FE29EF861" box="[151,246,1951,1977]" italics="true" pageId="4" pageNumber="30">Aktassia</emphasis>
|
||
<bibRefCitation id="EFFAA5B8FF99DB0915474C9FE3C7F861" author="Pritykina, L. N." box="[256,431,1951,1977]" pageId="4" pageNumber="30" pagination="26 - 55" refId="ref5650" refString="Pritykina, L. N. (1968) Strekozy Karatau (Odonata). [Dragonflies from Karatau (Odonata)]. In: Panfilov, D. V. (coord.), Yurskie Nasekomye Karatau. [Jurassic insects from Karatau]. Academy of Sciences of the S. S. S. R., Nauka, Moscow, pp. 26 - 55. [in Russian]" type="book chapter" year="1968">Pritykina, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6BA3CAFF99DB0915AC4C9FE1E8F861" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[491,896,1951,1977]" class="Insecta" family="Aktassiidae" genus="Aeschnogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F045BFF99DB0915AC4C9FE0C7F861" box="[491,687,1951,1977]" italics="true" pageId="4" pageNumber="30">Aeschnogomphus</emphasis>
|
||
<bibRefCitation id="EFFAA5B8FF99DB0916FE4C9FE1E8F861" author="Handlirsch, A." box="[697,896,1951,1977]" pageId="4" pageNumber="30" refId="ref4561" refString="Handlirsch, A. (1906 - 1908) Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch fur Palaontologen und Zoologen. Engelman, V. W. publ., Leipzig, 1430 pp. https: // doi. org / 10.5962 / bhl. title. 34145" type="book" year="1906">Handlirsch, 1906</bibRefCitation>
|
||
</taxonomicName>
|
||
). The new fossil and
|
||
<taxonomicName id="4C6BA3CAFF99DB09103B4C9FE77CF861" box="[1148,1300,1951,1977]" class="Insecta" family="Juragomphidae" genus="Juragomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F045BFF99DB09103B4C9FE77CF861" box="[1148,1300,1951,1977]" italics="true" pageId="4" pageNumber="30">Juragomphus</emphasis>
|
||
</taxonomicName>
|
||
differ from these taxa in the presence of a weak PsA, and of a curved CuP, vs. strong PsA and a straight CuP in the latter taxa (
|
||
<bibRefCitation id="EFFAA5B8FF98DB0814924B97E3EBFF69" author="Nel, A. & Bechly, G. & Jarzembowski, E. A. & Martinez-Delclos, X." box="[213,387,151,177]" pageId="5" pageNumber="31" pagination="1 - 68" refId="ref5361" refString="Nel, A., Bechly, G., Jarzembowski, E. A. & Martinez-Delclos, X. (1998) A revision of the fossil petalurid dragonflies (Insecta: Odonata: Anisoptera: Petalurida). Paleontologia Lombarda, (New Series), 10, 1 - 68." type="journal article" year="1998">
|
||
Nel
|
||
<emphasis id="B91F045BFF98DB0815404B97E328FF69" box="[263,320,151,177]" italics="true" pageId="5" pageNumber="31">et al.</emphasis>
|
||
1998
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFAA5B8FF98DB0815D74B97E00CFF69" author="Bechly, G. & Nel, A. & Martinez-Delclos, X. & Jarzembowski, E. A. & Coram, R. & Martill, D. & Fleck, G. & Escuillie, F. & Wisshak, M. M. & Maisch, M." box="[400,612,151,177]" pageId="5" pageNumber="31" pagination="1 - 219" refId="ref4008" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 219." type="journal article" year="2001">
|
||
Bechly
|
||
<emphasis id="B91F045BFF98DB0815A04B97E048FF69" box="[487,544,151,177]" italics="true" pageId="5" pageNumber="31">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
). The
|
||
<taxonomicName id="4C6BA3CAFF98DB0816EC4B97E119FF69" authorityName="Tillyard" authorityYear="1935" box="[683,881,151,177]" class="Insecta" family="Liassogomphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="31" phylum="Arthropoda" rank="family">Liassogomphidae</taxonomicName>
|
||
have a more transverse discoidal triangle than in
|
||
<taxonomicName id="4C6BA3CAFF98DB0814D04BBBE347FF0D" box="[151,303,187,213]" class="Insecta" family="Juragomphidae" genus="Juragomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F045BFF98DB0814D04BBBE347FF0D" box="[151,303,187,213]" italics="true" pageId="5" pageNumber="31">Juragomphus</emphasis>
|
||
</taxonomicName>
|
||
and the new fossil (
|
||
<bibRefCitation id="EFFAA5B8FF98DB0816574BBBE0E7FF0D" author="Bode, A." box="[528,655,186,213]" pageId="5" pageNumber="31" pagination="1 - 375" refId="ref4101" refString="Bode, A. (1953) Die Insektenfauna des Ostniedersachsischen oberen Lias. Palaeontographica, (A), 103 (1 - 4), 1 - 375." type="journal article" year="1953">Bode 1953</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFAA5B8FF98DB0816DC4BBBE12DFF0D" author="Nel, A. & Martinez-Delclos, X. & Paicheler, J. - C. & Henrotay, M." box="[667,837,187,213]" pageId="5" pageNumber="31" pagination="1 - 311" refId="ref5602" refString="Nel, A., Martinez-Delclos, X., Paicheler, J. - C. & Henrotay, M. (1993) Les ' Anisozygoptera' fossiles. Phylogenie et classification (Odonata). Martinia Numero Hors Serie, 3, 1 - 311." type="journal article" year="1993">
|
||
Nel
|
||
<emphasis id="B91F045BFF98DB08168D4BBBE16BFF0D" box="[714,771,187,213]" italics="true" pageId="5" pageNumber="31">et al.</emphasis>
|
||
1993
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF1488C1FF98DB0814D04C18E3B3F88B" ID-DOI="http://doi.org/10.5281/zenodo.10441297" ID-Zenodo-Dep="10441297" httpUri="https://zenodo.org/record/10441297/files/figure.png" pageId="5" pageNumber="31" startId="5.[151,250,1816,1840]" targetBox="[164,1423,237,1792]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BD4D849FF98DB0814D04C18E3B3F88B" blockId="5.[151,1436,1816,1876]" pageId="5" pageNumber="31">
|
||
<emphasis id="B91F045BFF98DB0814D04C18E37FF8E8" bold="true" box="[151,279,1816,1840]" pageId="5" pageNumber="31">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C6BA3CAFF98DB0815664C18E089F8E8" authority="Huang & Nel" authorityName="Huang & Xu & Zhao & Gao & Nel" authorityYear="2023" box="[289,737,1816,1840]" class="Insecta" family="Juragomphidae" genus="Oxfordgomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="31" phylum="Arthropoda" rank="species" species="trescellulae" status="gen. et sp. nov.">
|
||
<emphasis id="B91F045BFF98DB0815664C18E02BF8E8" box="[289,579,1816,1840]" italics="true" pageId="5" pageNumber="31">Oxfordgomphus trescellulae</emphasis>
|
||
Huang & Nel
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B91F045BFF98DB0816A84C18E1F0F8E8" bold="true" box="[751,920,1816,1840]" pageId="5" pageNumber="31">
|
||
<taxonomicNameLabel id="A22CB920FF98DB0816A84C18E1F0F8E8" box="[751,920,1816,1840]" pageId="5" pageNumber="31" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype NIGP202929, photographs.
|
||
<emphasis id="B91F045BFF98DB0811754C18E757F8E8" bold="true" box="[1330,1343,1816,1840]" pageId="5" pageNumber="31">a</emphasis>
|
||
, part;
|
||
<emphasis id="B91F045BFF98DB0811CF4C18E7FEF8E8" bold="true" box="[1416,1430,1816,1840]" pageId="5" pageNumber="31">b</emphasis>
|
||
, counterpart. Scale bars = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD4D849FF98DB0814804CA6E658F83C" blockId="5.[151,1436,1958,2021]" pageId="5" pageNumber="31">
|
||
The new fossil and the
|
||
<taxonomicName id="4C6BA3CAFF98DB0815964CA6E0FFF818" authorityName="Tillyard" authorityYear="1935" box="[465,663,1958,1984]" class="Insecta" family="Liassogomphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="31" phylum="Arthropoda" rank="family">Liassogomphidae</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="4C6BA3CAFF98DB0817644CA6E1D3F818" box="[803,955,1958,1984]" class="Insecta" family="Juragomphidae" genus="Juragomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F045BFF98DB0817644CA6E1D3F818" box="[803,955,1958,1984]" italics="true" pageId="5" pageNumber="31">Juragomphus</emphasis>
|
||
</taxonomicName>
|
||
in the less numerous cells in the discoidal triangle and the postdiscoidal area, and in the less numerous antenodal crossveins.
|
||
</paragraph>
|
||
<paragraph id="8BD4D849FF9BDB0B14804B97E1E6FF21" blockId="6.[151,1437,151,393]" pageId="6" pageNumber="32">
|
||
Thus, the affinities of the new fossil with the
|
||
<taxonomicName id="4C6BA3CAFF9BDB0B16A84B97E1DDFF69" authorityName="Tillyard" authorityYear="1935" box="[751,949,151,177]" class="Insecta" family="Liassogomphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="32" phylum="Arthropoda" rank="family">Liassogomphidae</taxonomicName>
|
||
vs. the
|
||
<taxonomicName id="4C6BA3CAFF9BDB0B105D4B97E6ADFF69" box="[1050,1221,151,177]" class="Insecta" family="Juragomphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="32" phylum="Arthropoda" rank="family">Juragomphidae</taxonomicName>
|
||
remain somewhat uncertain, although an attribution to the latter family could be more likely for a shared putative apomorphy. Thus we tentatively attribute it to this family, in a new genus and species.
|
||
</paragraph>
|
||
<paragraph id="8BD4D849FF9BDB0B14804A03E377FE51" blockId="6.[151,1437,151,393]" pageId="6" pageNumber="32">
|
||
<emphasis id="B91F045BFF9BDB0B14804A03E36EFEC5" bold="true" box="[199,262,259,285]" pageId="6" pageNumber="32">Note.</emphasis>
|
||
The
|
||
<taxonomicName id="4C6BA3CAFF9BDB0B15044A03E0DAFEC5" authority="Fleck et al., 2003" authorityName="Fleck" authorityYear="2003" box="[323,690,259,285]" class="Insecta" family="Henrotayiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="32" phylum="Arthropoda" rank="family">
|
||
Henrotayiidae
|
||
<bibRefCitation id="EFFAA5B8FF9BDB0B15AD4A03E0DAFEC5" author="Fleck, G. & Bechly, G. & Martinez-Delclos, X. & Jarzembowski, E. A. & Coram, R. & Nel, A." box="[490,690,259,285]" pageId="6" pageNumber="32" pagination="55 - 93" refId="ref4364" refString="Fleck, G., Bechly, G., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R. & Nel, A. (2003) Phylogeny and classification of the Stenophlebioptera (Odonata, Epiproctophora). Annales de la Societe Entomologique de France, (New Series), 39 (1), 55 - 93. https: // doi. org / 10.1080 / 00379271.2003.10697363" type="journal article" year="2003">
|
||
Fleck
|
||
<emphasis id="B91F045BFF9BDB0B16764A03E002FEC5" box="[561,618,259,285]" italics="true" pageId="6" pageNumber="32">et al.</emphasis>
|
||
, 2003
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C6BA3CAFF9BDB0B16844A03E67AFEC5" authority="Fleck et al., 2003" authorityName="Fleck" authorityYear="2003" box="[707,1042,259,285]" class="Insecta" family="Henrotayiidae" genus="Henrotayia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F045BFF9BDB0B16844A03E12AFEC4" box="[707,834,259,284]" italics="true" pageId="6" pageNumber="32">Henrotayia</emphasis>
|
||
<bibRefCitation id="EFFAA5B8FF9BDB0B170D4A03E67AFEC5" author="Fleck, G. & Bechly, G. & Martinez-Delclos, X. & Jarzembowski, E. A. & Coram, R. & Nel, A." box="[842,1042,259,285]" pageId="6" pageNumber="32" pagination="55 - 93" refId="ref4364" refString="Fleck, G., Bechly, G., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R. & Nel, A. (2003) Phylogeny and classification of the Stenophlebioptera (Odonata, Epiproctophora). Annales de la Societe Entomologique de France, (New Series), 39 (1), 55 - 93. https: // doi. org / 10.1080 / 00379271.2003.10697363" type="journal article" year="2003">
|
||
Fleck
|
||
<emphasis id="B91F045BFF9BDB0B17D64A03E1A3FEC5" box="[913,971,259,285]" italics="true" pageId="6" pageNumber="32">et al.</emphasis>
|
||
, 2003
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
known by a hind wing) is another Liassic anisopteran family sharing a subdiscoidal space similar to that of the new fossil but it strongly differs from it in the absence of antenodal crossveins of first row between C and ScP and a quite smaller discoidal triangle (
|
||
<bibRefCitation id="EFFAA5B8FF9BDB0B111B4A4BE37BFE51" author="Fleck, G. & Bechly, G. & Martinez-Delclos, X. & Jarzembowski, E. A. & Coram, R. & Nel, A." pageId="6" pageNumber="32" pagination="55 - 93" refId="ref4364" refString="Fleck, G., Bechly, G., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R. & Nel, A. (2003) Phylogeny and classification of the Stenophlebioptera (Odonata, Epiproctophora). Annales de la Societe Entomologique de France, (New Series), 39 (1), 55 - 93. https: // doi. org / 10.1080 / 00379271.2003.10697363" type="journal article" year="2003">
|
||
Fleck
|
||
<emphasis id="B91F045BFF9BDB0B14D04A6FE2A7FE51" box="[151,207,367,393]" italics="true" pageId="6" pageNumber="32">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |