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<mods:title id="44E1AF7D5660182321DBC1E2879A1CBE">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<emphasis id="A4DEEAF6FFFCFFD644A8F95E91B673E7" bold="true" box="[151,387,1759,1786]" italics="true" pageId="22" pageNumber="97">Tenupalpa biformis</emphasis>
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<bibRefCitation id="F23B4B15FFFCFFD645ACF961927773E7" author="Omelko, M. M." box="[403,578,1760,1786]" pageId="22" pageNumber="97" pagination="28 - 34" refId="ref23940" refString="Omelko, M. M. (1984) Description of three new species of Gelechiidae from South Primorye. The Fauna and Ecology of Insects in the South of the Far East, 112 (215), 28 - 34. [in Russian]" type="journal article" year="1984">Omelko, 1984</bibRefCitation>
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<figureCitation id="0E912A61FFFCFFD644A0F88590C37203" box="[159,246,1796,1822]" captionStart="FIGURES 1525" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 1525. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="22" pageNumber="97">Figs 19</figureCitation>
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<figureCitation id="0E912A61FFFCFFD6453DF88591157202" box="[258,288,1796,1823]" captionStart="FIGURES 2645" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 2645. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="22" pageNumber="97">35</figureCitation>
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<figureCitation id="0E912A61FFFCFFD64513F885917F7202" box="[300,330,1796,1823]" captionStart="FIGURES 5461" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 5461. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="22" pageNumber="97">55</figureCitation>
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<figureCitation id="0E912A61FFFCFFD64569F88591407203" box="[342,373,1796,1822]" captionStart="FIGURES 7277" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 7277. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="22" pageNumber="97">74</figureCitation>
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<figureCitation id="0E912A61FFFCFFD645BFF88591A87203" box="[384,413,1796,1822]" captionStart="FIGURES 8790" captionStartId="19.[151,264,1943,1967]" captionTargetBox="[229,1327,198,1905]" captionTargetId="figure-18@19.[196,1391,181,1920]" captionTargetPageId="19" captionText="FIGURES 8790. Female genitalia of Tenupalpa spp. 87. T. immaculata sp. nov., paratype, slide no. LGE18002; 88. T. isabellina sp. nov., paratype, slide no. LGE17283f; 89. T. flavitermina, slide no. LGE19021; 90. T. biformis, slide no. LLJ15251 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819832" httpUri="https://zenodo.org/record/5819832/files/figure.png" pageId="22" pageNumber="97">90</figureCitation>
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<emphasis id="A4DEEAF6FFFCFFD644A8F8C99146727D" box="[151,371,1864,1888]" italics="true" pageId="22" pageNumber="97">Cnaphostola biformis</emphasis>
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<emphasis id="A4DEEAF6FFFCFFD644A8F80791697283" box="[151,348,1926,1950]" italics="true" pageId="22" pageNumber="97">Thiotricha biformis</emphasis>
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Sohn
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., 2016: 116
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Kyaw
<emphasis id="A4DEEAF6FFFCFFD646ADF806928A7283" box="[658,703,1926,1950]" italics="true" pageId="22" pageNumber="97">et al</emphasis>
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<paragraph id="961536E4FFFCFFD644A8F825920E72A0" blockId="22.[151,1437,1864,1981]" box="[151,571,1956,1981]" pageId="22" pageNumber="97">
<treatmentCitationGroup id="B6BA11CAFFFCFFD644A8F825920E72A0" box="[151,571,1956,1981]" pageId="22" pageNumber="97">
<treatmentCitation id="170B10F5FFFCFFD644A8F825920272A0" author="Lee, G. - E. &amp; Han, T. M. &amp; Park, H. C. &amp; Qi, M. J. &amp; Li, H. H." box="[151,567,1956,1981]" page="371" pageId="22" pageNumber="97" year="2021">
<taxonomicName id="51AA4D67FFFCFFD644A8F825920272A0" ID-CoL="98YHZ" authority="Lee et al., 2021: 371" authorityName="Lee" authorityPageNumber="371" authorityYear="2021" box="[151,567,1956,1981]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="97" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFFCFFD644A8F825916972A1" box="[151,348,1956,1980]" italics="true" pageId="22" pageNumber="97">Tenupalpa biformis</emphasis>
<bibRefCitation id="F23B4B15FFFCFFD6455DF824920272A0" author="Lee, G. - E. &amp; Han, T. M. &amp; Park, H. C. &amp; Qi, M. J. &amp; Li, H. H." box="[354,567,1956,1981]" pageId="22" pageNumber="97" pagination="357 - 379" refId="ref23692" refString="Lee, G. - E., Han, T. M., Park, H. C., Qi, M. J. &amp; Li, H. H. (2021) A phylogeny of the subfamily Thiotrichinae (Lepidoptera: Gel- echiidae) with a revision of the generic classification based on molecular and morphological analyses. Systematic Entomology, 46, 357 - 379. https: // doi. org / 10.1111 / syen. 12466" type="journal article" year="2021">
Lee
<emphasis id="A4DEEAF6FFFCFFD645B0F82491F772A1" box="[399,450,1956,1980]" italics="true" pageId="22" pageNumber="97">et al.</emphasis>
, 2021: 371
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFDFFD744A8FF17951175E4" pageId="23" pageNumber="98" type="materials_examined">
<paragraph id="961536E4FFFDFFD744A8FF17951175E4" blockId="23.[151,1437,150,537]" pageId="23" pageNumber="98">
<emphasis id="A4DEEAF6FFFDFFD744A8FF17914B75AD" bold="true" box="[151,382,150,176]" pageId="23" pageNumber="98">Material examined.</emphasis>
<materialsCitation id="26C23CB9FFFDFFD745BDFF1793DC75C8" ID-GBIF-Occurrence="3427696304" accessionNumber="LLJ15140" collectingDate="2014-07-06" collectorName="Aihui Yin &amp; Xuemei Hu &amp; Qingyun Wang" country="China" elevation="555" latitude="30.33" location="Mt. Tianmu" longLatPrecision="733" longitude="119.4" municipality="Lao'an" pageId="23" pageNumber="98" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang">
<emphasis id="A4DEEAF6FFFDFFD745BDFF1792E575AD" bold="true" box="[386,720,150,176]" pageId="23" pageNumber="98">
<collectingCountry id="EEBD7674FFFDFFD745BDFF1791D175AD" box="[386,484,150,176]" name="China" pageId="23" pageNumber="98">CHINA</collectingCountry>
:
<collectingRegion id="546EF806FFFDFFD745D2FF1792FE75AD" box="[493,715,150,176]" country="China" name="Zhejiang" pageId="23" pageNumber="98">Zhejiang Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFFDFFD746EBFF1792C875AC" box="[724,765,150,177]" pageId="23" pageNumber="98" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFFDFFD74738FF19936675AE" box="[775,851,152,179]" pageId="23" pageNumber="98">Laoan</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFFDFFD74760FF1793F775AD" box="[863,962,150,176]" degrees="30.33" direction="north" orientation="latitude" pageId="23" pageNumber="98" precision="555" value="30.33">30.33°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFFDFFD747F5FF17940375AD" box="[970,1078,150,176]" degrees="119.40" direction="east" orientation="longitude" pageId="23" pageNumber="98" precision="555" value="119.4">119.40°E</geoCoordinate>
),
<quantity id="51529B01FFFDFFD74078FF1794BB75AC" box="[1095,1166,150,177]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="23" pageNumber="98" unit="m" value="555.0">
<elevation id="1D87D1D7FFFDFFD74078FF1794BB75AC" box="[1095,1166,150,177]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="23" pageNumber="98" unit="m" value="555.0">555 m</elevation>
</quantity>
,
<location id="9375603FFFFDFFD740A8FF16952E75AD" LSID="urn:lsid:plazi:treatment:1E0387F2FFFCFFD744A8F95E93067705:9375603FFFFDFFD740A8FF16952E75AD" box="[1175,1307,150,177]" country="China" latitude="30.33" longLatPrecision="733" longitude="119.4" municipality="Lao'an" name="Mt. Tianmu" pageId="23" pageNumber="98" stateProvince="Zhejiang">Mt.Tianmu</location>
,
<date id="E2141024FFFDFFD7411BFF1795AD75AD" box="[1316,1432,150,176]" pageId="23" pageNumber="98" value="2014-07-06">
<collectingDate id="F250E9CCFFFDFFD7411BFF1795AD75AD" box="[1316,1432,150,176]" pageId="23" pageNumber="98" value="2014-07-06">6.vii.2014</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFFDFFD744FAFF3B910175C9" box="[197,308,186,212]" pageId="23" pageNumber="98">Aihui Yin</collectorName>
,
<collectorName id="3B5F5332FFFDFFD74500FF3B918A75C8" box="[319,447,186,213]" pageId="23" pageNumber="98">Xuemei Hu</collectorName>
and
<collectorName id="3B5F5332FFFDFFD745CBFF3B92AA75C8" box="[500,671,186,213]" pageId="23" pageNumber="98">Qingyun Wang</collectorName>
, genitalia slide no. LLJ15140
</materialsCitation>
;
<materialsCitation id="26C23CB9FFFDFFD747CBFF3B951575E4" ID-GBIF-Occurrence="3427696401" accessionNumber="LLJ15251" collectingDate="2015-07-17" collectorName="Aihui Yin &amp; Kang Lou &amp; Tao Wang" country="China" elevation="789" latitude="30.37" location="Mt. Tianmu" longLatPrecision="733" longitude="119.43" municipality="Sanmuping" pageId="23" pageNumber="98" specimenCount="1" specimenCount-female="1" stateProvince="Zhejiang">
<specimenCount id="80ACFD6DFFFDFFD747CBFF3B942875C8" box="[1012,1053,186,213]" pageId="23" pageNumber="98" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="7671AC9EFFFDFFD74016FF3B949C75C9" box="[1065,1193,186,212]" pageId="23" pageNumber="98">Sanmuping</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFFDFFD74088FF3B952C75C9" box="[1207,1305,186,212]" degrees="30.37" direction="north" orientation="latitude" pageId="23" pageNumber="98" precision="555" value="30.37">30.37°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFFDFFD7411BFF3B95BA75C9" box="[1316,1423,186,212]" degrees="119.43" direction="east" orientation="longitude" pageId="23" pageNumber="98" precision="555" value="119.43">119.43°E</geoCoordinate>
),
<quantity id="51529B01FFFDFFD744A8FF5F90D575E4" box="[151,224,222,249]" metricMagnitude="2" metricUnit="m" metricValue="7.89" pageId="23" pageNumber="98" unit="m" value="789.0">
<elevation id="1D87D1D7FFFDFFD744A8FF5F90D575E4" box="[151,224,222,249]" metricMagnitude="2" metricUnit="m" metricValue="7.89" pageId="23" pageNumber="98" unit="m" value="789.0">789 m</elevation>
</quantity>
,
<location id="9375603FFFFDFFD744D3FF5E914675E5" LSID="urn:lsid:plazi:treatment:1E0387F2FFFCFFD744A8F95E93067705:9375603FFFFDFFD744D3FF5E914675E5" box="[236,371,222,249]" country="China" latitude="30.37" longLatPrecision="733" longitude="119.43" municipality="Sanmuping" name="Mt. Tianmu" pageId="23" pageNumber="98" stateProvince="Zhejiang">Mt. Tianmu</location>
,
<date id="E2141024FFFDFFD74541FF5F923475E4" box="[382,513,222,249]" pageId="23" pageNumber="98" value="2015-07-17">
<collectingDate id="F250E9CCFFFDFFD74541FF5F923475E4" box="[382,513,222,249]" pageId="23" pageNumber="98" value="2015-07-17">17.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFFDFFD74605FF5F929E75E5" box="[570,683,222,248]" pageId="23" pageNumber="98">Aihui Yin</collectorName>
,
<collectorName id="3B5F5332FFFDFFD74688FF5E931275E4" box="[695,807,223,249]" pageId="23" pageNumber="98">Kang Lou</collectorName>
and
<collectorName id="3B5F5332FFFDFFD74762FF5E93E475E4" box="[861,977,223,249]" pageId="23" pageNumber="98">Tao Wang</collectorName>
, genitalia slide no. LLJ15251
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFDFFD744F8FE83957A74B0" pageId="23" pageNumber="98" type="diagnosis">
<paragraph id="961536E4FFFDFFD744F8FE83957A74B0" blockId="23.[151,1437,150,537]" pageId="23" pageNumber="98">
<emphasis id="A4DEEAF6FFFDFFD744F8FE8391777401" bold="true" box="[199,322,258,284]" pageId="23" pageNumber="98">Diagnosis.</emphasis>
Adult wingspan 8.510.0 mm.
<taxonomicName id="51AA4D67FFFDFFD7469FFE83934C7401" box="[672,889,258,284]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="98" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFFDFFD7469FFE83934C7401" box="[672,889,258,284]" italics="true" pageId="23" pageNumber="98">Tenupalpa biformis</emphasis>
</taxonomicName>
is diagnosed by the glossy white forewing with a broad dark grey area in distal half, an orange distal patch and a black apical spot. The male genitalia are characterized by the apically pointed, large thumb-like anellus lobes and the juxta with a short-forked projection. The female genitalia are similar to
<taxonomicName id="51AA4D67FFFDFFD745A9FEEE92147495" baseAuthorityName="Omelko" baseAuthorityYear="1984" box="[406,545,366,392]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="98" phylum="Tracheophyta" rank="species" species="venustalis">
<emphasis id="A4DEEAF6FFFDFFD745A9FEEE92147495" box="[406,545,366,392]" italics="true" pageId="23" pageNumber="98">T. venustalis</emphasis>
</taxonomicName>
as seen in the short apophyses anteriores and the corpus bursae without signum, but
<taxonomicName id="51AA4D67FFFDFFD744FDFE12910F74B1" box="[194,314,402,428]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="98" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFFDFFD744FDFE12910F74B1" box="[194,314,402,428]" italics="true" pageId="23" pageNumber="98">T. biformis</emphasis>
</taxonomicName>
can be distinguished from the latter by the medially elongate anterior margin of tergum VIII.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFDFFD744F8FE3794F774CD" box="[199,1218,438,465]" pageId="23" pageNumber="98" type="distribution">
<paragraph id="961536E4FFFDFFD744F8FE3794F774CD" blockId="23.[151,1437,150,537]" box="[199,1218,438,465]" pageId="23" pageNumber="98">
<emphasis id="A4DEEAF6FFFDFFD744F8FE37916C74CD" bold="true" box="[199,345,438,464]" pageId="23" pageNumber="98">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFFDFFD74558FE37919F74CD" box="[359,426,438,464]" name="China" pageId="23" pageNumber="98">China</collectingCountry>
(
<collectingRegion id="546EF806FFFDFFD74586FE37921674CC" box="[441,547,438,465]" country="China" name="Zhejiang" pageId="23" pageNumber="98">Zhejiang</collectingRegion>
),
<collectingCountry id="EEBD7674FFFDFFD74609FE36924C74CC" box="[566,633,439,465]" name="Japan" pageId="23" pageNumber="98">Japan</collectingCountry>
,
<collectingCountry id="EEBD7674FFFDFFD746BCFE3792E474CD" box="[643,721,438,464]" name="Russia" pageId="23" pageNumber="98">Russia</collectingCountry>
. This species is newly recorded from
<collectingCountry id="EEBD7674FFFDFFD74047FE37948A74CD" box="[1144,1215,438,464]" name="China" pageId="23" pageNumber="98">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFDFFD744F8FE5B93067705" pageId="23" pageNumber="98" type="discussion">
<paragraph id="961536E4FFFDFFD744F8FE5B93067705" blockId="23.[151,1437,150,537]" pageId="23" pageNumber="98">
<emphasis id="A4DEEAF6FFFDFFD744F8FE5B910E74E9" bold="true" box="[199,315,474,500]" pageId="23" pageNumber="98">Remarks.</emphasis>
The
<typeStatus id="49118846FFFDFFD7454AFE5A919074E8" box="[373,421,475,501]" pageId="23" pageNumber="98">type</typeStatus>
materials deposited in ZIRAS of
<collectingCountry id="EEBD7674FFFDFFD74720FE5B935F74E9" box="[799,874,474,500]" name="Russia" pageId="23" pageNumber="98">Russia</collectingCountry>
were not examined in this study. A detailed re-description of
<taxonomicName id="51AA4D67FFFDFFD74521FE7E91A27705" box="[286,407,510,536]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="98" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFFDFFD74521FE7E91A27705" box="[286,407,510,536]" italics="true" pageId="23" pageNumber="98">T. biformis</emphasis>
</taxonomicName>
was provided in
<bibRefCitation id="F23B4B15FFFDFFD74667FE7E931A7705" author="Kyaw, K. M. M. &amp; Yagi, S. &amp; Oku, J. &amp; Sakamaki, Y. &amp; Hirowatari, T." box="[600,815,510,537]" pageId="23" pageNumber="98" pagination="67 - 99" refId="ref23620" refString="Kyaw, K. M. M., Yagi, S., Oku, J., Sakamaki, Y. &amp; Hirowatari, T. (2019) Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys, 897, 67 - 99." type="journal article" year="2019">
Kyaw
<emphasis id="A4DEEAF6FFFDFFD7469DFE7E92E67705" box="[674,723,510,536]" italics="true" pageId="23" pageNumber="98">et al</emphasis>
. (2019)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>