155 lines
18 KiB
XML
155 lines
18 KiB
XML
<document id="C07F2665BF45612AE7E5A34A8440C33C" ENCODING="UTF8" ID-CLB-Dataset="52775" ID-GBIF-Dataset="49f5a507-5a72-4c59-a638-203bb61a4a9a" ModsDocID="22782" checkinTime="1249509740462" checkinUser="christiana" docAuthor="Shattuck, S. O." docDate="2009" docId="57301B7B708B4B13F98661CD45DC99B0" docLanguage="en" docName="22782.xml" docOrigin="Zootaxa 2146" docSource="http://hol.osu.edu/reference-full.html?id=22782" docTitle="Myrmecina difficulta Shattuck, 2009, sp. n." docType="treatment" docUuid="48886DE1-0866-4FC9-98F1-83987EEA86CA" docUuidSource="ZooBank" docVersion="16" lastPageNumber="10" masterDocId="F8262E6E0A515024BA501A49CF21E327" masterDocTitle="A revision the Australian species of the ant genus Myrmecina (Hymenoptera: Formicidae)." masterLastPageNumber="21" masterPageNumber="1" pageNumber="8" updateTime="1732479109242" updateUser="ExternalLinkService">
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<mods:mods id="163CF884C55E3A3283D7C0C8C5857902" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="19565F4A8FBC08CC3D6ECFA9ABB2D023">
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<mods:title id="A5B2DE65FBB0E93EA92B55D1B8064FA1">A revision the Australian species of the ant genus Myrmecina (Hymenoptera: Formicidae).</mods:title>
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</mods:titleInfo>
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<mods:name id="42D7CFE48CBD619E74940DCC3DB714F9" type="personal">
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<mods:role id="810C3D986E096D07B0EB348B27562D81">
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<mods:roleTerm id="4580CC43179DB4BE54E1A9BAA55633CA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="925A59FA8D994FB33D22F0BEBBDD570F">Shattuck, S. O.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5FBAF46711B436163FFE17906C7FDC18">text</mods:typeOfResource>
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<mods:titleInfo id="79CE7317CFAFCBABEA200FD2FEF99FB8">
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<mods:title id="EF0883FF1B6203BF50B5D88B0F3C846C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B718CA9AB5B996103FF6A4F54070060E">
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<mods:date id="8299238C32720D2D4DE52229F6B64CB1">2009</mods:date>
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<mods:detail id="FD7F946F7D91A6DEE317071906AB1A95" type="volume">
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<mods:number id="31B28CC224D2FDD71E57C9FE5D5225E5">2146</mods:number>
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<mods:start id="8C0611AE41592656113D12657F443F12">1</mods:start>
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<mods:end id="FAA6E416D73BC8BAB072C8F8153CEF2A">21</mods:end>
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<mods:location id="2D7F4C15E6FBB4ECDEE2BE2637888314">
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<mods:url id="38E2BBB81D90E5A335A569A59D6CA020">http://hol.osu.edu/reference-full.html?id=22782</mods:url>
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</mods:location>
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<mods:classification id="2C7A36E1815253AAEAC8534A5A772329">journal article</mods:classification>
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<mods:identifier id="69FF9D65701E6032E4947419EB7AAD79" type="HNS-Pub">22782</mods:identifier>
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<mods:identifier id="E6DDB8FE0B99D7268CCAF29E18FC275A" type="ZooBank">C666693E-9FDE-4897-A20D-CBCE9B4F6D78</mods:identifier>
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</mods:mods>
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<treatment id="57301B7B708B4B13F98661CD45DC99B0" ID-DOI="http://doi.org/10.5281/zenodo.6223601" ID-GBIF-Taxon="100122469" ID-Zenodo-Dep="6223601" LSID="urn:lsid:zoobank.org:act:48886DE1-0866-4FC9-98F1-83987EEA86CA" httpUri="http://treatment.plazi.org/id/57301B7B708B4B13F98661CD45DC99B0" lastPageNumber="10" pageNumber="8" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="91A41DF0DE12DD40B05A9B196B3A779B" type="nomenclature">
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<paragraph id="DA2F4BB1C65A938F26156C54D21ECD2A" pageNumber="8">
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<taxonomicName id="2396809485F20A57C4AB2167A7546E35" ID-CoL="CMC9Z" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:251840" LSID-ZBK="urn:lsid:zoobank.org:act:48886DE1-0866-4FC9-98F1-83987EEA86CA" class="Insecta" family="Formicidae" genus="Myrmecina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmecina difficulta" lsidName-ZBK="Myrmecina difficulta Shattuck 2009:8" order="Hymenoptera" pageNumber="8" phylum="Arthropoda" rank="species" species="difficulta">Myrmecina difficulta</taxonomicName>
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<taxonomicNameLabel id="94BB043A589374DC7B8238F1AD369063" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="A7CE5C7165BF136D0353E460CD97D094" type="description">
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<paragraph id="BE16B846773899641518033E3C04B2CA" pageNumber="8">(Figs 6, 12-16, 49)</paragraph>
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</subSubSection>
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<subSubSection id="754E88CF7646682A8114CEB6E554217C" type="materials_examined">
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<paragraph id="4B941CA60CD4C42352EBAA4A2990B163" pageNumber="8">
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<materialsCitation id="17770AF847AE38C4BCBD1A4B8FBFEDE1" ID-GBIF-Occurrence="923892984" collectingDate="1971-07-07" collectionCode="ANIC" collectorName="R. W. Taylor & J. Feehan" country="Australia" location="Summit, Mt. Bellenden-Ker, 1700 m, Queensland" specimenCode="ANIC 32 - 047353" typeStatus="Holotype">
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Types.
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<typeStatus id="556BB5B7D4894A00C8444D4EE285077D">Holotype</typeStatus>
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worker from
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<location id="EFA780EBE912CE3D115078B8D6F26642" LSID="urn:lsid:plazi:treatment:57301B7B708B4B13F98661CD45DC99B0:EFA780EBE912CE3D115078B8D6F26642" country="Australia" name="Summit, Mt. Bellenden-Ker, 1700 m, Queensland">Summit, Mt. Bellenden-Ker, 1700m, Queensland</location>
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,
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<date id="3D68E30E493233C55877D5E42FBBB3B3" day="07" month="07" value="1971-07-07" year="1971">
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<collectingDate id="AD0DAD2CB7FCFE7AE34202D6E3734E92" day="07" month="07" value="1971-07-07" year="1971">7 July 1971</collectingDate>
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</date>
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,
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<collectorName id="256908536D10272E0A83DCCC4598FEF3">R. W. Taylor & J. Feehan</collectorName>
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, moss forest (
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<collectionCode id="5EEC531A0763689A6FD3CEBC1D2DAEB9" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">ANIC</collectionCode>
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,
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<specimenCode id="9862F67BFCA141B2507F58BEE0C2137D">ANIC 32-047353</specimenCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="C3EC1BE57F087E909BF7BED36EDF9CEA" ID-GBIF-Occurrence="923892992" collectionCode="ANIC, BMNH, MCZC" country="Australia" specimenCode="ANIC 32 - 047507" typeStatus="paratype">
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8
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<typeStatus id="5E53D5E504EADD8974F3EDC99C85BCB2">paratype</typeStatus>
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workers, same data as holotype (
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<collectionCode id="BA86B9AAC0E24299131CE06B4D30F1CB" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">ANIC</collectionCode>
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,
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<collectionCode id="600D7877C6C9498B246D3E826E4A0514" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]">BMNH</collectionCode>
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,
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<collectionCode id="8D42D38F1851CC417994E46E8DEF8D90" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
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,
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<specimenCode id="9386174789EC4A6D3ABCA3CB6BE11BF8">ANIC 32-047507</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="5CC5E55BBBE4DB4E7B77C42D649C68EB" type="diagnosis">
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<paragraph id="A6FE068B2B85341298A25F147C456A5E" pageNumber="8">Diagnosis. Dorsal surface of mesosoma with continuous longitudinal carinae running from pronotum to propodeum; carinae extending continuously from the posterodorsal surface onto the lateral surfaces of the anterior mesosoma (the carinae extend from the mesonotum to the sides of the pronotum), the sculpturing well developed on sides of pronotum. The form of the mesosomal sculpturing combined with the smooth region behind the compound eyes will separate this species from others known from Australia.</paragraph>
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</subSubSection>
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<subSubSection id="256DA2EA7244998B7E0F62D504216673" type="description">
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<paragraph id="28C3D1A766D5328FB6F4F12CCBF0B3D7" lastPageNumber="9" pageNumber="8">
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Worker description. Antennal scapes smooth or with low ridges. First segment of funiculus cone-shaped. Sides of head behind compound eyes smooth. Sculpturing on dorsal surface of mesosoma varying from running longitudinally and generally with the central carina (or carinae) running nearly the length (the
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<pageStartToken id="F58A1E30EF74563C7C31DB65233B3919" pageNumber="9">pronotum</pageStartToken>
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not differentiated from the mesonotum) to weakly "V"-shaped anteriorly. Carinae extending continuously from the dorsal surface onto the lateral surfaces of the mesosoma. Metanotal spines very short and broadly anglular. Propodeal spines long. Erect hairs abundant, straight. Colour dark brown-black, gaster slightly lighter, antennae, mandibles, legs and tip of gaster yellow.
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</paragraph>
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<caption id="0D032A28A6F5BE502B53D7882524EBEC">
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<paragraph id="AD9BE2FF1EA25488D3B78BB2E796D5D2" pageNumber="9">
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FIGURES 12-16.
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<taxonomicName id="97F6A04B99EF3FE386B864A10ABEE344" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:251840" LSID-ZBK="urn:lsid:zoobank.org:act:48886DE1-0866-4FC9-98F1-83987EEA86CA" class="Insecta" family="Formicidae" genus="Myrmecina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmecina difficulta" lsidName-ZBK="Myrmecina difficulta Shattuck 2009:8" order="Hymenoptera" pageNumber="9" phylum="Arthropoda" rank="species" species="difficulta">Myrmecina difficulta</taxonomicName>
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<taxonomicNameLabel id="6586BFD0610516E257F76C7965E30244" rank="species">sp. n.</taxonomicNameLabel>
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worker. Fig. 12, front of head; Fig. 13, lateral view of body; Fig. 14, dorsal view of body; Fig. 15, dorsal view of body; Fig. 16, dorsal view of body.
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</paragraph>
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</caption>
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<paragraph id="FDBE943C071F5BBA3AB596B8CE8A556C" pageNumber="9">Measurements. Worker (n = 24)- CI 93-107; HL 0.59-0.92; HW 0.54-0.91; MTL 0.28-0.49; SI 79-97; SL 0.45-0.82; WL 0.67-1.15.</paragraph>
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</subSubSection>
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<subSubSection id="F6D2BB9441B8A7762940366236F4806D" type="materials_examined">
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<paragraph id="EF16325F2D93291243749360A0E85300" lastPageNumber="10" pageNumber="9">
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<materialsCitation id="B39F296BB029785AB7EA46760F87DF75" ID-GBIF-Occurrence="923892990" country="Australia">
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Additional material examined (
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<collectionCode id="2BE2B8771E915D8CD152D40FA1D9574F" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">ANIC</collectionCode>
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except where noted). Queensland: 2.0km WNW Cape Tribulation (Site 2) (Monteith, Yeates & Thompson);
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</materialsCitation>
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<materialsCitation id="A60548A9D9CE9395998890B4A8502E14" ID-GBIF-Occurrence="923892997" country="Australia">4km E Lake Barrine (Taylor,R.W. & Feehan,J.E.);</materialsCitation>
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<materialsCitation id="0E2B23B45FD91981CE67735DEBAE9E21" ID-GBIF-Occurrence="923893006" country="Australia">
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4.3km NE Mt. Perseverance (Burwell,C.) (
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<collectionCode id="0B165C9B576125E056AAB00B755C08D1" collectionName="Australia, Queensland, South Brisbane, Queensland Museum">QMBA</collectionCode>
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);
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</materialsCitation>
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<materialsCitation id="EEED804E80FF7141C38265B447065F24" ID-GBIF-Occurrence="923892998" country="Australia">6.5km SSE Eungella (Gillison,A.);</materialsCitation>
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<materialsCitation id="F32158AECD5516A4F27E4910AEF734BA" ID-GBIF-Occurrence="923892993" country="Australia">Alexandra Bay (Taylor,R.W. & Feehan,J.);</materialsCitation>
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<materialsCitation id="35339C7C1E846D8C5750EFF1DC3317B8" ID-GBIF-Occurrence="923892994" country="Australia">Bellenden Ker Range, 1km S Cable Tower 6 (Earthwatch, Qld Museum);</materialsCitation>
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<materialsCitation id="5A08673F44906DFCCFAED50ABB729271" ID-GBIF-Occurrence="923893000" country="Australia">Bellenden Ker Range, Summit TV Stn (Earthwatch, Qld Museum;</materialsCitation>
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<materialsCitation id="7C46F142B435BDE7815AE73FFD7A0D3C" ID-GBIF-Occurrence="923892987" country="Australia">Monteith, Yeates & Thompson; Monteith,G.B. & Yeates,D.K.);</materialsCitation>
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<materialsCitation id="A4DEBFEAC3D50DA3A43E8F97F437802E" ID-GBIF-Occurrence="923892986" country="Australia">Bellenden Ker Summit (Monteith,G.B.);</materialsCitation>
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<materialsCitation id="CDF9DA8343FDCC254D5B561D29E83EFB" ID-GBIF-Occurrence="923892985" country="Australia">Bellenden Ker, Centre Peak Summit (Monteith,G.B.);</materialsCitation>
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<materialsCitation id="AD30087D807B87DA1792CABF5F64D41F" ID-GBIF-Occurrence="923892995" country="Australia">Brandy Ck. Rd., Conway S.F. (Monteith,G.B.);</materialsCitation>
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<materialsCitation id="C21D5E22F70E9148C7389718DE6C2027" ID-GBIF-Occurrence="923892989" country="Australia">Crystal Cascades, via Redlynch (Monteith,G.B.);</materialsCitation>
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<materialsCitation id="663A9CB950A1B4544589DE314D17323D" ID-GBIF-Occurrence="923893007" country="Australia">Hinchinbrook Is., Gayundah Ck. (Monteith, Davies, Thompson & Gallon);</materialsCitation>
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<materialsCitation id="DB1B2B276A221D4890E04F396F508EE2" ID-GBIF-Occurrence="923892996" country="Australia">
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Kuranda, Black
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<pageStartToken id="142D43AC45FE9FFD24D9A124DAF4B898" pageNumber="10">Mt</pageStartToken>
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. Road (Taylor,R.W. & Feehan,J.);
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</materialsCitation>
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<materialsCitation id="93F9ABFB79EABE827ED5902A8365D9DA" ID-GBIF-Occurrence="923893003" country="Australia">Mt. Bartle Frere, below NW Peak Summit (Monteith,G.);</materialsCitation>
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<materialsCitation id="378E75F09EB4CBDF6D1FE56C26B2184F" ID-GBIF-Occurrence="923893002" country="Australia">Mt. Bartle Frere, Summit NW Peak (Monteith,G. & Monteith,S.);</materialsCitation>
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<materialsCitation id="174C73E7653320D93E563B61EF570027" ID-GBIF-Occurrence="923893004" country="Australia">Mt. Dryander (Monteith,G.);</materialsCitation>
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<materialsCitation id="18DE31541134A6533F7FF6EE28040991" ID-GBIF-Occurrence="923893005" country="Australia">Mt. Elliot (Taylor,R.W.);</materialsCitation>
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<materialsCitation id="4F091815B7530D21D37331F15AE237C1" ID-GBIF-Occurrence="923893008" country="Australia">Mt. Fisher, 7km SW Millaa Millaa (Monteith, Yeates & Cook);</materialsCitation>
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<materialsCitation id="BED724D17449723E011B48B3987FCDE0" ID-GBIF-Occurrence="923893015" country="Australia">Mt. Tyson, 2km W Tully (Yeates,D.K.);</materialsCitation>
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<materialsCitation id="0E1C96B64ECDC1BB4176DD43A7E26539" ID-GBIF-Occurrence="923893027" country="Australia">Mt. Windsor Tableland (Taylor,R.W.);</materialsCitation>
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<materialsCitation id="18485FB89105ED5C2A80B5D28CA0198C" ID-GBIF-Occurrence="923893026" country="Australia">Mulgrave River Road, 7km WbyS Bellenden Ker (Calder,A. & Weir,T.);</materialsCitation>
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<materialsCitation id="A69A5A40A6914BEED3E31AF71014C0A0" ID-GBIF-Occurrence="923893030" country="Australia">Noah Creek (Taylor & Feehan);</materialsCitation>
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<materialsCitation id="C897894FCC24B65012819AF03AA26207" ID-GBIF-Occurrence="923893020" country="Australia">Noah Creek, 7km ENE Thornton Peak (Calder,A. & Weir,T.);</materialsCitation>
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<materialsCitation id="89E3E8181DB4372DE7D356D0EA11640F" ID-GBIF-Occurrence="923893016" country="Australia">Nob Creek, Byfield (Monteith,G.B.);</materialsCitation>
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<materialsCitation id="02618E05D0406270C2163EA7F1F822F3" ID-GBIF-Occurrence="923893009" country="Australia">Summit, Mt. Bellenden-Ker (Taylor,R.W. & Feehan,J.);</materialsCitation>
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<materialsCitation id="FDB52C3EA426A9666A51E19B7344E734" ID-GBIF-Occurrence="923893011" country="Australia">Thornton Range (Taylor,R.W. & Feehan,J.);</materialsCitation>
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<materialsCitation id="B70F2995DD104093BD31B6F357C8702C" ID-GBIF-Occurrence="923893028" country="Australia">Tully Falls Nat. Pk (Taylor,R.W. & Feehan,J.E.).</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B5E500A3220567B4179987903C2E6415" type="discussion">
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<paragraph id="FD7B123809DC249069E8F64ED817034F" pageNumber="10">Comments. This rainforest and palm forest species has been collected from leaf litter and under stones.</paragraph>
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<paragraph id="D4785B41FBB589ACD0029CB8F100138E" pageNumber="10">
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<taxonomicName id="921BAABD47552C71436F8253399D637B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:251840" LSID-ZBK="urn:lsid:zoobank.org:act:48886DE1-0866-4FC9-98F1-83987EEA86CA" class="Insecta" family="Formicidae" genus="Myrmecina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmecina difficulta" lsidName-ZBK="Myrmecina difficulta Shattuck 2009:8" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="species" species="difficulta">Myrmecina difficulta</taxonomicName>
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shows considerable morphological variation in a number of character systems, including the size, number and shape/configuration of carinae on the head and mesosoma, the development of sculpturing in the area behind the compound eye, the length of the metanotal spines and the overall body size. In some cases these characters show as much variation as that seen between other species in the genus, strongly suggesting that multiple closely related species are involved. For example the density of carinae on the mesonotum shows considerable variation, ranging from large and widely spaced to smaller and more dense. Similarly, body size is extreme in this species, showing more variation than that found in all other species combined - a highly unexpected and surprising condition.
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</paragraph>
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<paragraph id="ED4AA89E55C1D7572E2509C8FE6D65B3" pageNumber="10">However, this variation was found to be largely continuous and lacking obvious subdivisions. Additionally, there are numerous examples where significant variation is seen among specimens collected from the same litter sample, suggesting that this variation is intra- rather than inter-specific.</paragraph>
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<paragraph id="9C23BC40E3B43D8FE1BCF019C8848ECE" pageNumber="10">Additionally, the variation across characters showed little correlation. For example, many of the smaller specimens show a tendency to have the carinae on the dorsum of the pronotum in the form of a "V", a pattern also seen in some of the largest workers but absent from the majority of intermediately sized specimens (where the carinae are more or less linear).</paragraph>
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<paragraph id="2E16C1900BEB34A487B9BD07769D83DC" pageNumber="10">Combined, the lack of congruence among characters and the considerable variation found within individual characters make it difficult or impossible to subdivide the specimens treated here into diagnosable subsets. As a result these specimens are treated as belonging to a single heterogeneous species. It would be worth revisiting this conclusion as additional material becomes available.</paragraph>
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</subSubSection>
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</treatment>
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</document> |