165 lines
22 KiB
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165 lines
22 KiB
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<document id="540E93655F6A1E5179F1E0785857406A" ID-DOI="10.11646/zootaxa.4755.1.7" ID-GBIF-Dataset="01f9a310-e503-4558-945b-dce61bdc2dbf" ID-ISSN="1175-5326" ID-Zenodo-Dep="3724311" ID-ZooBank="FFDB122C-E34B-486A-AC99-88B240AAC4DD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584968616145" checkinUser="plazi" docAuthor="Lehnert, Helmut & Stone, Robert P." docDate="2020" docId="0E138793FF94AF21FF2DF6E10F3DA44B" docLanguage="en" docName="zootaxa.4755.1.7.pdf" docOrigin="Zootaxa 4755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hymedesmiidae Topsent 1928" docType="treatment" docVersion="5" lastPageNumber="157" masterDocId="F22AFFEBFF96AF23FFBAF6000D3CA11B" masterDocTitle="Acantorna-a new genus of Hymedesmiidae (Porifera, Demospongiae Poecilosclerida) from the western Aleutian Islands of Alaska with description of Acantorna tahoma n. sp." masterLastPageNumber="162" masterPageNumber="155" pageNumber="157" updateTime="1698730803449" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="63DF84CB977C80EFB8C7ED0C63E49AA8">Acantorna-a new genus of Hymedesmiidae (Porifera, Demospongiae Poecilosclerida) from the western Aleutian Islands of Alaska with description of Acantorna tahoma n. sp.</mods:title>
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<mods:namePart id="19C47FA0EFB0D5E9BD5187C03B5EF3F8">Lehnert, Helmut</mods:namePart>
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<mods:namePart id="043241D83D7CA3D4964D07E224666F26">Stone, Robert P.</mods:namePart>
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<mods:title id="04E77438C15E7C1106FAB798D7D1C2DA">Zootaxa</mods:title>
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<mods:part id="5B6F46CE365FF9D4BD11EE6130CEC826">
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<mods:date id="4A981546DB42F97D5674AB8CA1645B1B">2020</mods:date>
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<mods:number id="1E4E5C7A8B555EC2811C462273BEAAAA">2020-03-23</mods:number>
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<mods:number id="B2C7F047F9F07B21A337216602C32A77">4755</mods:number>
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<treatment id="0E138793FF94AF21FF2DF6E10F3DA44B" ID-DOI="http://doi.org/10.5281/zenodo.3728370" ID-GBIF-Taxon="162696947" ID-Zenodo-Dep="3728370" LSID="urn:lsid:plazi:treatment:0E138793FF94AF21FF2DF6E10F3DA44B" httpUri="http://treatment.plazi.org/id/0E138793FF94AF21FF2DF6E10F3DA44B" lastPageNumber="157" pageId="2" pageNumber="157">
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<subSubSection id="CEA0650EFF94AF21FF2DF6E10804A0ED" pageId="2" pageNumber="157" type="nomenclature">
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<paragraph id="86053685FF94AF21FF2DF6E10F4EA1E0" blockId="2.[151,626,153,252]" box="[151,626,225,252]" pageId="2" pageNumber="157">
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<heading id="DD4D81E9FF94AF21FF2DF6E10F4EA1E0" bold="true" box="[151,626,225,252]" fontSize="11" level="1" pageId="2" pageNumber="157" reason="1">
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<emphasis id="B4CEEA97FF94AF21FF2DF6E10F4EA1E0" bold="true" box="[151,626,225,252]" pageId="2" pageNumber="157">
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Family
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<taxonomicName id="41BA4D06FF94AF21FF4DF6E10F4EA1E0" ID-CoL="84K9R" authority="Topsent, 1928" authorityName="Topsent" authorityYear="1928" box="[247,626,225,252]" class="Demospongiae" family="Hymedesmiidae" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="family">
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Hymedesmiidae
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<bibRefCitation id="E22B4B74FF94AF21FE7CF6E20F4EA1E0" author="Topsent, E." box="[454,626,225,252]" pageId="2" pageNumber="157" pagination="1 - 376" refId="ref4797" refString="Topsent, E. (1928) Spongiaires de l'Atlantique et de la Mediterranee provenant des croisieres du Prince Albert ler de Monaco. Resultats des campagnes scientifiques accomplies par le Prince Albert I, Monaco, 74, 1 - 376, pls. I-XI." type="journal article" year="1928">Topsent, 1928</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="86053685FF94AF21FF2DF7290804A0ED" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
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<emphasis id="B4CEEA97FF94AF21FF2DF7290C24A05F" bold="true" box="[151,280,297,324]" pageId="2" pageNumber="157">Definition:</emphasis>
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(amended from Van Soest 2002:575):
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<taxonomicName id="41BA4D06FF94AF21FD7FF7290E4CA058" authorityName="Topsent" authorityYear="1928" box="[709,880,297,323]" class="Demospongiae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="order">Poecilosclerida</taxonomicName>
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with smooth or acanthose ectosomal tornotes (basically diactinal, oxeote, but may be anisotornotes or styles), a peripheral choanosomal plumose skeleton of bundles or tracts of smooth or acanthose tornotes, a basal skeleton of acanthostyles, singly or in plumose bundles (may be absent), echinating acanthostyles (may be absent) or, a choanosmal confused arrangement of all spicule categories. Additional non-echinating acanthostyles or acanthoxeas may occasionally be present. Microscleres include arcuate chelas and sigmas, but may be absent. Ectosomal surfaces often but, not always with areolated porefields.
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</paragraph>
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</subSubSection>
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<subSubSection id="CEA0650EFF94AF21FF7DF7FF0F3DA44B" pageId="2" pageNumber="157" type="discussion">
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<paragraph id="86053685FF94AF21FF7DF7FF0F62A3EB" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
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<emphasis id="B4CEEA97FF94AF21FF7DF7FF0C73A301" bold="true" box="[199,335,511,538]" pageId="2" pageNumber="157">Discussion:</emphasis>
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To accommodate the new genus in
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<taxonomicName id="41BA4D06FF94AF21FD6DF4000EB1A301" authorityName="Topsent" authorityYear="1928" box="[727,909,512,538]" class="Demospongiae" family="Hymedesmiidae" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="family">Hymedesmiidae</taxonomicName>
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the definition (Van Soest 2002, p. 575) requires expanding to allow for the possession of acanthose ectosomal tornotes (other hymedesmiids have only smooth tornotes) and a confusedly plumose choanosomal arrangement of spicules which include the choanosomal tornotes. In addition, the ectosome is often but not always with areolated pore-fields. At present the family
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<taxonomicName id="41BA4D06FF94AF21FB5CF46B08A0A39E" authorityName="Topsent" authorityYear="1928" box="[1254,1436,619,645]" class="Demospongiae" family="Hymedesmiidae" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="family">Hymedesmiidae</taxonomicName>
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contains 10 genera, none with acanthose tornotes or with a more confusedly plumose choanosomal arrangement of spicules. We present additional differences between
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<taxonomicName id="41BA4D06FF94AF21FD5CF4B30E60A3D6" authority="Lehnert & Stone, 2020" authorityName="Lehnert & Stone" authorityYear="2020" box="[742,860,691,717]" class="Demospongiae" family="Hymedesmiidae" genus="Acantorna" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus" status="gen. nov.">
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<emphasis id="B4CEEA97FF94AF21FD5CF4B30E60A3D6" box="[742,860,691,717]" italics="true" pageId="2" pageNumber="157">Acantorna</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AFFD57ECFF94AF21FCDFF4B30E8EA3D6" box="[869,946,691,717]" pageId="2" pageNumber="157" rank="genus">n. gen.</taxonomicNameLabel>
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and the existing genera of
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<taxonomicName id="41BA4D06FF94AF21FB5CF4B208A0A3D7" authorityName="Topsent" authorityYear="1928" box="[1254,1436,690,716]" class="Demospongiae" family="Hymedesmiidae" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="family">Hymedesmiidae</taxonomicName>
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(according to Van Soest 2002: 576-593):
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</paragraph>
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<paragraph id="86053685FF94AF21FF7DF4FB0DC5A223" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
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<emphasis id="B4CEEA97FF94AF21FF7DF4FB0DECA20F" box="[199,208,763,788]" italics="true" pageId="2" pageNumber="157">-</emphasis>
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<taxonomicName id="41BA4D06FF94AF21FF60F4FA0F1FA20F" authority="Topsent, 1927" authorityName="Topsent" authorityYear="1927" box="[218,547,762,788]" class="Demospongiae" family="Hymedesmiidae" genus="Acanthancora" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
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<emphasis id="B4CEEA97FF94AF21FF60F4FA0C45A20F" box="[218,377,762,788]" italics="true" pageId="2" pageNumber="157">Acanthancora</emphasis>
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<bibRefCitation id="E22B4B74FF94AF21FE38F4FA0F1FA20F" author="Topsent, E." box="[386,547,762,788]" pageId="2" pageNumber="157" pagination="1 - 19" refId="ref4766" refString="Topsent, E. (1927) Diagnoses d'Eponges nouvelles recueillies par le Prince Albert ler de Monaco. Bulletin de l'Institut oceanographique Monaco, 502, 1 - 19." type="journal article" year="1927">Topsent, 1927</bibRefCitation>
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</taxonomicName>
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has modified spined tuberculate chelae and a hymedesmioid arrangement of spicules.
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</paragraph>
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<paragraph id="86053685FF94AF21FF7DF542085EA240" blockId="2.[151,1437,297,1360]" box="[199,1378,833,860]" pageId="2" pageNumber="157">
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-
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<taxonomicName id="41BA4D06FF94AF21FF62F5420CF0A240" authority="Gray, 1867" authorityName="Gray" authorityYear="1867" box="[216,460,833,859]" class="Demospongiae" family="Hymedesmiidae" genus="Hamigera" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
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<emphasis id="B4CEEA97FF94AF21FF62F5420C76A240" box="[216,330,834,859]" italics="true" pageId="2" pageNumber="157">Hamigera</emphasis>
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<bibRefCitation id="E22B4B74FF94AF21FEEAF5410CF0A240" author="Gray, J. E." box="[336,460,833,859]" pageId="2" pageNumber="157" pagination="492 - 558" refId="ref4073" refString="Gray, J. E. (1867) Notes on the Arrangement of Sponges, with the Descriptions of some New Genera. Proceedings of the Zoological Society of London, 1867 (2), 492 - 558, pls. XXVII-XXVIII. [https: // biodiversitylibrary. org / page / 29533932]" type="journal article" year="1867">Gray, 1867</bibRefCitation>
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</taxonomicName>
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has strongyles ot tylotes as tornotes, choanosomal smooth styles or subtylostyles.
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</paragraph>
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<paragraph id="86053685FF94AF21FF7DF5660FFDA2B8" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
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<emphasis id="B4CEEA97FF94AF21FF7DF5660DECA264" box="[199,208,870,895]" italics="true" pageId="2" pageNumber="157">-</emphasis>
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<taxonomicName id="41BA4D06FF94AF21FF60F5650F39A264" authority="Burton, 1934" authorityName="Burton" authorityYear="1934" box="[218,517,869,896]" class="Demospongiae" family="Hymedesmiidae" genus="Hemimycale" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
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<emphasis id="B4CEEA97FF94AF21FF60F5650C5AA264" box="[218,358,869,895]" italics="true" pageId="2" pageNumber="157">Hemimycale</emphasis>
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<bibRefCitation id="E22B4B74FF94AF21FED5F5660F39A264" author="Burton, M." box="[367,517,869,896]" pageId="2" pageNumber="157" pagination="513 - 621" refId="ref3752" refString="Burton, M. (1934) Sponges. Scientific Reports of the Great Barrier Reef Expedition 1928 - 29, 4 (14), 513 - 621, pls. 1 - 2. [https: // www. biodiversitylibrary. org / page / 49517195 # page / 667 / mode / 1 up]" type="journal article" year="1934">Burton, 1934</bibRefCitation>
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</taxonomicName>
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has no acanthostyles and no microscleres. Smooth strongyles and styles are not separated in ectosomal and choanosomal regions.
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</paragraph>
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<paragraph id="86053685FF94AF21FF7DF5AD0854A2DC" blockId="2.[151,1437,297,1360]" box="[199,1384,940,967]" pageId="2" pageNumber="157">
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<emphasis id="B4CEEA97FF94AF21FF7DF5AD0DECA2DD" box="[199,208,941,966]" italics="true" pageId="2" pageNumber="157">-</emphasis>
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<taxonomicName id="41BA4D06FF94AF21FF6DF5AD0F0AA2DD" authority="Bowerbank, 1864" authorityName="Bowerbank" authorityYear="1864" box="[215,566,940,967]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
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<emphasis id="B4CEEA97FF94AF21FF6DF5AD0C54A2DC" box="[215,360,941,967]" italics="true" pageId="2" pageNumber="157">Hymedesmia</emphasis>
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<bibRefCitation id="E22B4B74FF94AF21FED5F5AC0F0AA2DD" author="Bowerbank, J. S." box="[367,566,940,966]" pageId="2" pageNumber="157" refId="ref3640" refString="Bowerbank, J. S. (1864) A Monograph of the British Spongiadae. Vol. 1. Ray Society, London, xx + 290 pp., XXXVII pls. https: // doi. org / 10.5962 / bhl. title. 2038" type="book" year="1864">Bowerbank, 1864</bibRefCitation>
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</taxonomicName>
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are all thinly encrusting and have a hymedesmioid skeletal arrangement.
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</paragraph>
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<paragraph id="86053685FF94AF21FF7DF5D10F2CA515" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
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<emphasis id="B4CEEA97FF94AF21FF7DF5D10DECA2F1" box="[199,208,977,1002]" italics="true" pageId="2" pageNumber="157">-</emphasis>
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<taxonomicName id="41BA4D06FF94AF21FF6DF5D00F2FA2F1" authority="Topsent, 1912" authorityName="Topsent" authorityYear="1912" box="[215,531,976,1003]" class="Demospongiae" family="Hymedesmiidae" genus="Kirkpatrickia" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
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<emphasis id="B4CEEA97FF94AF21FF6DF5D00C52A2F1" box="[215,366,976,1002]" italics="true" pageId="2" pageNumber="157">Kirkpatrickia</emphasis>
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<bibRefCitation id="E22B4B74FF94AF21FECFF5D10F2FA2F1" author="Topsent, E." box="[373,531,976,1003]" pageId="2" pageNumber="157" pagination="1 - 7" refId="ref4729" refString="Topsent, E. (1912) Sur une grande Tedania abyssale des Acores (Tedania phacellina, n. sp.). Bulletin de l'Institut oceanographique, Monaco, 252, 1 - 7." type="journal article" year="1912">Topsent, 1912</bibRefCitation>
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</taxonomicName>
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has strongylote tornotes, lacks microscleres and “smooth or spined styles in plumose but anastomosing bundles.”
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</paragraph>
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<paragraph id="86053685FF94AF21FF7DF2190F0EA54E" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
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<emphasis id="B4CEEA97FF94AF21FF7DF2190DECA529" box="[199,208,1049,1074]" italics="true" pageId="2" pageNumber="157">-</emphasis>
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<taxonomicName id="41BA4D06FF94AF21FF6CF2180CD0A529" authority="Burton, 1929" authorityName="Burton" authorityYear="1929" box="[214,492,1048,1074]" class="Demospongiae" family="Hymedesmiidae" genus="Myxodoryx" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
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<emphasis id="B4CEEA97FF94AF21FF6CF2180C68A529" box="[214,340,1048,1074]" italics="true" pageId="2" pageNumber="157">Myxodoryx</emphasis>
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<bibRefCitation id="E22B4B74FF94AF21FEE0F2180CD0A529" author="Burton, M." box="[346,492,1048,1074]" pageId="2" pageNumber="157" pagination="393 - 458" refId="ref3688" refString="Burton, M. (1929) Porifera. Part II. Antarctic sponges. British Antarctic (' Terra Nova') Expedition, 1910. Natural History Report, British Museum (Natural History), London, Zoology, 6 (4), 393 - 458, pls. I-V. https: // biodiversitylibrary. org / page / 49467888" type="journal article" year="1929">Burton, 1929</bibRefCitation>
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</taxonomicName>
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has strongylote tornotes, two categories of acanthostyles “in plumose but anastomosing bundles” and lacks microscleres.
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</paragraph>
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<paragraph id="86053685FF94AF21FF7DF2600831A562" blockId="2.[151,1437,297,1360]" box="[199,1293,1119,1146]" pageId="2" pageNumber="157">
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<emphasis id="B4CEEA97FF94AF21FF7DF2600DECA562" box="[199,208,1120,1145]" italics="true" pageId="2" pageNumber="157">-</emphasis>
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<taxonomicName id="41BA4D06FF94AF21FF6DF25F0F97A562" authority="Duchassaing & Michelotti, 1864" authorityName="Duchassaing & Michelotti" authorityYear="1864" box="[215,683,1119,1145]" class="Demospongiae" family="Hymedesmiidae" genus="Phorbas" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
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<emphasis id="B4CEEA97FF94AF21FF6DF25F0C0AA562" box="[215,310,1119,1145]" italics="true" pageId="2" pageNumber="157">Phorbas</emphasis>
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<bibRefCitation id="E22B4B74FF94AF21FE87F25F0F97A562" author="Duchassaing de Fonbressin, P. & Michelotti, G." box="[317,683,1119,1145]" pageId="2" pageNumber="157" pagination="1 - 124" refId="ref3989" refString="Duchassaing de Fonbressin, P. & Michelotti, G. (1864) Spongiaires de la mer Caraibe. Natuurkundige verhandelingen van de Hollandsche maatschappij der wetenschappen te Haarlem, 21 (2), 1 - 124, pls. I-XXV. [https: // books. google. nl / books / about / Natuurkundige _ verhandelingen _ van _ de _ Holl. html? id = djVFAAAAcAAJ & redir _ esc = y & hl = nl]" type="journal article" year="1864">Duchassaing & Michelotti, 1864</bibRefCitation>
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</taxonomicName>
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has “plumose tracts of tornotes and/or acanthostyles.”
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</paragraph>
|
|
<paragraph id="86053685FF94AF21FF7DF2840E37A5DA" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
|
|
<emphasis id="B4CEEA97FF94AF21FF7DF2840DECA586" box="[199,208,1156,1181]" italics="true" pageId="2" pageNumber="157">-</emphasis>
|
|
<taxonomicName id="41BA4D06FF94AF21FF6DF2830F24A586" authority="Topsent, 1927" authorityName="Topsent" authorityYear="1927" box="[215,536,1155,1181]" class="Demospongiae" family="Hymedesmiidae" genus="Plocamionida" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
|
|
<emphasis id="B4CEEA97FF94AF21FF6DF2830C48A586" box="[215,372,1155,1181]" italics="true" pageId="2" pageNumber="157">Plocamionida</emphasis>
|
|
<bibRefCitation id="E22B4B74FF94AF21FEC0F2830F24A586" author="Topsent, E." box="[378,536,1155,1181]" pageId="2" pageNumber="157" pagination="1 - 19" refId="ref4766" refString="Topsent, E. (1927) Diagnoses d'Eponges nouvelles recueillies par le Prince Albert ler de Monaco. Bulletin de l'Institut oceanographique Monaco, 502, 1 - 19." type="journal article" year="1927">Topsent, 1927</bibRefCitation>
|
|
</taxonomicName>
|
|
has a “plocamiid basal skeleton. Megascleres mucronate tornotes, long acanthostyles, and basal acanthostrongyles.” Sigmas are absent.
|
|
</paragraph>
|
|
<paragraph id="86053685FF94AF21FF7DF2CB0C92A413" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
|
|
<emphasis id="B4CEEA97FF94AF21FF7DF2CB0DECA5FF" box="[199,208,1227,1252]" italics="true" pageId="2" pageNumber="157">-</emphasis>
|
|
<taxonomicName id="41BA4D06FF94AF21FF66F2CA0F65A5FF" authority="Carter, 1886" authorityName="Carter" authorityYear="1886" box="[220,601,1226,1253]" class="Demospongiae" family="Hymedesmiidae" genus="Pseudohalichondria" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
|
|
<emphasis id="B4CEEA97FF94AF21FF66F2CA0C83A5FF" box="[220,447,1226,1252]" italics="true" pageId="2" pageNumber="157">Pseudohalichondria</emphasis>
|
|
<bibRefCitation id="E22B4B74FF94AF21FE70F2CB0F65A5FF" author="Carter, H. J." box="[458,601,1226,1253]" pageId="2" pageNumber="157" pagination="454 - 455" refId="ref3909" refString="Carter, H. J. (1886) Supplement to the Descriptions of Mr. J. Bracebridge Wilson's Australian Sponges. Annals and Magazine of Natural History, Series 5, 18 (106), 271 - 290 + 369 - 379 + 445 - 466, pl. X, pp. 454 - 455, pl. C, figs. 6 - 9. https: // doi. org / 10.1080 / 00222938609459970" type="journal article" year="1886">Carter, 1886</bibRefCitation>
|
|
</taxonomicName>
|
|
has “plumose columns of smooth tornotes, without acanthostyles, with modified spined chelae.”
|
|
</paragraph>
|
|
<paragraph id="86053685FF94AF21FF7DF3130F3DA44B" blockId="2.[151,1437,297,1360]" pageId="2" pageNumber="157">
|
|
<emphasis id="B4CEEA97FF94AF21FF7DF3130DECA437" box="[199,208,1299,1324]" italics="true" pageId="2" pageNumber="157">-</emphasis>
|
|
<taxonomicName id="41BA4D06FF94AF21FF60F3120F39A437" authority="Topsent, 1890" authorityName="Topsent" authorityYear="1890" box="[218,517,1298,1324]" class="Demospongiae" family="Hymedesmiidae" genus="Spanioplon" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="157" phylum="Porifera" rank="genus">
|
|
<emphasis id="B4CEEA97FF94AF21FF60F3120C66A437" box="[218,346,1298,1324]" italics="true" pageId="2" pageNumber="157">Spanioplon</emphasis>
|
|
<bibRefCitation id="E22B4B74FF94AF21FED9F3120F39A437" author="Topsent, E." box="[355,517,1298,1324]" pageId="2" pageNumber="157" pagination="26 - 32" refId="ref4678" refString="Topsent, E. (1890) Notice preliminaire sur les spongiaires recueillis durant les campagnes de l'Hirondelle. Bulletin de la Societe zoologique de France, 15, 26 - 32 + 65 - 71. https: // doi. org / 10.5962 / bhl. part. 18721" type="journal article" year="1890">Topsent, 1890</bibRefCitation>
|
|
</taxonomicName>
|
|
“has non-echinating accessory acanthostyles or acanthoxeas. No chelae.” Ectosomal anisotornotes are smooth.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
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</document> |