treatments-xml/data/DC/7E/E8/DC7EE8592704FFD0F6BFFC05FDA7FE88.xml

<document id="79F13810BDE581383282E1CB12F1BA6C" ID-CLB-Dataset="24418" ID-DOI="10.11646/zootaxa.4718.3.7" ID-GBIF-Dataset="22a109e7-7d0f-41d4-adf0-6f5be8a7374e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3602550" ID-ZooBank="51AB6842-F063-4580-B30F-65102ABA797C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578565066580" checkinUser="plazi" docAuthor="Joshi, Sunil, Jose, Bindu K., Gullan, Penny, Sajeev, T. V. &amp; Anoop, E. V." docDate="2020" docId="DC7EE8592704FFD0F6BFFC05FDA7FE88" docLanguage="en" docName="zootaxa.4718.3.7.pdf" docOrigin="Zootaxa 4718 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="396" masterDocId="204790212707FFD5F628FFC5FFAFFFEF" masterDocTitle="A new species of mealybug (Hemiptera: Coccomorpha: Pseudococcidae) from Tectona grandis L. f. (Lamiaceae) in southern India" masterLastPageNumber="400" masterPageNumber="391" pageNumber="394" updateTime="1698776724599" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CD532F7C4DF09C28A95C806FF9D1138A">A new species of mealybug (Hemiptera: Coccomorpha: Pseudococcidae) from Tectona grandis L. f. (Lamiaceae) in southern India</mods:title>
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<mods:namePart id="4686346B1D8EA557D81CD45410FBD719">Joshi, Sunil</mods:namePart>
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<mods:namePart id="B0E07A1C20267B62E4DBA42FB7C932F1">Jose, Bindu K.</mods:namePart>
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<mods:namePart id="B636EBC652FEE003BE62495B2FC04911">Gullan, Penny</mods:namePart>
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<mods:namePart id="E11C01E0BE7DA397AAEF0D268A5F7D43">Sajeev, T. V.</mods:namePart>
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<mods:namePart id="BAB953FA3895A84529A9C5BDE8234861">Anoop, E. V.</mods:namePart>
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<emphasis id="66A3855D2704FFD6F6BFFC05FC87FC35" bold="true" box="[151,808,960,987]" pageId="3" pageNumber="394">
<taxonomicName id="93D722CC2704FFD6F6BFFC05FD71FC35" authority="Joshi, Bindu &amp; Gullan" authorityName="Joshi, Bindu &amp; Gullan. A. Body" authorityYear="2020" box="[151,734,960,987]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="tectonae" status="sp. nov.">
<emphasis id="66A3855D2704FFD6F6BFFC05FE12FC34" bold="true" box="[151,445,960,987]" italics="true" pageId="3" pageNumber="394">Formicococcus tectonae</emphasis>
Joshi, Bindu &amp; Gullan
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urn:lsid:zoobank.org:act: 
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<emphasis id="66A3855D2704FFD6F6BFFBE9FEEAFBA9" bold="true" box="[151,325,1067,1094]" pageId="3" pageNumber="394">Type material.</emphasis>
<materialsCitation id="E4BF53122704FFD6F764FBE9FDBBFB61" ID-GBIF-Occurrence="2545451401" collectingDate="2018-12-07" collectionCode="ICAR, NBAIR" collectorName="Bindu Jose" country="India" location="Thrissur" municipality="Vettigapadam" pageId="3" pageNumber="394" specimenCount="1" specimenCount-female="1" stateProvince="Kerala" typeStatus="holotype">
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<typeStatus id="8B6CE7ED2704FFD6F764FBE9FE16FBA9" box="[332,441,1068,1094]" pageId="3" pageNumber="394" type="holotype">Holotype</typeStatus>
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adult 
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on a slide together with four paratypes, 
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, 
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/ 
<collectingMunicipality id="B40CC3352704FFD6F2BFFBE9FA9BFBA9" box="[1175,1332,1068,1094]" pageId="3" pageNumber="394">Vettigapadam</collectingMunicipality>
, 
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/ 
<taxonomicName id="93D722CC2704FFD6F68EFB94FEF4FB85" box="[166,347,1104,1130]" class="Magnoliopsida" family="Lamiaceae" genus="Tectona" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="394" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="66A3855D2704FFD6F68EFB94FEF4FB85" box="[166,347,1104,1130]" italics="true" pageId="3" pageNumber="394">Tectona grandis</emphasis>
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, 
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<collectingDate id="302D86672704FFD6F741FB95FE5DFB85" box="[361,498,1104,1130]" pageId="3" pageNumber="394" value="2018-12-07">07.XII.2018</collectingDate>
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/ 
<collectorName id="F9223C992704FFD6F421FB95FD2BFB84" box="[521,644,1104,1131]" pageId="3" pageNumber="394">Bindu Jose</collectorName>
leg. / [
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/ 
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/PSEUDO/Form/71218-01]; holotype marked by encircling with permanent marker
</materialsCitation>
. 
<materialsCitation id="E4BF53122704FFD6F435FBB1FAD2FB5D" ID-GBIF-Occurrence="2545451402" collectingDate="2018-12-07" collectionCode="ICAR, NBAIR" collectorName="Bindu Jose" country="India" location="Thrissur" municipality="Vettigapadam" pageId="3" pageNumber="394" specimenCount="9" specimenCount-female="9" stateProvince="Kerala" typeStatus="paratype">
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<typeStatus id="8B6CE7ED2704FFD6F435FBB1FD3AFB61" box="[541,661,1140,1166]" pageId="3" pageNumber="394" type="paratype">Paratypes</typeStatus>
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: 
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, data same as holotype; four paratypes on the same slide as the holotype; the remaining five paratypes are mounted on a separate slide [
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/ 
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/PSEUDO/Form/71218-02]
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</paragraph>
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<emphasis id="66A3855D2704FFD6F6EFFB7EFD90FB39" bold="true" box="[199,575,1211,1238]" pageId="3" pageNumber="394">Description of the adult female.</emphasis>
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<paragraph id="5468594F2704FFD6F6EFFB24FBCCFA40" blockId="3.[151,1437,1067,2031]" pageId="3" pageNumber="394">
<emphasis id="66A3855D2704FFD6F6EFFB24FE25FB15" box="[199,394,1249,1274]" italics="true" pageId="3" pageNumber="394">Live appearance.</emphasis>
Body oval or rotund, peach-pink; legs and antennae yellowish brown; bodies of nymphs and adult female covered by flocculent white mealy wax (
<figureCitation id="CCEC45CA2704FFD6F4DAFAC1FC25FAF1" box="[754,906,1284,1311]" captionStart="FIGURE 1" captionStartId="5.[151,250,1860,1885]" captionTargetBox="[160,1427,373,1840]" captionTargetId="figure@5.[160,1427,373,1840]" captionTargetPageId="5" captionText="FIGURE 1. A. Site of observations in the Palappalli forest range; B. Teak tree affected with trunk borer, Cossus cadambae; C. Entry holes made by the larvae of trunk borer; D. Infestation of Formicococcus tectonae Joshi, Bindu &amp; Gullan at the entry hole of the borer and an ant attending the mealybugs; E. Eggs of F. tectonae; F. Nymph hidden under the loose bark; G. Adult female." figureDoi="http://doi.org/10.5281/zenodo.3602552" httpUri="https://zenodo.org/record/3602552/files/figure.png" pageId="3" pageNumber="394">Fig. 1 F &amp; G</figureCitation>
), without bare areas on dorsum; intersegmental areas without mealy wax; very short lateral wax filaments present on abdominal segments, the pair on the last abdominal segment longer and curved; wax filaments absent from thorax and head. Ovisac absent; eggs gleaming cream (
<figureCitation id="CCEC45CA2704FFD6F6C3FAB5FEE5FA64" box="[235,330,1392,1419]" captionStart="FIGURE 1" captionStartId="5.[151,250,1860,1885]" captionTargetBox="[160,1427,373,1840]" captionTargetId="figure@5.[160,1427,373,1840]" captionTargetPageId="5" captionText="FIGURE 1. A. Site of observations in the Palappalli forest range; B. Teak tree affected with trunk borer, Cossus cadambae; C. Entry holes made by the larvae of trunk borer; D. Infestation of Formicococcus tectonae Joshi, Bindu &amp; Gullan at the entry hole of the borer and an ant attending the mealybugs; E. Eggs of F. tectonae; F. Nymph hidden under the loose bark; G. Adult female." figureDoi="http://doi.org/10.5281/zenodo.3602552" httpUri="https://zenodo.org/record/3602552/files/figure.png" pageId="3" pageNumber="394">Fig. 1 E</figureCitation>
), laid singly, not covered with mealy wax powder. All life stages occurring on above-ground parts of host, under the bark in borer tunnels, and are sometimes attended by ants (
<figureCitation id="CCEC45CA2704FFD6F5DDFA51FBF8FA40" box="[1013,1111,1428,1455]" captionStart="FIGURE 1" captionStartId="5.[151,250,1860,1885]" captionTargetBox="[160,1427,373,1840]" captionTargetId="figure@5.[160,1427,373,1840]" captionTargetPageId="5" captionText="FIGURE 1. A. Site of observations in the Palappalli forest range; B. Teak tree affected with trunk borer, Cossus cadambae; C. Entry holes made by the larvae of trunk borer; D. Infestation of Formicococcus tectonae Joshi, Bindu &amp; Gullan at the entry hole of the borer and an ant attending the mealybugs; E. Eggs of F. tectonae; F. Nymph hidden under the loose bark; G. Adult female." figureDoi="http://doi.org/10.5281/zenodo.3602552" httpUri="https://zenodo.org/record/3602552/files/figure.png" pageId="3" pageNumber="394">Fig. 1 D</figureCitation>
).
</paragraph>
<paragraph id="5468594F2704FFD1F6EFFA7DFE22FD68" blockId="3.[151,1437,1067,2031]" lastBlockId="4.[151,1437,153,2046]" lastPageId="4" lastPageNumber="395" pageId="3" pageNumber="394">
<emphasis id="66A3855D2704FFD6F6EFFA7DFE54FA3D" box="[199,507,1464,1490]" italics="true" pageId="3" pageNumber="394">Slide-mounted adult female</emphasis>
(
<figureCitation id="CCEC45CA2704FFD6F422FA7DFDE0FA3D" box="[522,591,1464,1490]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="3" pageNumber="394">Fig. 2</figureCitation>
). Measurements based on 
<specimenCount id="42D192C62704FFD6F55FFA7DFC65FA3C" box="[887,970,1464,1491]" pageId="3" pageNumber="394" type="generic">10 type</specimenCount>
specimens. Body of adult female broadly oval to almost circular (
<figureCitation id="CCEC45CA2704FFD6F799FA19FDB8FA19" box="[433,535,1500,1526]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="3" pageNumber="394">Fig. 2 A</figureCitation>
, 
<figureCitation id="CCEC45CA2704FFD6F40DFA19FD24FA19" box="[549,651,1500,1526]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="3" pageNumber="394">Fig. 3 A</figureCitation>
), membranous; largest specimen 
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long and 
<quantity id="932FF4AA2704FFD6F2DBFA19FAF4FA18" box="[1267,1371,1500,1527]" metricMagnitude="-3" metricUnit="m" metricValue="2.37" pageId="3" pageNumber="394" unit="mm" value="2.37">2.37 mm</quantity>
wide (
<typeStatus id="8B6CE7ED2704FFD6F6B7F9C5FEACF9F5" box="[159,259,1536,1562]" pageId="3" pageNumber="394" type="holotype">holotype</typeStatus>
<quantity id="932FF4AA2704FFD6F721FA3AFEC2F9F4" box="[265,365,1535,1563]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="3" pageNumber="394" unit="mm" value="2.05">2.05 mm</quantity>
long, 
<quantity id="932FF4AA2704FFD6F79AFA3AFDBAF9F4" box="[434,533,1535,1563]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="394" unit="mm" value="1.5">1.50 mm</quantity>
wide). Eyespots each 
<quantity id="932FF4AA2704FFD6F526F9C5FC2FF9F4" box="[782,896,1536,1563]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="3" pageNumber="394" unit="mm" value="38.5" valueMax="40.0" valueMin="37.0">37–40 µm</quantity>
in diameter, not associated with discoidal pores. Antennae 7 segmented each about 
<quantity id="932FF4AA2704FFD6F435F9E1FDDBF9D0" box="[541,628,1572,1599]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="394" unit="mm" value="320.0">320 µm</quantity>
long (
<figureCitation id="CCEC45CA2704FFD6F494F9E1FCB5F9D0" box="[700,794,1572,1599]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="3" pageNumber="394">Fig. 2 B</figureCitation>
, 
<figureCitation id="CCEC45CA2704FFD6F50EF9E1FC2CF9D0" box="[806,899,1572,1599]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="3" pageNumber="394">Fig. 3 B</figureCitation>
); apical segment 
<quantity id="932FF4AA2704FFD6F26FF9E1FB15F9D0" box="[1095,1210,1572,1599]" metricMagnitude="-2" metricUnit="m" metricValue="9.6" metricValueMax="9.8" metricValueMin="9.4" pageId="3" pageNumber="394" unit="mm" value="96.0" valueMax="98.0" valueMin="94.0">94–98 µm</quantity>
long and 
<quantity id="932FF4AA2704FFD6F301F9E6FA33F9D0" box="[1321,1436,1571,1599]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="3" pageNumber="394" unit="mm" value="26.0" valueMax="27.0" valueMin="25.0">25–27 µm</quantity>
wide. Labium 
<quantity id="932FF4AA2704FFD6F714F982FE1FF98C" box="[316,432,1607,1635]" metricMagnitude="-2" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="3" pageNumber="394" unit="mm" value="92.5" valueMax="95.0" valueMin="90.0">90–95 µm</quantity>
wide at base and 
<quantity id="932FF4AA2704FFD6F456F982FCA0F98C" box="[638,783,1607,1635]" metricMagnitude="-1" metricUnit="m" metricValue="2.075" metricValueMax="2.15" metricValueMin="2.0" pageId="3" pageNumber="394" unit="mm" value="207.5" valueMax="215.0" valueMin="200.0">200–215 µm</quantity>
long, noticeably longer than clypeolabral shield, at about 
<quantity id="932FF4AA2704FFD6F6BFF9A9FF5FF968" box="[151,240,1644,1671]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="394" unit="mm" value="190.0">190 µm</quantity>
long. Mesothoracic spiracles each 
<quantity id="932FF4AA2704FFD6F4A9F9AEFD5AF968" box="[641,757,1643,1671]" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="3" pageNumber="394" unit="mm" value="77.5" valueMax="80.0" valueMin="75.0">75–80 µm</quantity>
long including apodeme, 
<quantity id="932FF4AA2704FFD6F237F9A9FB3CF968" box="[1055,1171,1644,1671]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="3" pageNumber="394" unit="mm" value="39.5" valueMax="42.0" valueMin="37.0">37–42 µm</quantity>
wide across peritreme; metathoracic spiracles each 
<quantity id="932FF4AA2704FFD6F7FCF955FDE8F944" box="[468,583,1680,1707]" metricMagnitude="-2" metricUnit="m" metricValue="8.45" metricValueMax="8.7" metricValueMin="8.2" pageId="3" pageNumber="394" unit="mm" value="84.5" valueMax="87.0" valueMin="82.0">82–87 µm</quantity>
long including apodeme, 
<quantity id="932FF4AA2704FFD6F545F94AFC4FF944" box="[877,992,1679,1707]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="3" pageNumber="394" unit="mm" value="50.0" valueMax="55.0" valueMin="45.0">45–55 µm</quantity>
wide across peritreme. Anal lobes well developed, ventral surface of each lobe bearing stout apical seta about 
<quantity id="932FF4AA2704FFD6F582F976FBACF920" box="[938,1027,1715,1743]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="394" unit="mm" value="250.0">250 µm</quantity>
, bar seta 
<quantity id="932FF4AA2704FFD6F243F971FB56F920" box="[1131,1273,1716,1743]" metricMagnitude="-1" metricUnit="m" metricValue="1.21" metricValueMax="1.3" metricValueMin="1.1199999999999999" pageId="3" pageNumber="394" unit="mm" value="121.0" valueMax="130.0" valueMin="112.0">112–130 µm</quantity>
long, and anal lobe bar about 
<quantity id="932FF4AA2704FFD6F76AF91DFE35F91C" box="[322,410,1752,1779]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="394" unit="mm" value="100.0">100 µm</quantity>
long (
<figureCitation id="CCEC45CA2704FFD6F7CDF91DFDEAF91D" box="[485,581,1752,1778]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="3" pageNumber="394">Fig. 2 C</figureCitation>
, 
<figureCitation id="CCEC45CA2704FFD6F47AF91DFD1DF91D" box="[594,690,1752,1778]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="3" pageNumber="394">Fig. 3 C</figureCitation>
), developed mainly forwards from bar seta. Legs well developed; hind trochanter + femur about 
<quantity id="932FF4AA2704FFD6F7C4F939FDD4F8F8" box="[492,635,1788,1815]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="3" pageNumber="394" unit="mm" value="275.0" valueMax="280.0" valueMin="270.0">270–280 µm</quantity>
long, hind tibia + tarsus 
<quantity id="932FF4AA2704FFD6F5B8F939FBB0F8F8" box="[912,1055,1788,1815]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="3" pageNumber="394" unit="mm" value="250.0" valueMax="260.0" valueMin="240.0">240–260 µm</quantity>
long; claw stout, about 
<quantity id="932FF4AA2704FFD6F302F939FA33F8F8" box="[1322,1436,1788,1815]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="3" pageNumber="394" unit="mm" value="38.5" valueMax="40.0" valueMin="37.0">37–40 µm</quantity>
long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur about 0.96. Ratio of lengths of hind tibia to tarsus about 1.50. Hind trochanter with longest seta 
<quantity id="932FF4AA2704FFD6F4F4F881FCE1F8B0" box="[732,846,1860,1887]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" metricValueMax="7.0" metricValueMin="6.2" pageId="3" pageNumber="394" unit="mm" value="66.0" valueMax="70.0" valueMin="62.0">62–70 µm</quantity>
long; tarsal digitules on each leg capitate, 
<quantity id="932FF4AA2704FFD6F302F886FA33F8B0" box="[1322,1436,1859,1887]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="3" pageNumber="394" unit="mm" value="52.5" valueMax="55.0" valueMin="50.0">50–55 µm</quantity>
long; claw digitules capitate, 
<quantity id="932FF4AA2704FFD6F7C9F8ADFDFAF86C" box="[481,597,1896,1923]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="3" pageNumber="394" unit="mm" value="31.5" valueMax="33.0" valueMin="30.0">30–33 µm</quantity>
long; claw denticle not discernible (
<figureCitation id="CCEC45CA2704FFD6F5C6F8ADFBFFF86C" box="[1006,1104,1896,1923]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="3" pageNumber="394">Fig. 2 D</figureCitation>
). A total of 37–40 translucent pores present on anterior and posterior surfaces of hind coxa (22–26 on anterior surface and 14–15 on posterior surface) (
<figureCitation id="CCEC45CA2704FFD6F728F875FEF2F824" box="[256,349,1968,1995]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="3" pageNumber="394">Fig. 2 E</figureCitation>
, 
<figureCitation id="CCEC45CA2704FFD6F741F875FE67F824" box="[361,456,1968,1995]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="3" pageNumber="394">Fig. 3 D</figureCitation>
, and translucent pores on posterior surface of hind tibia rare (2 out of 
<specimenCount id="42D192C62704FFD6F2FDF875FAE0F825" box="[1237,1359,1968,1994]" pageId="3" pageNumber="394" type="female">10 females</specimenCount>
with 6 to 7 pores on posterior surface) or absent (
<figureCitation id="CCEC45CA2704FFD6F45AF811FD63F800" box="[626,716,2004,2031]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="3" pageNumber="394">Fig. 2 F</figureCitation>
, 
<figureCitation id="CCEC45CA2704FFD6F4FFF811FC82F801" box="[727,813,2004,2030]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="3" pageNumber="394">Fig. 3E</figureCitation>
). Circulus present between abdominal segments III and IV irregular in shape (
<figureCitation id="CCEC45CA2703FFD1F7BAFF5CFE5BFF5C" box="[402,500,153,179]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 G</figureCitation>
), 
<quantity id="932FF4AA2703FFD1F421FF5CFD36FF5C" box="[521,665,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="4" pageNumber="395" unit="mm" value="110.0" valueMax="120.0" valueMin="100.0">100–120 µm</quantity>
wide, divided by an intersegmental line. Trilocular pores unusual in appearance, with a smaller triangle containing 3 swirled loculi situated within a larger outer triangle (
<figureCitation id="CCEC45CA2703FFD1F2DFFF78FAFCFF38" box="[1271,1363,189,215]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 L</figureCitation>
). Both pairs of ostioles well developed (
<figureCitation id="CCEC45CA2703FFD1F424FF24FDC0FF14" box="[524,623,225,251]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 H</figureCitation>
); anterior ostioles 
<quantity id="932FF4AA2703FFD1F517FF25FC60FF14" box="[831,975,224,251]" metricMagnitude="-1" metricUnit="m" metricValue="1.125" metricValueMax="1.25" metricValueMin="1.0" pageId="4" pageNumber="395" unit="mm" value="112.5" valueMax="125.0" valueMin="100.0">100–125 µm</quantity>
wide, each lip with 8–9 setae and 32–50 trilocular pores; posterior ostiole 
<quantity id="932FF4AA2703FFD1F43DFEC1FD25FEF0" box="[533,650,260,287]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" metricValueMax="6.2" metricValueMin="5.2" pageId="4" pageNumber="395" unit="mm" value="57.0" valueMax="62.0" valueMin="52.0">52–62 µm</quantity>
wide, each lip with 7–8 setae and 40–51 trilocular. Anal ring 55–75 μm long and 65–85 μm wide (
<figureCitation id="CCEC45CA2703FFD1F7C6FEECFDE4FEAC" box="[494,587,297,323]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 F</figureCitation>
), situated at apex of abdomen; anal ring with 6 setae, each 90–95 μm long. Number of cerarii highly variable, their number varying from 11 to 15 pairs (only 1 out of 
<specimenCount id="42D192C62703FFD1F283FE88FA88FE88" box="[1195,1319,333,359]" pageId="4" pageNumber="395" type="female">10 females</specimenCount>
examined possessed 18 clearly defined pairs of cerarii); all cerarian setae conical with flagellate tips. Anal lobe cerarii each bearing 4 or 5 conical setae (
<figureCitation id="CCEC45CA2703FFD1F7FEFE50FD85FE40" box="[470,554,405,431]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">
Fig. 
<date id="20697F8F2703FFD1F422FE50FD85FE40" box="[522,554,405,431]" pageId="4" pageNumber="395">2 I</date>
</figureCitation>
, 
<figureCitation id="CCEC45CA2703FFD1F41DFE50FD22FE40" box="[565,653,405,431]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3G</figureCitation>
), each 
<quantity id="932FF4AA2703FFD1F4F3FE50FCE2FE40" box="[731,845,405,431]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="4" pageNumber="395" unit="mm" value="25.0" valueMax="28.0" valueMin="22.0">22–28 µm</quantity>
long and about 
<quantity id="932FF4AA2703FFD1F5D7FE51FBE0FE40" box="[1023,1103,404,431]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="395" unit="mm" value="7.5">7.5 µm</quantity>
wide at base, with 1 auxiliary seta and a few trilocular pores, all situated on a membranous area. Penultimate cerarii (C 
<subScript id="C8535B0A2703FFD1F25AFE0FFB2DFE37" attach="left" box="[1138,1154,458,472]" fontSize="6" pageId="4" pageNumber="395">17</subScript>
) each with 7 or 8 conical setae, 1 or 2 auxiliary setae (
<figureCitation id="CCEC45CA2703FFD1F7C9FE18FD95FE18" box="[481,570,477,503]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 J</figureCitation>
, 
<figureCitation id="CCEC45CA2703FFD1F460FE18FD30FE18" box="[584,671,477,503]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3H</figureCitation>
) and a few trilocular pores. Other abdominal cerarii as far forward as abdominal segment II each containing 2–5 conical setae, 1 or 2 auxiliary setae and few trilocular pores. Cerarii on thorax not discernible. Frontal cerarii each with 4 conical setae (
<figureCitation id="CCEC45CA2703FFD1F5A5FDE0FC41FDD0" box="[909,1006,549,575]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 K</figureCitation>
), preocular cerarii each with 3 conical setae and ocular cerarii each with 2 conical and 1 auxiliary setae (
<figureCitation id="CCEC45CA2703FFD1F55DFD8CFC7EFD8C" box="[885,977,585,611]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 L</figureCitation>
, 
<figureCitation id="CCEC45CA2703FFD1F5F5FD8CFB85FD8C" box="[989,1066,585,611]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">
Fig. 
<date id="20697F8F2703FFD1F238FD8CFB85FD8C" box="[1040,1066,585,611]" pageId="4" pageNumber="395">3I</date>
</figureCitation>
). All cerarii each associated with a few trilocular pores.
</paragraph>
<paragraph id="5468594F2703FFD1F6EFFD54FB8FFCB0" blockId="4.[151,1437,153,2046]" pageId="4" pageNumber="395">
Dorsal surface with thick, stiff setae each with a flagellate tip (
<figureCitation id="CCEC45CA2703FFD1F585FD54FBB6FD44" box="[941,1049,657,683]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 M</figureCitation>
, 
<figureCitation id="CCEC45CA2703FFD1F20EFD54FB2DFD44" box="[1062,1154,657,683]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 J</figureCitation>
), of variable size, fairly numerous on median area. Smaller setae, each 12–17 μm long, medium setae each 25–30 μm long, and longer setae mostly each 35–37 μm long (
<figureCitation id="CCEC45CA2703FFD1F7F5FD1CFD99FD1C" box="[477,566,729,755]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3K</figureCitation>
). Many setae each with 1 or 2 trilocular pores around setal collar. Even longer setae present dorsally just anterior to clypeolabral shield between antennal bases, each 55–65 μm long. Trilocular pores numerous (
<figureCitation id="CCEC45CA2703FFD1F770FCE4FE1AFCD4" box="[344,437,801,827]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 L</figureCitation>
), evenly distributed, each about 
<quantity id="932FF4AA2703FFD1F508FCE5FCF5FCD4" box="[800,858,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="395" unit="mm" value="5.0">5 µm</quantity>
wide. Discoidal pores (
<figureCitation id="CCEC45CA2703FFD1F24BFCE4FB67FCD4" box="[1123,1224,801,827]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 M</figureCitation>
), each as wide as a trilocular pore, scattered. Oral collar tubular ducts and multilocular pores absent.
</paragraph>
<paragraph id="5468594F2703FFD1F6EFFCACFECFFAE1" blockId="4.[151,1437,153,2046]" pageId="4" pageNumber="395">
Ventral surface with flagellate setae (
<figureCitation id="CCEC45CA2703FFD1F440FCACFD65FC6C" box="[616,714,873,899]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 N</figureCitation>
, 
<figureCitation id="CCEC45CA2703FFD1F4FEFCACFC99FC6C" box="[726,822,873,899]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 N</figureCitation>
), mostly each 
<quantity id="932FF4AA2703FFD1F5F1FCACFBE2FC6C" box="[985,1101,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="6.7" metricValueMin="2.0" pageId="4" pageNumber="395" unit="mm" value="43.5" valueMax="67.0" valueMin="20.0">20–67 µm</quantity>
long, but some on abdominal margins each 
<quantity id="932FF4AA2703FFD1F71CFC49FE07FC48" box="[308,424,908,935]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="4" pageNumber="395" unit="mm" value="62.5" valueMax="65.0" valueMin="60.0">60–65 µm</quantity>
long. Ventral setae on head, just anterior to mouthparts, each about 
<quantity id="932FF4AA2703FFD1F286FC49FB57FC48" box="[1198,1272,908,935]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="395" unit="mm" value="75.0">75 µm</quantity>
long, whereas median areas of thorax and abdomen with setae each 25 to 67 μm long. Cisanal setae stout at base, each about 
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long; obanal setae each about 
<quantity id="932FF4AA2703FFD1F40AFC11FDC3FC00" box="[546,620,980,1007]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="395" unit="mm" value="50.0">50 µm</quantity>
long. Multilocular disc pores each 9.8–10.0 µm in diameter (
<figureCitation id="CCEC45CA2703FFD1F31CFC10FA37FC00" box="[1332,1432,981,1007]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 O</figureCitation>
, 
<figureCitation id="CCEC45CA2703FFD1F6BFFC3CFF5AFBFC" box="[151,245,1017,1043]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 O</figureCitation>
), present posterior to vulva and medially at posterior edges of abdominal segments IV-VII; occasional pores also present on anterior edges of segments and on submargins of abdomen. Multilocular disc pores forming multiple transverse rows on abdominal segments VI and VII, a double row on V and a single row on segment IV. Trilocular pores same size and structure as those on dorsum but less numerous, evenly distributed. Discoidal pores scattered. Oral collar tubular ducts of 2 slightly different sizes present, both 
<typeStatus id="8B6CE7ED2703FFD1F554FB4CFC18FB4C" box="[892,951,1161,1187]" pageId="4" pageNumber="395">types</typeStatus>
narrower than a trilocular pore; wider 
<typeStatus id="8B6CE7ED2703FFD1F344FB4CFA33FB4C" box="[1388,1436,1161,1187]" pageId="4" pageNumber="395">type</typeStatus>
(
<figureCitation id="CCEC45CA2703FFD1F688FB69FF57FB28" box="[160,248,1196,1223]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 P</figureCitation>
, 
<figureCitation id="CCEC45CA2703FFD1F72CFB69FEF2FB28" box="[260,349,1196,1223]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 P</figureCitation>
) present submarginally on abdominal segments VI and VII; and slender 
<typeStatus id="8B6CE7ED2703FFD1F2A3FB68FB14FB28" box="[1163,1211,1197,1223]" pageId="4" pageNumber="395">type</typeStatus>
(
<figureCitation id="CCEC45CA2703FFD1F2E2FB69FA84FB29" box="[1226,1323,1196,1222]" captionStart="FIGURE 2" captionStartId="6.[151,250,1770,1795]" captionTargetBox="[151,1436,216,1684]" captionTargetId="figure@6.[151,1436,216,1687]" captionTargetPageId="6" captionText="FIGURE 2. Diagnostic features of a slide-mounted adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Female broadly oval to rotund; B. Antenna with seven segments; C. Anal lobe bar with bar seta; D. Claw without a denticle; E. Hind coxa with translucent pores; F. Hind tibia without translucent pores; G. Circulus irregular in shape; H. Ostiole well developed; I. Anal lobe cerarius with 4–5 conical setae and 1 auxiliary seta; J. Penultimate cerarius with 7 or 8 conical and 1 or 2 auxiliary setae; K. Frontal cerarius with 4 conical setae; L. Ocular cerarius with 2 conical and 1 auxiliary seta; M. Dorsal seta thick and stiff; N. Ventral setae flagellate; O. Multilocular disc pores; P. Submarginal oral collar tubular ducts; Q. Median oral collar tubular ducts." figureDoi="http://doi.org/10.5281/zenodo.3602554" httpUri="https://zenodo.org/record/3602554/files/figure.png" pageId="4" pageNumber="395">Fig. 2 Q</figureCitation>
, 
<figureCitation id="CCEC45CA2703FFD1F31EFB69FA3BFB29" box="[1334,1428,1196,1222]" captionStart="FIGURE 3" captionStartId="7.[151,250,1841,1866]" captionTargetBox="[289,1347,284,1730]" captionTargetId="figure@7.[151,1435,171,1757]" captionTargetPageId="7" captionText="FIGURE 3. Taxonomic illustration of an adult female of Formicococcus tectonae Joshi, Bindu &amp; Gullan. A. Body derm; B. Antenna; C. Ventral surface of anal lobe showing anal lobe bar; D. Coxa with translucent pores; E. Tibia without translucent pores; F. Anal ring with 3 pairs of setae; G. Anal lobe cerarius; H. Penultimate cerarius; I. Preocular cerarius; J. Dorsal seta (shorter type); K. Dorsal seta (longer type); L. Trilocular pores (dorsal and ventral); M. Discoidal pores (dorsal and ventral); N. Ventral seta; O. Multilocular disc pore; P. Tubular duct on marginal area; Q. Tubular duct on median area." figureDoi="http://doi.org/10.5281/zenodo.3602556" httpUri="https://zenodo.org/record/3602556/files/figure.png" pageId="4" pageNumber="395">Fig. 3 Q</figureCitation>
) distributed across middle of abdominal segments V and VI, posterior to vulva, and in marginal groups on abdominal segments V–VIII.
</paragraph>
</subSubSection>
<subSubSection id="1CCD0AC42703FFD1F6EFFAD2FC05F930" pageId="4" pageNumber="395" type="discussion">
<paragraph id="5468594F2703FFD1F6EFFAD2FE37F99B" blockId="4.[151,1437,153,2046]" pageId="4" pageNumber="395">
<emphasis id="66A3855D2703FFD1F6EFFAD2FEFDFADD" bold="true" box="[199,338,1303,1330]" pageId="4" pageNumber="395">Comments.</emphasis>
<taxonomicName id="93D722CC2703FFD1F774FADDFDD7FADE" authorityName="Joshi, Bindu &amp; Gullan. A. Body" authorityYear="2020" box="[348,632,1304,1329]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="tectonae">
<emphasis id="66A3855D2703FFD1F774FADDFDD7FADE" box="[348,632,1304,1329]" italics="true" pageId="4" pageNumber="395">Formicococcus tectonae</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="93D722CC2703FFD1F532FADDFB0BFADD" authority="(Ezzat &amp; McConnell)" baseAuthorityName="Ezzat &amp; McConnell" box="[794,1188,1303,1330]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="66A3855D2703FFD1F532FADDFC35FADD" box="[794,922,1304,1330]" italics="true" pageId="4" pageNumber="395">F. robustus</emphasis>
(Ezzat &amp; McConnell)
</taxonomicName>
in having more than 3 conical setae in most cerarii including the anal lobe pair; cerarii on the abdomen containing auxiliary setae; and similar distributions of ventral oral collar tubular ducts and 
<typeStatus id="8B6CE7ED2703FFD1F559FAA5FC03FA95" box="[881,940,1376,1402]" pageId="4" pageNumber="395">types</typeStatus>
of dorsal and ventral setae. 
<taxonomicName id="93D722CC2703FFD1F2D8FAA5FF59FA72" authorityName="Joshi, Bindu &amp; Gullan. A. Body" authorityYear="2020" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="tectonae">
<emphasis id="66A3855D2703FFD1F2D8FAA5FF59FA72" italics="true" pageId="4" pageNumber="395">Formicococcus tectonae</emphasis>
</taxonomicName>
can be distinguished (characters of 
<taxonomicName id="93D722CC2703FFD1F4B9FA41FCA2FA72" box="[657,781,1411,1437]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="66A3855D2703FFD1F4B9FA41FCA2FA72" box="[657,781,1411,1437]" italics="true" pageId="4" pageNumber="395">F. robustus</emphasis>
</taxonomicName>
given in parentheses) by having (i) multilocular pores on segments IV–VIII (V–VIII); (ii) cerarian setae conical with flagellate tips (conical but sometimes 1 or more setae with flagellate tips); and (iii) by the absence or rarity of translucent pores on the hind tibia (present). 
<taxonomicName id="93D722CC2703FFD1F2DAFA0EFF59F9E7" authorityName="Joshi, Bindu &amp; Gullan. A. Body" authorityYear="2020" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="tectonae">
<emphasis id="66A3855D2703FFD1F2DAFA0EFF59F9E7" italics="true" pageId="4" pageNumber="395">Formicococcus tectonae</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName id="93D722CC2703FFD1F7E1FA2AFD62F9E7" authority="Williams" authorityName="Williams" authorityYear="2004" box="[457,717,1518,1544]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="polysperes">
<emphasis id="66A3855D2703FFD1F7E1FA2AFDF1F9E7" box="[457,606,1518,1544]" italics="true" pageId="4" pageNumber="395">F. polysperes</emphasis>
Williams
</taxonomicName>
in having multilocular disc pores as far forward as abdominal segment IV, but differs (characters of 
<taxonomicName id="93D722CC2703FFD1F414F9D6FD19F9C3" authorityName="Joshi, Bindu &amp; Gullan. A. Body" authorityYear="2020" box="[572,694,1555,1580]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="tectonae">
<emphasis id="66A3855D2703FFD1F414F9D6FD19F9C3" box="[572,694,1555,1580]" italics="true" pageId="4" pageNumber="395">F. tectonae</emphasis>
</taxonomicName>
given in parentheses) in having stout, stiff dorsal setae (thick, stiff setae with a flagellate tip) and in having oral collar tubular ducts on the thorax and head (oral collar ducts absent from thorax and head).
</paragraph>
<paragraph id="5468594F2703FFD1F6EFF9B8FC05F930" blockId="4.[151,1437,153,2046]" pageId="4" pageNumber="395">
In view of the unusual structure of the trilocular pores in 
<taxonomicName id="93D722CC2703FFD1F54BF9BBFC4DF978" authority="Joshi &amp; Jose &amp; Gullan &amp; Sajeev &amp; Anoop, 2020" authorityName="Joshi &amp; Jose &amp; Gullan &amp; Sajeev &amp; Anoop" authorityYear="2020" box="[867,994,1662,1687]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="tectonae" status="sp. nov.">
<emphasis id="66A3855D2703FFD1F54BF9BBFC4DF978" box="[867,994,1662,1687]" italics="true" pageId="4" pageNumber="395">F. tectonae</emphasis>
</taxonomicName>
<emphasis id="66A3855D2703FFD1F5C2F9B8FB83F978" bold="true" box="[1002,1068,1661,1687]" pageId="4" pageNumber="395">
<taxonomicNameLabel id="7D9038262703FFD1F5C2F9B8FB83F978" box="[1002,1068,1661,1687]" pageId="4" pageNumber="395" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, those of some other species of 
<taxonomicName id="93D722CC2703FFD1F6BFF964FEEBF955" authorityName="Takahashi" authorityYear="1928" box="[151,324,1697,1722]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="66A3855D2703FFD1F6BFF964FEEBF955" box="[151,324,1697,1722]" italics="true" pageId="4" pageNumber="395">Formicococcus</emphasis>
</taxonomicName>
were examined to see if this feature is shared with congeneric species. Trilocular pores with a similar structure were observed in 
<taxonomicName id="93D722CC2703FFD1F7EEF900FD90F931" authorityName="Williams" authorityYear="2004" box="[454,575,1732,1758]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="lingnani">
<emphasis id="66A3855D2703FFD1F7EEF900FD90F931" box="[454,575,1732,1758]" italics="true" pageId="4" pageNumber="395">F. lingnani</emphasis>
</taxonomicName>
, 
<taxonomicName id="93D722CC2703FFD1F465F900FD4FF931" authorityName="Williams" authorityYear="2004" box="[589,736,1732,1758]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="polysperes">
<emphasis id="66A3855D2703FFD1F465F900FD4FF931" box="[589,736,1732,1758]" italics="true" pageId="4" pageNumber="395">F. polysperes</emphasis>
</taxonomicName>
and 
<taxonomicName id="93D722CC2703FFD1F53EF900FC0CF930" box="[790,931,1733,1759]" class="Insecta" family="Pseudococcidae" genus="Formicococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="formicarii">
<emphasis id="66A3855D2703FFD1F53EF900FC0CF930" box="[790,931,1733,1759]" italics="true" pageId="4" pageNumber="395">F. formicarii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="1CCD0AC42703FFD0F6EFF92DFDA7FE88" lastPageId="5" lastPageNumber="396" pageId="4" pageNumber="395" type="etymology">
<paragraph id="5468594F2703FFD1F6EFF92DFD8BF8C9" blockId="4.[151,1437,153,2046]" pageId="4" pageNumber="395">
<emphasis id="66A3855D2703FFD1F6EFF92DFEE2F8ED" bold="true" box="[199,333,1768,1794]" pageId="4" pageNumber="395">Etymology.</emphasis>
We have named this new species after the genus of the host plant (
<taxonomicName id="93D722CC2703FFD1F235F92CFBDCF8ED" box="[1053,1139,1769,1794]" class="Magnoliopsida" family="Lamiaceae" genus="Tectona" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="395" phylum="Tracheophyta" rank="genus">
<emphasis id="66A3855D2703FFD1F235F92CFBDCF8ED" box="[1053,1139,1769,1794]" italics="true" pageId="4" pageNumber="395">Tectona</emphasis>
</taxonomicName>
) on which it was collected; the name is in the genitive singular.
</paragraph>
<paragraph id="5468594F2703FFD0F6EFF8EAFE83FF38" blockId="4.[151,1437,153,2046]" lastBlockId="5.[151,1436,153,359]" lastPageId="5" lastPageNumber="396" pageId="4" pageNumber="395">
<emphasis id="66A3855D2703FFD1F6EFF8EAFE26F8A5" bold="true" box="[199,393,1839,1866]" pageId="4" pageNumber="395">Biological notes.</emphasis>
At the first location (
<figureCitation id="CCEC45CA2703FFD1F450F8EAFD7FF8A5" box="[632,720,1839,1866]" captionStart="FIGURE 1" captionStartId="5.[151,250,1860,1885]" captionTargetBox="[160,1427,373,1840]" captionTargetId="figure@5.[160,1427,373,1840]" captionTargetPageId="5" captionText="FIGURE 1. A. Site of observations in the Palappalli forest range; B. Teak tree affected with trunk borer, Cossus cadambae; C. Entry holes made by the larvae of trunk borer; D. Infestation of Formicococcus tectonae Joshi, Bindu &amp; Gullan at the entry hole of the borer and an ant attending the mealybugs; E. Eggs of F. tectonae; F. Nymph hidden under the loose bark; G. Adult female." figureDoi="http://doi.org/10.5281/zenodo.3602552" httpUri="https://zenodo.org/record/3602552/files/figure.png" pageId="4" pageNumber="395">Fig. 1A</figureCitation>
), 36 of 45 trees (80.0 %) were infested by trunk borer to various degrees, whereas at the second location 40 of 55 trees (72.7 %) were infested. The trunks of borer-infested trees had a twisted appearance (
<figureCitation id="CCEC45CA2703FFD1F7A2F8B2FE48F87D" box="[394,487,1911,1938]" captionStart="FIGURE 1" captionStartId="5.[151,250,1860,1885]" captionTargetBox="[160,1427,373,1840]" captionTargetId="figure@5.[160,1427,373,1840]" captionTargetPageId="5" captionText="FIGURE 1. A. Site of observations in the Palappalli forest range; B. Teak tree affected with trunk borer, Cossus cadambae; C. Entry holes made by the larvae of trunk borer; D. Infestation of Formicococcus tectonae Joshi, Bindu &amp; Gullan at the entry hole of the borer and an ant attending the mealybugs; E. Eggs of F. tectonae; F. Nymph hidden under the loose bark; G. Adult female." figureDoi="http://doi.org/10.5281/zenodo.3602552" httpUri="https://zenodo.org/record/3602552/files/figure.png" pageId="4" pageNumber="395">Fig. 1 B</figureCitation>
). Each moth larva was found to make four to five entry holes in the trunk and produce tunnels while feeding on the heartwood (
<figureCitation id="CCEC45CA2703FFD1F46AF85EFD3CF85A" box="[578,659,1947,1973]" captionStart="FIGURE 1" captionStartId="5.[151,250,1860,1885]" captionTargetBox="[160,1427,373,1840]" captionTargetId="figure@5.[160,1427,373,1840]" captionTargetPageId="5" captionText="FIGURE 1. A. Site of observations in the Palappalli forest range; B. Teak tree affected with trunk borer, Cossus cadambae; C. Entry holes made by the larvae of trunk borer; D. Infestation of Formicococcus tectonae Joshi, Bindu &amp; Gullan at the entry hole of the borer and an ant attending the mealybugs; E. Eggs of F. tectonae; F. Nymph hidden under the loose bark; G. Adult female." figureDoi="http://doi.org/10.5281/zenodo.3602552" httpUri="https://zenodo.org/record/3602552/files/figure.png" pageId="4" pageNumber="395">Fig. 1C</figureCitation>
). At both locations, all the borer-infested trees had mealybug populations. Mealybugs were found only at the entrance of the passageways made by the moth larvae. It is doubtful if there is any reciprocal relationship or even a dependency of the mealybug on the moth larvae, although the tunnels made by the borer provide suitable habitat for the mealybugs. Further studies are needed to investigate the association of these insects.
</paragraph>
<paragraph id="5468594F2702FFD0F6EFFF24FDA7FE88" blockId="5.[151,1436,153,359]" pageId="5" pageNumber="396">
No parasitoids or predators of the mealybugs were obtained from the collections made in the present study. In the field, ants of an unidentified species of 
<taxonomicName id="93D722CC2702FFD0F442FEC3FC9DFEF0" authority="Foerster" authorityName="Foerster" box="[618,818,261,287]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="66A3855D2702FFD0F442FEC3FD7AFEF0" box="[618,725,262,287]" italics="true" pageId="5" pageNumber="396">Tapinoma</emphasis>
Foerster
</taxonomicName>
were found attending the mealybugs. In the first location, ants were found on 12 out of 36 mealybug-infested trees (33.3 %), while at second location, ants were found on 9 out of 40 infested trees (22.5 %).
</paragraph>
</subSubSection>
</treatment>
</document>