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<mods:title id="16109F28363EDF5FB6AF3589C286B2E6">A new species of Thinophilus Wahlberg (Diptera: Dolichopodidae: Hydrophorinae) from coastal India with re-description of Thinophilus tesselatus Becker (Diptera: Dolichopodidae: Hydrophorinae)</mods:title>
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Mukherjee &amp; Pramanik
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(
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Thinophilus maritimus sp. nov., holotype, ♂, lateral habitus (inset: hind coxa without strong lateral seta)." captionText-1="FIGURE 2. Thinophilus maritimus sp. nov., holotype, ♂, dorsal habitus." captionText-10="FIGURE 11. Thinophilus maritimus sp. nov., paratype, female terminalia." captionText-11="FIGURE 12. Two male individuals of Thinophilus maritimus sp. nov. on moist sandy beach." captionText-2="FIGURE 3. Thinophilus maritimus sp. nov., holotype, ♂, head A. Frontal view; B. Lateral view; C. Magnified lateral view showing antennae." captionText-3="FIGURE 4. Thinophilus maritimus sp. nov., holotype, ♂, lateral view of thorax." captionText-4="FIGURE 5. Thinophilus maritimus sp. nov., holotype, ♂, wing venation." captionText-5="FIGURE 6. Thinophilus maritimus sp. nov., holotype, ♂, leg. A. Fore and mid legs; B. Hind tibia and tarsus" captionText-6="FIGURE 7. Thinophilus maritimus sp. nov., holotype, ♂, abdomen. A. Ventral view; B. Dorsal view." captionText-7="FIGURE 8. Thinophilus maritimus sp. nov., holotype, ♂, male terminalia A. Lateral view; B. Dorsal view.Abbreviations: Cerc: Cerci, Ph: Phallus, Sur: Surstylus." captionText-8="FIGURE 9. Thinophilus maritimus sp. nov., holotype, ♂, male terminalia A. Lateral view of phallus and phallosome; B. Dorsal view of the tip of surstylus; C. Dorsal view of cercus." captionText-9="FIGURE 10. Thinophilus maritimus sp. nov., paratype, lateral habitus of female." figureDoi-0="http://doi.org/10.5281/zenodo.10202834" figureDoi-1="http://doi.org/10.5281/zenodo.10202836" figureDoi-10="http://doi.org/10.5281/zenodo.10202852" figureDoi-11="http://doi.org/10.5281/zenodo.10202854" figureDoi-2="http://doi.org/10.5281/zenodo.10202838" figureDoi-3="http://doi.org/10.5281/zenodo.10202840" figureDoi-4="http://doi.org/10.5281/zenodo.10202842" figureDoi-5="http://doi.org/10.5281/zenodo.10202844" figureDoi-6="http://doi.org/10.5281/zenodo.10202846" figureDoi-7="http://doi.org/10.5281/zenodo.10202848" figureDoi-8="http://doi.org/10.5281/zenodo.10202850" figureDoi-9="http://doi.org/10.5281/zenodo.10277925" httpUri-0="https://zenodo.org/record/10202834/files/figure.png" httpUri-1="https://zenodo.org/record/10202836/files/figure.png" httpUri-10="https://zenodo.org/record/10202852/files/figure.png" httpUri-11="https://zenodo.org/record/10202854/files/figure.png" httpUri-2="https://zenodo.org/record/10202838/files/figure.png" httpUri-3="https://zenodo.org/record/10202840/files/figure.png" httpUri-4="https://zenodo.org/record/10202842/files/figure.png" httpUri-5="https://zenodo.org/record/10202844/files/figure.png" httpUri-6="https://zenodo.org/record/10202846/files/figure.png" httpUri-7="https://zenodo.org/record/10202848/files/figure.png" httpUri-8="https://zenodo.org/record/10202850/files/figure.png" httpUri-9="https://zenodo.org/record/10277925/files/figure.png" pageId="3" pageNumber="481">Figs 1–12</figureCitation>
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/H 
<materialsCitation id="A7D76E3EFFB9130FFE38FD08FD48FD13" ID-GBIF-Occurrence="4441575303" collectingDate="2021-08-07" collectorName="K. Mukherjee &amp; D. Pramanik." country="India" latitude="21.556719" location="Bakkhali" longLatPrecision="1" longitude="88.27095" municipality="Bay of Bengal" pageId="3" pageNumber="481" specimenCount="6" specimenCount-female="6" stateProvince="West Bengal" typeStatus="paratype">
<specimenCount id="01B9AFEAFFB9130FFE38FD08FE0DFD30" box="[452,493,705,732]" count="6" pageId="3" pageNumber="481" type="female">6 ♀</specimenCount>
, 
<location id="126032B8FFB9130FFE07FD08FD83FD37" LSID="urn:lsid:plazi:treatment:9F16D575FFB91309FF6BFDC4FE11FEF1:126032B8FFB9130FFE07FD08FD83FD37" box="[507,611,705,731]" country="India" latitude="21.556719" longLatPrecision="1" longitude="88.27095" municipality="Bay of Bengal" name="Bakkhali" pageId="3" pageNumber="481" stateProvince="West Bengal">Bakkhali</location>
, 
<collectingMunicipality id="F764FE19FFB9130FFD91FD0BFCF1FD37" box="[621,785,705,732]" pageId="3" pageNumber="481">Bay of Bengal</collectingMunicipality>
, 
<geoCoordinate id="728B02A4FFB9130FFCE0FD0AFC2DFD32" box="[796,973,707,734]" degrees="21" direction="north" minutes="33" orientation="latitude" pageId="3" pageNumber="481" precision="1" seconds="24.19" value="21.556719">21°33′24.19″ N</geoCoordinate>
, 
<geoCoordinate id="728B02A4FFB9130FFC2DFD0AFB60FD30" box="[977,1152,706,734]" degrees="88" direction="east" minutes="16" orientation="longitude" pageId="3" pageNumber="481" precision="1" seconds="15.40" value="88.27095">88°16′15.40″ E</geoCoordinate>
, alt 0 m, sweep netting, 
<date id="630142A3FFB9130FFA72FD0BFEF0FD13" pageId="3" pageNumber="481" value="2021-08-07">
<collectingDate id="7345BB4BFFB9130FFA72FD0BFEF0FD13" pageId="3" pageNumber="481" value="2021-08-07">7 Aug. 2021</collectingDate>
</date>
, coll. 
<collectorName id="BA4A01B5FFB9130FFEAFFD2FFE0FFD13" box="[339,495,741,768]" pageId="3" pageNumber="481">K. Mukherjee</collectorName>
&amp; 
<collectorName id="BA4A01B5FFB9130FFDEFFD2FFD48FD13" box="[531,680,741,768]" pageId="3" pageNumber="481">D. Pramanik.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5FA537E8FFB9130FFF3BFCC0FBE8FC3B" pageId="3" pageNumber="481" type="diagnosis">
<paragraph id="17006463FFB9130FFF3BFCC0FBE8FC3B" blockId="3.[151,1437,633,2028]" pageId="3" pageNumber="481">
<emphasis id="25CBB871FFB9130FFF3BFCC0FEA2FCCF" bold="true" box="[199,322,777,803]" pageId="3" pageNumber="481">Diagnosis.</emphasis>
A large species, mostly dark metallic green with whitish greyish tomentum. Scape, pedicel and postpedicel mostly brown; bulbous postpedicel; arista white except for brown base. 9–10 dorsocentral bristle decreasing in length anteriorly. All coxae metallic green with whitish tomentum; hind coxa lacks lateral setae. All tarsomeres orange-yellow with dark brown bands at the tip of each; terminal tarsomere mostly dark. Mid basitarsus with a protuberance. Hind femur twice as long as fore and mid femora; hind tibia moderately swollen with modified basitarsus which appears as a bifurcated process at the apical end of hind tibia.
</paragraph>
</subSubSection>
<subSubSection id="5FA537E8FFB91308FF3BFC28FC8AFCF9" lastPageId="4" lastPageNumber="482" pageId="3" pageNumber="481" type="description">
<paragraph id="17006463FFB91308FF3BFC28FC8AFCF9" blockId="3.[151,1437,633,2028]" lastBlockId="4.[151,1437,151,789]" lastPageId="4" lastPageNumber="482" pageId="3" pageNumber="481">
<emphasis id="25CBB871FFB9130FFF3BFC28FE7AFC17" bold="true" box="[199,410,993,1019]" pageId="3" pageNumber="481">Description. Male</emphasis>
(
<figureCitation id="8F8478E6FFB9130FFE5AFC28FDEAFC17" box="[422,522,993,1019]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[152,255,1945,1969]" captionStartId-1="2.[152,255,1979,2003]" captionTargetBox-0="[169,1419,887,1914]" captionTargetBox-1="[170,1418,963,1950]" captionTargetId-0="figure-270@1.[164,1423,882,1919]" captionTargetId-1="figure-353@2.[164,1423,957,1956]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Thinophilus maritimus sp. nov., holotype, ♂, lateral habitus (inset: hind coxa without strong lateral seta)." captionText-1="FIGURE 2. Thinophilus maritimus sp. nov., holotype, ♂, dorsal habitus." figureDoi-0="http://doi.org/10.5281/zenodo.10202834" figureDoi-1="http://doi.org/10.5281/zenodo.10202836" httpUri-0="https://zenodo.org/record/10202834/files/figure.png" httpUri-1="https://zenodo.org/record/10202836/files/figure.png" pageId="3" pageNumber="481">Figs 1–2</figureCitation>
). Body length 6.0– 
<quantity id="D047C986FFB9130FFD26FC28FCCFFC10" box="[730,815,993,1020]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="481" unit="mm" value="6.8">6.8 mm</quantity>
, wing length 
<quantity id="D047C986FFB9130FFC43FC28FBA7FC10" box="[959,1095,993,1020]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.2" metricValueMin="4.5" pageId="3" pageNumber="481" unit="mm" value="4.85" valueMax="5.2" valueMin="4.5">4.5–5.2 mm</quantity>
. 
<emphasis id="25CBB871FFB9130FFBB3FC28FB6FFC17" bold="true" box="[1103,1167,993,1019]" pageId="3" pageNumber="481">Head</emphasis>
(
<figureCitation id="8F8478E6FFB9130FFB60FC28FB3EFC17" box="[1180,1246,993,1019]" captionStart="FIGURE 3" captionStartId="4.[152,255,1155,1179]" captionTargetBox="[151,1435,813,1131]" captionTargetId="figure-317@4.[151,1435,813,1131]" captionTargetPageId="4" captionText="FIGURE 3. Thinophilus maritimus sp. nov., holotype, ♂, head A. Frontal view; B. Lateral view; C. Magnified lateral view showing antennae." figureDoi="http://doi.org/10.5281/zenodo.10202838" httpUri="https://zenodo.org/record/10202838/files/figure.png" pageId="3" pageNumber="481">Fig. 3</figureCitation>
): Eyes dichoptic, red with dense white pubescence, widely separated at ocellar region; postcranium dark metallic green, white pollinose; vertex slightly sunken, metallic green, whitish tomentum. Metallic green frons with violet reflections and whitish tomentum denser than on vertex; 2 proclinate ocellar setae present; 6 black upper postocular setae present along with dense white hairs on postgena; pair of inner vertical setae present; subtriangular clypeus with white tomentum; white triangular palpus with white hairs (
<figureCitation id="8F8478E6FFB9130FFD16FB5CFC85FB43" box="[746,869,1173,1199]" captionStart="FIGURE 3" captionStartId="4.[152,255,1155,1179]" captionTargetBox="[151,1435,813,1131]" captionTargetId="figure-317@4.[151,1435,813,1131]" captionTargetPageId="4" captionText="FIGURE 3. Thinophilus maritimus sp. nov., holotype, ♂, head A. Frontal view; B. Lateral view; C. Magnified lateral view showing antennae." figureDoi="http://doi.org/10.5281/zenodo.10202838" httpUri="https://zenodo.org/record/10202838/files/figure.png" pageId="3" pageNumber="481">Fig. 3A, B</figureCitation>
). Antennal scape brown along basal three-fourths, distally yellowish, apex curved inside; pedicel brown with white tomentum bearing small white hairs at base; postpedicel bulbous, brown with white tomentum, longer than wide; arista dorsal, bare, white, base swollen and dark brown (
<figureCitation id="8F8478E6FFB9130FFED4FAC8FE60FAF7" box="[296,384,1281,1307]" captionStart="FIGURE 3" captionStartId="4.[152,255,1155,1179]" captionTargetBox="[151,1435,813,1131]" captionTargetId="figure-317@4.[151,1435,813,1131]" captionTargetPageId="4" captionText="FIGURE 3. Thinophilus maritimus sp. nov., holotype, ♂, head A. Frontal view; B. Lateral view; C. Magnified lateral view showing antennae." figureDoi="http://doi.org/10.5281/zenodo.10202838" httpUri="https://zenodo.org/record/10202838/files/figure.png" pageId="3" pageNumber="481">Fig. 3C</figureCitation>
); postpedicel equal in length to scape; length ratio of scape to pedicel to postpedicel to arista, 0.24/0.06/0.18/0.37. 
<emphasis id="25CBB871FFB9130FFE7EFAECFE3BFAD3" bold="true" box="[386,475,1317,1343]" pageId="3" pageNumber="481">Thorax</emphasis>
(
<figureCitation id="8F8478E6FFB9130FFE10FAECFDD4FAD3" box="[492,564,1317,1343]" captionStart="FIGURE 4" captionStartId="5.[152,255,1296,1320]" captionTargetBox="[151,1435,309,1273]" captionTargetId="figure-76@5.[151,1435,309,1273]" captionTargetPageId="5" captionText="FIGURE 4. Thinophilus maritimus sp. nov., holotype, ♂, lateral view of thorax." figureDoi="http://doi.org/10.5281/zenodo.10202840" httpUri="https://zenodo.org/record/10202840/files/figure.png" pageId="3" pageNumber="481">Fig. 4</figureCitation>
): Metallic green with purple reflections, whitish tomentum throughout; dark green pleura with denser whitish tomentum than scutum; acrostichals absent; 9–10 dorsocentral bristles, decreasing in length anteriorly; 1 humeral; 1 posthumeral; 1 notopleural; 2 supra-alars; postalar absent; anterior spiracle white; scutellum concolorous with thorax, 2 strong longer marginals and 2 shorter hair-like laterals. 
<emphasis id="25CBB871FFB9130FFB4FFA58FB14FA47" bold="true" box="[1203,1268,1425,1451]" pageId="3" pageNumber="481">Wing</emphasis>
(
<figureCitation id="8F8478E6FFB9130FFAF8FA58FAA8FA40" box="[1284,1352,1425,1452]" captionStart="FIGURE 5" captionStartId="5.[152,255,2000,2024]" captionTargetBox="[156,1430,1350,1971]" captionTargetId="figure-94@5.[151,1435,1344,1976]" captionTargetPageId="5" captionText="FIGURE 5. Thinophilus maritimus sp. nov., holotype, ♂, wing venation." figureDoi="http://doi.org/10.5281/zenodo.10202842" httpUri="https://zenodo.org/record/10202842/files/figure.png" pageId="3" pageNumber="481">Fig. 5</figureCitation>
): Grey, simple, narrow; M 
<subScript id="8B3B6626FFB9130FFE94FA01FE90FA3A" attach="left" box="[360,368,1480,1494]" fontSize="6" pageId="3" pageNumber="481">1</subScript>
moderately sinuous after dm-m crossvein; R 
<subScript id="8B3B6626FFB9130FFC96FA01FC64FA3A" attach="left" box="[874,900,1480,1494]" fontSize="6" pageId="3" pageNumber="481">4+5</subScript>
and M 
<subScript id="8B3B6626FFB9130FFC2FFA01FC3BFA3A" attach="left" box="[979,987,1480,1494]" fontSize="6" pageId="3" pageNumber="481">1</subScript>
converge at apical area but do not meet each other; ratio of costal section between R 
<subScript id="8B3B6626FFB9130FFD7FFA25FD7CFA16" attach="left" box="[643,668,1516,1530]" fontSize="6" pageId="3" pageNumber="481">2+3</subScript>
and R 
<subScript id="8B3B6626FFB9130FFD18FA25FD1EFA16" attach="left" box="[740,766,1516,1530]" fontSize="6" pageId="3" pageNumber="481">4+5</subScript>
to costal section between R 
<subScript id="8B3B6626FFB9130FFBC8FA25FBADFA16" attach="left" box="[1076,1101,1516,1530]" fontSize="6" pageId="3" pageNumber="481">4+5</subScript>
and M 
<subScript id="8B3B6626FFB9130FFB67FA25FB43FA16" attach="left" box="[1179,1187,1516,1530]" fontSize="6" pageId="3" pageNumber="481">1</subScript>
6:4; ratio of crossvein dm-m to distal part of M 
<subScript id="8B3B6626FFB9130FFE55F9D9FE51F9F2" attach="left" box="[425,433,1552,1566]" fontSize="6" pageId="3" pageNumber="481">4</subScript>
6:5; lower calypter dirty yellow with white cilia, pilose; halter light yellow. 
<emphasis id="25CBB871FFB9130FFAFEFA34FADAF9FB" bold="true" box="[1282,1338,1533,1559]" pageId="3" pageNumber="481">Legs</emphasis>
(
<figureCitation id="8F8478E6FFB9130FFAB5FA34FA6EF9FB" box="[1353,1422,1533,1559]" captionStart="FIGURE 6" captionStartId="6.[152,255,698,722]" captionTargetBox="[166,1416,199,675]" captionTargetId="figure-19@6.[151,1435,199,675]" captionTargetPageId="6" captionText="FIGURE 6. Thinophilus maritimus sp. nov., holotype, ♂, leg. A. Fore and mid legs; B. Hind tibia and tarsus" figureDoi="http://doi.org/10.5281/zenodo.10202844" httpUri="https://zenodo.org/record/10202844/files/figure.png" pageId="3" pageNumber="481">Fig. 6</figureCitation>
): All coxae metallic green with whitish tomentum; fore and mid coxae with single black bristle, hind coxa lacks lateral setae; fore, mid and hind femora concolorous with coxae. Fore and hind tibiae concolorous with coxae and femora, mid tibia orange yellow. All tarsomeres orange yellow with dark brown bands at tip of each tarsomere, terminal tarsomere of all legs completely darkish brown. 
<emphasis id="25CBB871FFB9130FFD40F944FCF6F94B" bold="true" box="[700,790,1677,1703]" pageId="3" pageNumber="481">Foreleg</emphasis>
(
<figureCitation id="8F8478E6FFB9130FFCDAF944FC63F94B" box="[806,899,1677,1703]" captionStart="FIGURE 6" captionStartId="6.[152,255,698,722]" captionTargetBox="[166,1416,199,675]" captionTargetId="figure-19@6.[151,1435,199,675]" captionTargetPageId="6" captionText="FIGURE 6. Thinophilus maritimus sp. nov., holotype, ♂, leg. A. Fore and mid legs; B. Hind tibia and tarsus" figureDoi="http://doi.org/10.5281/zenodo.10202844" httpUri="https://zenodo.org/record/10202844/files/figure.png" pageId="3" pageNumber="481">Fig. 6A</figureCitation>
): Fore femur basally black and yellow apically, without MSSC, posterolaterally bearing row of black bristles from base to apex, apical 4 bristles larger; ventrally with scattered bristly hairs of various lengths. Fore tibia light brown except for yellowish basal one-third, bearing yellowish hairs all over with black apical bristle both dorsally and ventrally. All tarsomeres yellowish with blackish apex; basitarsus equal to combined length of remaining segments, bearing row of long yellowish bristles along entire length (MSSC); row of setae dorsally and rows of setae ventrally; claws symmetrical; length of femur, tibia and tarsomeres (in mm): 1.17/0.88/0.88/0.23/0.21/0.22/0.21. 
<emphasis id="25CBB871FFB9130FFCB7F8ACFC7FF893" bold="true" box="[843,927,1893,1919]" pageId="3" pageNumber="481">Midleg</emphasis>
(
<figureCitation id="8F8478E6FFB9130FFC4CF8ACFBEDF893" box="[944,1037,1893,1919]" captionStart="FIGURE 6" captionStartId="6.[152,255,698,722]" captionTargetBox="[166,1416,199,675]" captionTargetId="figure-19@6.[151,1435,199,675]" captionTargetPageId="6" captionText="FIGURE 6. Thinophilus maritimus sp. nov., holotype, ♂, leg. A. Fore and mid legs; B. Hind tibia and tarsus" figureDoi="http://doi.org/10.5281/zenodo.10202844" httpUri="https://zenodo.org/record/10202844/files/figure.png" pageId="3" pageNumber="481">Fig. 6A</figureCitation>
): Mid femur concolorous with fore femur, with two ventral rows of black small bristles, 3–4 larger bristles at base and 2–4 bristles at apex. Mid tibia yellow, ventrally bearing row of yellowish hairs along entire length, forming very long hairs at apex. Mid basitarsus with protuberance at apical third bearing row of black spines anteriorly and posteriorly; other tarsomeres with black and yellow hairs; claws same as on foreleg; length of femur, tibia and tarsomeres (in mm): 1.46/1.40/1.04/0.23/0.2 0/0.17/0.27. 
<emphasis id="25CBB871FFBE1308FED8FF73FE63FF38" bold="true" box="[292,387,186,212]" pageId="4" pageNumber="482">Hindleg</emphasis>
(
<figureCitation id="8F8478E6FFBE1308FE6EFF72FE0AFF39" box="[402,490,187,213]" captionStart="FIGURE 6" captionStartId="6.[152,255,698,722]" captionTargetBox="[166,1416,199,675]" captionTargetId="figure-19@6.[151,1435,199,675]" captionTargetPageId="6" captionText="FIGURE 6. Thinophilus maritimus sp. nov., holotype, ♂, leg. A. Fore and mid legs; B. Hind tibia and tarsus" figureDoi="http://doi.org/10.5281/zenodo.10202844" httpUri="https://zenodo.org/record/10202844/files/figure.png" pageId="4" pageNumber="482">Fig. 6B</figureCitation>
): Hind femur twice length of fore and mid femora, curved in middle, without MSSC, bare of bristles. Hind tibia metallic darkish green with white tomentum, moderately swollen with apical process; basitarsus shaped as stout curved process with bifurcation at apex, foot-shaped terminally with long yellowish hairs on outer branch on basal third; other branch bare (MSSC). Tarsomere 2 yellow with row of long yellow hairs ventrally; other 3 tarsomeres together equal in length to tarsomere 2; claws same as on foreleg; length of femur, tibia and tarsomeres (in mm): 2.71/1.54/0.69/0.73/0.26/0.27/0.31. 
<emphasis id="25CBB871FFBE1308FCC2FEA7FC53FE64" bold="true" box="[830,947,366,392]" pageId="4" pageNumber="482">Abdomen</emphasis>
(
<figureCitation id="8F8478E6FFBE1308FC3DFEA6FBE5FE65" box="[961,1029,367,393]" captionStart="FIGURE 7" captionStartId="6.[152,255,1294,1318]" captionTargetBox="[151,1435,783,1271]" captionTargetId="figure-44@6.[151,1435,783,1271]" captionTargetPageId="6" captionText="FIGURE 7. Thinophilus maritimus sp. nov., holotype, ♂, abdomen. A. Ventral view; B. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10202846" httpUri="https://zenodo.org/record/10202846/files/figure.png" pageId="4" pageNumber="482">Fig. 7</figureCitation>
): First and second segments dorsally dark green with silvery and black pubescence, remaining segments darker with whitish pubescence; ventrally dull yellowish with white pubescence. 
<emphasis id="25CBB871FFBE1308FDEFFE7EFD5DFE3C" bold="true" box="[531,701,438,465]" pageId="4" pageNumber="482">Male genitalia</emphasis>
(
<figureCitation id="8F8478E6FFBE1308FD36FE7EFCD0FE3D" box="[714,816,439,465]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="6.[152,255,1878,1902]" captionStartId-1="7.[152,255,746,770]" captionTargetBox-0="[157,1420,1379,1850]" captionTargetBox-1="[151,1435,181,722]" captionTargetId-0="figure-65@6.[151,1435,1379,1855]" captionTargetId-1="figure-19@7.[151,1435,181,722]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 8. Thinophilus maritimus sp. nov., holotype, ♂, male terminalia A. Lateral view; B. Dorsal view.Abbreviations: Cerc: Cerci, Ph: Phallus, Sur: Surstylus." captionText-1="FIGURE 9. Thinophilus maritimus sp. nov., holotype, ♂, male terminalia A. Lateral view of phallus and phallosome; B. Dorsal view of the tip of surstylus; C. Dorsal view of cercus." figureDoi-0="http://doi.org/10.5281/zenodo.10202848" figureDoi-1="http://doi.org/10.5281/zenodo.10202850" httpUri-0="https://zenodo.org/record/10202848/files/figure.png" httpUri-1="https://zenodo.org/record/10202850/files/figure.png" pageId="4" pageNumber="482">Figs 8–9</figureCitation>
): Dark brown pair of surstylus with furcated and pincershaped tips; cerci not fused, pale whitish brown with small pale white hairs throughout and larger hairs along edges; phallus strap-shaped, strongly curved, not coiled. 
<emphasis id="25CBB871FFBE1308FD3BFE36FCFEFDF5" bold="true" box="[711,798,511,537]" pageId="4" pageNumber="482">Female</emphasis>
(
<figureCitation id="8F8478E6FFBE1308FCD2FE36FC63FDF5" box="[814,899,511,537]" captionStart="FIGURE 10" captionStartId="7.[152,255,1917,1941]" captionTargetId="figure-55@7.[151,1435,867,1894]" captionTargetPageId="7" captionText="FIGURE 10. Thinophilus maritimus sp. nov., paratype, lateral habitus of female." figureDoi="http://doi.org/10.5281/zenodo.10277925" httpUri="https://zenodo.org/record/10277925/files/figure.png" pageId="4" pageNumber="482">Fig. 10</figureCitation>
): Body length 
<quantity id="D047C986FFBE1308FBD5FE36FB50FDF5" box="[1065,1200,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="4" pageNumber="482" unit="mm" value="6.65" valueMax="6.7" valueMin="6.6">6.6–6.7 mm</quantity>
Wing length 
<quantity id="D047C986FFBE1308FAB4FE37FF26FDD1" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.2" metricValueMin="5.1" pageId="4" pageNumber="482" unit="mm" value="5.15" valueMax="5.2" valueMin="5.1">5.1–5.2 mm</quantity>
. Similar to male except lacking secondary sexual characters, otherwise as follows: All coxae and trochanters black with yellowish-white tomentum; fore femur black, with yellowish apex; mid femur black on basal half, remaining yellowish; hind femur more than 1.5 times longer than preceding femora, distinctly curved, black except yellowish apex; all femora covered with yellowish and black pubescence; all tibia yellow except black infuscation at apex of hind tibia, with hind tibia slightly longer than other tibiae; all tibia covered with black and pale yellow pubescence; all tarsomeres yellow with black apex, with apical 2 tarsomeres blacker; 6 acanthophorite setae present on each hemitergite 9+10; female genitalia as in figure (
<figureCitation id="8F8478E6FFBE1308FCF7FD32FCBCFCF9" box="[779,860,763,789]" captionStart="FIGURE 11" captionStartId="8.[152,255,1337,1361]" captionTargetBox="[165,1431,186,1311]" captionTargetId="figure-19@8.[151,1436,181,1313]" captionTargetPageId="8" captionText="FIGURE 11. Thinophilus maritimus sp. nov., paratype, female terminalia." figureDoi="http://doi.org/10.5281/zenodo.10202852" httpUri="https://zenodo.org/record/10202852/files/figure.png" pageId="4" pageNumber="482">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="43C034EBFFBE1308FF64FB4AFEB5FB52" ID-DOI="http://doi.org/10.5281/zenodo.10202838" ID-Zenodo-Dep="10202838" httpUri="https://zenodo.org/record/10202838/files/figure.png" pageId="4" pageNumber="482" startId="4.[152,255,1155,1179]" targetBox="[151,1435,813,1131]" targetPageId="4" targetType="figure">
<paragraph id="17006463FFBE1308FF64FB4AFEB5FB52" blockId="4.[151,1436,1155,1215]" pageId="4" pageNumber="482">
<emphasis id="25CBB871FFBE1308FF64FB4AFEFAFB77" bold="true" box="[152,282,1155,1179]" pageId="4" pageNumber="482">FIGURE 3.</emphasis>
<taxonomicName id="D0BF1FE0FFBE1308FEDEFB4AFDEAFB77" authority="Mukherjee &amp; Pramanik &amp; Naskar &amp; Banerjee, 2023" authorityName="Mukherjee &amp; Pramanik &amp; Naskar &amp; Banerjee" authorityYear="2023" box="[290,522,1155,1179]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="25CBB871FFBE1308FEDEFB4AFDEAFB77" box="[290,522,1155,1179]" italics="true" pageId="4" pageNumber="482">Thinophilus maritimus</emphasis>
</taxonomicName>
<emphasis id="25CBB871FFBE1308FDEDFB4AFD83FB77" bold="true" box="[529,611,1155,1179]" pageId="4" pageNumber="482">
<taxonomicNameLabel id="3EF8050AFFBE1308FDEDFB4AFD83FB77" box="[529,611,1155,1179]" pageId="4" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ♂, head 
<emphasis id="25CBB871FFBE1308FCCEFB4AFCAAFB77" bold="true" box="[818,842,1155,1179]" pageId="4" pageNumber="482">A.</emphasis>
Frontal view; 
<emphasis id="25CBB871FFBE1308FC1FFB4AFC1AFB77" bold="true" box="[995,1018,1155,1179]" pageId="4" pageNumber="482">B.</emphasis>
Lateral view; 
<emphasis id="25CBB871FFBE1308FB6EFB4AFB4AFB77" bold="true" box="[1170,1194,1155,1179]" pageId="4" pageNumber="482">C.</emphasis>
Magnified lateral view showing antennae.
</paragraph>
</caption>
<subSubSection id="5FA537E8FFBE1308FF3BFB24FCD3F8AB" pageId="4" pageNumber="482" type="etymology">
<paragraph id="17006463FFBE1308FF3BFB24FEECFAC7" blockId="4.[151,1437,1261,2044]" pageId="4" pageNumber="482">
<emphasis id="25CBB871FFBE1308FF3BFB24FEAEFAEB" bold="true" box="[199,334,1261,1287]" pageId="4" pageNumber="482">Etymology.</emphasis>
From the Latin 
<taxonomicName id="D0BF1FE0FFBE1308FDFDFB24FD9CFAEB" authorityName="Mukherjee &amp; Pramanik &amp; Naskar &amp; Banerjee" authorityYear="2023" box="[513,636,1261,1287]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="species" species="maritimus">
‘ 
<emphasis id="25CBB871FFBE1308FDF6FB27FD9CFAEB" box="[522,636,1262,1287]" italics="true" pageId="4" pageNumber="482">maritimus</emphasis>
</taxonomicName>
’ (of the sea), as the specimens were collected along the coast of the Bay of Bengal.
</paragraph>
<paragraph id="17006463FFBE1308FF3BFAFCFEEDF9CB" blockId="4.[151,1437,1261,2044]" pageId="4" pageNumber="482">
<emphasis id="25CBB871FFBE1308FF3BFAFCFEA8FAA3" bold="true" box="[199,328,1333,1359]" pageId="4" pageNumber="482">Behaviour.</emphasis>
The specimens were seen in clusters or groups in areas where the sands were moist and along the edges of seawater (
<figureCitation id="8F8478E6FFBE1308FE91FA90FE20FA9F" box="[365,448,1369,1395]" captionStart="FIGURE 12" captionStartId="9.[152,255,1170,1194]" captionTargetBox="[151,1436,181,1146]" captionTargetId="figure-19@9.[151,1436,181,1146]" captionTargetPageId="9" captionText="FIGURE 12. Two male individuals of Thinophilus maritimus sp. nov. on moist sandy beach." figureDoi="http://doi.org/10.5281/zenodo.10202854" httpUri="https://zenodo.org/record/10202854/files/figure.png" pageId="4" pageNumber="482">Fig. 12</figureCitation>
). They were not seen on dry sands. During the survey, the females were relatively easier to catch than males, mostly because as soon as the males were approached closer than approximately 
<quantity id="D047C986FFBE1308FAFFFAB4FA81FA74" box="[1283,1377,1405,1432]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="482" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 m</quantity>
, they mostly jumped on the sea waves, kept floating on the waves, perfectly balancing themselves even on larger waves, and returned back to the shore only when the said distance was maintained again. Females were not seen showing this behaviour. Therefore, we hypothesize that the modified hind limbs of males might have a role in the floating behaviour.
</paragraph>
<paragraph id="17006463FFBE1308FF3BF9F8FCD3F8AB" blockId="4.[151,1437,1261,2044]" pageId="4" pageNumber="482">
<emphasis id="25CBB871FFBE1308FF3BF9F8FEAAF9A7" bold="true" box="[199,330,1585,1611]" pageId="4" pageNumber="482">Barcoding.</emphasis>
DNA barcoding of 
<specimenCount id="01B9AFEAFFBE1308FDC7F9FBFD45F9A7" box="[571,677,1585,1612]" count="1" pageId="4" pageNumber="482" type="male">one male</specimenCount>
(Accession no: ON255696) and female 
<typeStatus id="C804DAC1FFBE1308FB79F9FBFB05F9A0" box="[1157,1253,1586,1612]" pageId="4" pageNumber="482" type="paratype">paratype</typeStatus>
(Accession no: ON255707) of our species was obtained, and both specimens displayed 100% similarity, confirming that the sexes are correctly associated. The closest and most similar sequence for both the specimens are identified as “ 
<taxonomicName id="D0BF1FE0FFBE1308FB14F9B0FA7CF97F" box="[1256,1436,1657,1683]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="family">
<emphasis id="25CBB871FFBE1308FB14F9B0FA7CF97F" box="[1256,1436,1657,1683]" italics="true" pageId="4" pageNumber="482">Dolichopodidae</emphasis>
</taxonomicName>
sp” sharing only 87% similarity in BOLD. In the constructed phylogenetic tree (
<figureCitation id="8F8478E6FFBE1308FBCCF954FB65F95B" box="[1072,1157,1693,1719]" captionStart="FIGURE 13" captionStartId="9.[152,255,1903,1927]" captionTargetBox="[155,1432,1223,1875]" captionTargetId="figure-38@9.[151,1435,1218,1879]" captionTargetPageId="9" captionText="FIGURE 13. Phylogenetic tree where our specimen clusters with individuals of the genus Thinophilus where the other 3 genera, Scellus, Hydrophorus and Thambemyia show definite distinction from our specimen. Despite the clustering of our specimen with that of the genus Thinophilus, their distinction within the group is also evident. Numbers next to each branch of the tree indicate the percentage of replicate trees where associated taxa clustered together in the bootstrap." figureDoi="http://doi.org/10.5281/zenodo.10202856" httpUri="https://zenodo.org/record/10202856/files/figure.png" pageId="4" pageNumber="482">Fig. 13</figureCitation>
), 
<taxonomicName id="D0BF1FE0FFBE1308FB60F954FA7DF95B" authority="Mukherjee &amp; Pramanik &amp; Naskar &amp; Banerjee, 2023" authorityName="Mukherjee &amp; Pramanik &amp; Naskar &amp; Banerjee" authorityYear="2023" box="[1180,1437,1693,1719]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="25CBB871FFBE1308FB60F954FA7DF95B" box="[1180,1437,1693,1719]" italics="true" pageId="4" pageNumber="482">Thinophilus maritimus</emphasis>
</taxonomicName>
<emphasis id="25CBB871FFBE1308FF6BF908FF10F937" bold="true" box="[151,240,1729,1755]" pageId="4" pageNumber="482">
<taxonomicNameLabel id="3EF8050AFFBE1308FF6BF908FF10F937" box="[151,240,1729,1755]" pageId="4" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
clusters within the genus 
<taxonomicName id="D0BF1FE0FFBE1308FDEFF908FD79F937" authorityName="Wahlberg" authorityYear="1844" box="[531,665,1729,1755]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="25CBB871FFBE1308FDEFF908FD79F937" box="[531,665,1729,1755]" italics="true" pageId="4" pageNumber="482">Thinophilus</emphasis>
</taxonomicName>
where the other three genera, 
<taxonomicName id="D0BF1FE0FFBE1308FC16F908FB61F930" authority="Loew" authorityName="Loew" authorityYear="1857" box="[1002,1153,1729,1756]" class="Insecta" family="Dolichopodidae" genus="Scellus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="25CBB871FFBE1308FC16F908FBD9F937" box="[1002,1081,1729,1755]" italics="true" pageId="4" pageNumber="482">Scellus</emphasis>
Loew
</taxonomicName>
, 
<taxonomicName id="D0BF1FE0FFBE1308FB70F908FA8DF937" authority="Fallen" authorityName="Fallen" authorityYear="1823" box="[1164,1389,1729,1755]" class="Insecta" family="Dolichopodidae" genus="Hydrophorus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="25CBB871FFBE1308FB70F908FAC0F937" box="[1164,1312,1729,1755]" italics="true" pageId="4" pageNumber="482">Hydrophorus</emphasis>
Fallén
</taxonomicName>
and 
<taxonomicName id="D0BF1FE0FFBE1308FF6BF92CFE6CF913" authority="Oldroyd" authorityName="Oldroyd" authorityYear="1956" box="[151,396,1765,1791]" class="Insecta" family="Dolichopodidae" genus="Thambemyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="25CBB871FFBE1308FF6BF92CFEC8F913" box="[151,296,1765,1791]" italics="true" pageId="4" pageNumber="482">Thambemyia</emphasis>
Oldroyd
</taxonomicName>
show definite distinction from our specimen. In spite of the clustering of our specimen with that of the genus 
<taxonomicName id="D0BF1FE0FFBE1308FE9CF8C0FE06F8CF" authorityName="Wahlberg" authorityYear="1844" box="[352,486,1801,1827]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="25CBB871FFBE1308FE9CF8C0FE06F8CF" box="[352,486,1801,1827]" italics="true" pageId="4" pageNumber="482">Thinophilus</emphasis>
</taxonomicName>
, their distinction with sequences of available 
<taxonomicName id="D0BF1FE0FFBE1308FC0AF8C0FB36F8CF" box="[1014,1238,1801,1827]" pageId="4" pageNumber="482">
<emphasis id="25CBB871FFBE1308FC0AF8C0FB9CF8CF" box="[1014,1148,1801,1827]" italics="true" pageId="4" pageNumber="482">Thinophilus</emphasis>
species
</taxonomicName>
in BOLD is also evident, thereby confirming our specimen as a new species.
</paragraph>
</subSubSection>
<subSubSection id="5FA537E8FFBE1308FF3BF898FDA4F887" box="[199,580,1873,1900]" pageId="4" pageNumber="482" type="distribution">
<paragraph id="17006463FFBE1308FF3BF898FDA4F887" blockId="4.[151,1437,1261,2044]" box="[199,580,1873,1900]" pageId="4" pageNumber="482">
<emphasis id="25CBB871FFBE1308FF3BF898FE81F887" bold="true" box="[199,353,1873,1899]" pageId="4" pageNumber="482">Distribution.</emphasis>
<collectingRegion id="D57BAA81FFBE1308FE9BF89BFE17F887" box="[359,503,1873,1900]" country="India" name="West Bengal" pageId="4" pageNumber="482">West Bengal</collectingRegion>
, 
<collectingCountry id="6FA824F3FFBE1308FDFFF898FDA0F887" box="[515,576,1873,1899]" name="India" pageId="4" pageNumber="482">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="5FA537E8FFBE1309FF3BF8BCFE11FEF1" lastPageId="5" lastPageNumber="483" pageId="4" pageNumber="482" type="discussion">
<paragraph id="17006463FFBE1309FF3BF8BCFE11FEF1" blockId="4.[151,1437,1261,2044]" lastBlockId="5.[151,1437,151,285]" lastPageId="5" lastPageNumber="483" pageId="4" pageNumber="482">
<emphasis id="25CBB871FFBE1308FF3BF8BCFEDBF863" bold="true" box="[199,315,1909,1935]" pageId="4" pageNumber="482">Remarks.</emphasis>
<taxonomicName id="D0BF1FE0FFBE1308FEB9F8BCFDA7F863" authority="Mukherjee &amp; Pramanik &amp; Naskar &amp; Banerjee, 2023" authorityName="Mukherjee &amp; Pramanik &amp; Naskar &amp; Banerjee" authorityYear="2023" box="[325,583,1909,1935]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="25CBB871FFBE1308FEB9F8BCFDA7F863" box="[325,583,1909,1935]" italics="true" pageId="4" pageNumber="482">Thinophilus maritimus</emphasis>
</taxonomicName>
<emphasis id="25CBB871FFBE1308FDADF8BCFD4EF863" bold="true" box="[593,686,1909,1935]" pageId="4" pageNumber="482">
<taxonomicNameLabel id="3EF8050AFFBE1308FDADF8BCFD4EF863" box="[593,686,1909,1935]" pageId="4" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected from the sandy delta beaches of Sagar Island and Bakkhali along the coast of the Bay of Bengal of South 24 Parganas district of 
<collectingRegion id="D57BAA81FFBE1308FBFDF853FB71F85F" box="[1025,1169,1945,1972]" country="India" name="West Bengal" pageId="4" pageNumber="482">West Bengal</collectingRegion>
, 
<collectingCountry id="6FA824F3FFBE1308FB60F850FB39F85F" box="[1180,1241,1945,1971]" name="India" pageId="4" pageNumber="482">India</collectingCountry>
. According to the key to New World genera (
<bibRefCitation id="732E1992FFBE1308FE36F874FDB9F83B" author="Bickel, D. J." box="[458,601,1981,2007]" pageId="4" pageNumber="482" refId="ref6324" refString="Bickel, D. J. (2009) Dolichopodidae (long-legged flies). In: Bickel, D. J., Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, N. E. &amp; Zumbado, M. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, Ontario, Canada, pp. 671 - 694." type="book" year="2009">Bickel 2009</bibRefCitation>
), the species belongs to the subfamily 
<taxonomicName id="D0BF1FE0FFBE1308FBECF874FB5AF83B" box="[1040,1210,1981,2007]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="subFamily" subFamily="Hydrophorinae">Hydrophorinae</taxonomicName>
and ends at couplet 55 with the genus 
<taxonomicName id="D0BF1FE0FFBE1308FE8DF828FE17F817" authorityName="Wahlberg" authorityYear="1844" box="[369,503,2017,2043]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="25CBB871FFBE1308FE8DF828FE17F817" box="[369,503,2017,2043]" italics="true" pageId="4" pageNumber="482">Thinophilus</emphasis>
</taxonomicName>
, but it completely lacks lateral setae on the hind coxa. However, we assign this species to 
<taxonomicName id="D0BF1FE0FFBF1309FEF0FF5EFE72FF5D" authorityName="Wahlberg" authorityYear="1844" box="[268,402,151,177]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="25CBB871FFBF1309FEF0FF5EFE72FF5D" box="[268,402,151,177]" italics="true" pageId="5" pageNumber="483">Thinophilus</emphasis>
</taxonomicName>
due to several morphological similarities, and its clustering within the 
<taxonomicName id="D0BF1FE0FFBF1309FB4DFF5EFAD7FF5D" authorityName="Wahlberg" authorityYear="1844" box="[1201,1335,151,177]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="25CBB871FFBF1309FB4DFF5EFAD7FF5D" box="[1201,1335,151,177]" italics="true" pageId="5" pageNumber="483">Thinophilus</emphasis>
</taxonomicName>
group in the phylogenetic tree. Some other species such as 
<taxonomicName id="D0BF1FE0FFBF1309FD2CFF72FB13FF39" authority="Samoh, Satasook &amp; Grootaert, 2017" authorityName="Samoh, Satasook &amp; Grootaert" authorityYear="2017" box="[720,1267,187,213]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="483" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="25CBB871FFBF1309FD2CFF72FCAEFF39" box="[720,846,187,213]" italics="true" pageId="5" pageNumber="483">T. parvulus</emphasis>
<bibRefCitation id="732E1992FFBF1309FCAAFF72FB13FF39" author="Samoh, A. &amp; Satasook, C. &amp; Grootaert, P." box="[854,1267,187,213]" pageId="5" pageNumber="483" pagination="1 - 40" refId="ref7450" refString="Samoh, A., Satasook, C. &amp; Grootaert, P. (2017) Eight new species of marine dolichopodid flies of Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae) from peninsular Thailand. European Journal of Taxonomy, 329, 1 - 40. https: // doi. org / 10.5852 / ejt. 2017.329" type="journal article" year="2017">Samoh, Satasook &amp; Grootaert, 2017</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="D0BF1FE0FFBF1309FAFCFF72FDC9FF15" authority="Samoh, Satasook et Grootaert, 2017" authorityName="Samoh, Satasook et Grootaert" authorityYear="2017" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="483" phylum="Arthropoda" rank="species" species="spinatoides">
<emphasis id="25CBB871FFBF1309FAFCFF72FA7DFF39" box="[1280,1437,187,213]" italics="true" pageId="5" pageNumber="483">T. spinatoides</emphasis>
<bibRefCitation id="732E1992FFBF1309FF6BFF16FDC9FF15" author="Samoh, A. &amp; Satasook, C. &amp; Grootaert, P." box="[151,553,223,249]" pageId="5" pageNumber="483" pagination="1 - 40" refId="ref7450" refString="Samoh, A., Satasook, C. &amp; Grootaert, P. (2017) Eight new species of marine dolichopodid flies of Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae) from peninsular Thailand. European Journal of Taxonomy, 329, 1 - 40. https: // doi. org / 10.5852 / ejt. 2017.329" type="journal article" year="2017">Samoh, Satasook et Grootaert, 2017</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="D0BF1FE0FFBF1309FDC8FF16FC74FF15" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[564,916,223,249]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="483" phylum="Arthropoda" rank="species" species="subpalpatus">
<emphasis id="25CBB871FFBF1309FDC8FF16FD35FF15" box="[564,725,223,249]" italics="true" pageId="5" pageNumber="483">T. subpalpatus</emphasis>
Grichanov, 2023
</taxonomicName>
also lack lateral setae on the hind coxa (
<bibRefCitation id="732E1992FFBF1309FAB1FF16FEF1FEF1" author="Samoh, A. &amp; Satasook, C. &amp; Grootaert, P." pageId="5" pageNumber="483" pagination="1 - 40" refId="ref7450" refString="Samoh, A., Satasook, C. &amp; Grootaert, P. (2017) Eight new species of marine dolichopodid flies of Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae) from peninsular Thailand. European Journal of Taxonomy, 329, 1 - 40. https: // doi. org / 10.5852 / ejt. 2017.329" type="journal article" year="2017">
Samoh 
<emphasis id="25CBB871FFBF1309FF6BFECAFF28FEF1" box="[151,200,259,285]" italics="true" pageId="5" pageNumber="483">et al</emphasis>
. 2017
</bibRefCitation>
; 
<bibRefCitation id="732E1992FFBF1309FEE1FECAFE05FEF1" author="Grichanov, I. Y." box="[285,485,259,285]" pageId="5" pageNumber="483" pagination="1 - 17" refId="ref6574" refString="Grichanov, I. Y. (2023 b) Four new species of the genus Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae) with new records and a key to species known from India and adjacent regions. Far Eastern Entomologist, 472, 1 - 17. https: // doi. org / 10.25221 / fee. 472.1" type="journal article" year="2023">Grichanov 2023b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>