271 lines
23 KiB
XML
271 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1066.69697" ID-GBIF-Dataset="89be094e-6732-48a6-b5aa-74053d16f748" ID-GBIF-Taxon="189599992" ID-Pensoft-Pub="1313-2970-1066-1" ID-Pensoft-UUID="BD84F4C3157550C9B64120B2BE53F01A" ID-ZooBank="133CE040A5AF44F1BC9A558C2F06A8AA" LSID="urn:lsid:zoobank.org:act:5020B866-48D5-4686-8BDA-7769397C7D5A" ModsDocID="1313-2970-1066-1" checkinTime="1635480455043" checkinUser="pensoft" docAuthor="Filander, Zoleka N., Kitahara, Marcelo V., Cairns, Stephen D., Sink, Kerry J. & Lombard, Amanda T." docDate="2021" docId="508D6D3691F1EBE1108FF6E86F430C2E" docLanguage="en" docName="ZooKeys 1066: 1-198" docOrigin="ZooKeys 1066" docPubDate="2021-10-28" docSource="http://dx.doi.org/10.3897/zookeys.1066.69697" docTitle="Heterocyathus monileseptatum Filander & Kitahara 2021, sp. nov." docType="treatment" docUuid="5020B866-48D5-4686-8BDA-7769397C7D5A" docUuidSource="ZooBank" docVersion="3" id="BD84F4C3157550C9B64120B2BE53F01A" lastPageNumber="1" masterDocId="BD84F4C3157550C9B64120B2BE53F01A" masterDocTitle="Azooxanthellate Scleractinia (Cnidaria, Anthozoa) from South Africa" masterLastPageNumber="198" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1635513010010" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Azooxanthellate Scleractinia (Cnidaria, Anthozoa) from South Africa</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Filander, Zoleka N.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6905-4440</mods:nameIdentifier>
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<mods:affiliation>Biodiversity and Coastal Research, Oceans and Coasts, Department of Environment, Forestry, and Fisheries, Cape Town, South Africa & Zoology Department, Nelson Mandela University, Port Elizabeth, South Africa</mods:affiliation>
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<mods:nameIdentifier type="email">zfilander@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kitahara, Marcelo V.</mods:namePart>
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<mods:affiliation>Universidade Federal de Sao Paulo, Departamento de Ciencias do Mar, Santos, Brazil & Centro de Biologia Marinha, Universidade de Sao Paulo, Sao Sebastiao, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cairns, Stephen D.</mods:namePart>
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<mods:affiliation>Department of Invertebrate Zoology, Smithsonian Institution, Washington DC, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sink, Kerry J.</mods:namePart>
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<mods:affiliation>South African National Biodiversity Institute, Cape Town, South Africa & Institute for Coastal and Marine Research, Nelson Mandela University, Port Elizabeth, South Africa</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lombard, Amanda T.</mods:namePart>
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<mods:affiliation>Institute for Coastal and Marine Research, Nelson Mandela University, Port Elizabeth, South Africa</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-10-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1066</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>198</mods:end>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1066.69697</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1066.69697</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1066-1</mods:identifier>
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<mods:identifier type="ZooBank">133CE040A5AF44F1BC9A558C2F06A8AA</mods:identifier>
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<mods:identifier type="Pensoft-UUID">BD84F4C3157550C9B64120B2BE53F01A</mods:identifier>
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<treatment ID-GBIF-Taxon="189599992" LSID="urn:lsid:zoobank.org:act:5020B866-48D5-4686-8BDA-7769397C7D5A" httpUri="http://treatment.plazi.org/id/508D6D3691F1EBE1108FF6E86F430C2E" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/5020B866-48D5-4686-8BDA-7769397C7D5A" authority="Filander & Kitahara" authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum" status="sp. nov.">Heterocyathus monileseptatum Filander & Kitahara</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, B Heterocyathus alternatus (ORI _ DIIIe 1 _ 1, locality data unknown) A calicular view B lateral view C, D Heterocyathus monileseptatum sp. nov. (SAM _ H 1431, off Durban Harbour, 99 m) C calicular view D lateral view E, F Heterocyathus sulcatus (SAMC _ A 073123, off Shaka's Rock, 100 - 105 m) E calicular view F lateral view G, H Labyrinthocyathus delicatus (SAM _ H 2836, off East London, 146 - 238 m) G calicular view H lateral view I, K Monohedotrochus capensis comb. nov. I, J (SAMC _ A 088924, off Kidds Beach, 247 - 147 m) I calicular view J lateral view K (SAM _ H 3210, off Scottburgh, 690 m) calicular view L, M Polycyathus sp. (USNM 91677, off Port Dunford, 69 m) L calicular view M full view N, O Rhizosmilia robusta (USNM 91689, off Kosi Bay Estuary, 74 m) N calicular view O lateral view P Solenosmilia variabilis (SAM _ H 3158, off Cintsa, 630 m) P full view. Scale bars: 10 mm (A-G, I-O); 100 mm (H, P)." figureDoi="10.3897/zookeys.1066.69697.figure4" httpUri="https://binary.pensoft.net/fig/604107" pageId="0" pageNumber="1">Fig. 4C, D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">Off Durban Harbour, South Africa, 99 m.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material/.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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SAM_H1431A: eastern margin, 5 km from Durban/7 km off Umgeni Estuary, (RV 'Pieter
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<normalizedToken originalValue="Faure’">Faure'</normalizedToken>
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:
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<geoCoordinate degrees="29" direction="south" minutes="52" orientation="latitude" precision="1" seconds="00.00" value="-29.866667">29°52'00.00"S</geoCoordinate>
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,
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<geoCoordinate degrees="31" direction="east" minutes="03" orientation="longitude" precision="1" seconds="00.00" value="31.05">31°03'00.00"E</geoCoordinate>
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); 99 m.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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SAM_H1431B (4 specimens): eastern margin, 5 km from Durban/7 km off Umgeni Estuary, (RV 'Pieter
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<normalizedToken originalValue="Faure’">Faure'</normalizedToken>
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:
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<geoCoordinate degrees="29" direction="south" minutes="52" orientation="latitude" precision="1" seconds="00.00" value="-29.866667">29°52'00.00"S</geoCoordinate>
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,
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<geoCoordinate degrees="31" direction="east" minutes="03" orientation="longitude" precision="1" seconds="00.00" value="31.05">31°03'00.00"E</geoCoordinate>
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); 99 m. SAM_H1246 (8 specimens): Locality data unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The species name
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<taxonomicName authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum">
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<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus monileseptatum</emphasis>
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</taxonomicName>
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(derived from Latin
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<emphasis italics="true" pageId="0" pageNumber="1">monile</emphasis>
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meaning "string of beads" and
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<emphasis italics="true" pageId="0" pageNumber="1">septum</emphasis>
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meaning "enclosure, wall, fence") which alludes to the beaded septal margins.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Corallum unattached and tall, with lateral efferent pores ranging in diameter from 1.00 ≤ 2.00 mm. Base flat, but a prominent basal pore results in an asymmetric corallum with an irregularly shaped base (BD = 1.1-1.6). Calice circular to elliptical (GCD:LCD = 1.0-1.1), with serrate calicular margin. Holotype (SAM_H1431) 11.0
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<normalizedToken originalValue="×">x</normalizedToken>
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9.7 mm in CD, 11.1
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<normalizedToken originalValue="×">x</normalizedToken>
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6.9 mm in BD, and 10.6 mm in H. Paratypes having one or two aboral pores randomly positioned. Costae equidistant and progressively decreasing in width towards base. All costae finely granulated. C4 bears distinctive low spine-like granules. At base each costae become a row of granules. Intercostal striae equal in width and depth. Corallum white, with theca and columella with blackish pigment.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Septa thin, spaced out, delicate, and hexamerally arranged in four complete cycles (which follow a
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<normalizedToken originalValue="Pourtalès">Pourtales</normalizedToken>
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plan) according to the formula: S1 ≥ S2> S4> S3 (48 septa). S1 most exsert and extend to columella with straight and smooth axial margins. S2 only slight less exsert and ca. as wide as S1. S2 axial margins slightly sinuous. S3 least exsert septa and 2/3 the size of S2. In each half-system, a pair of S4 joins common S3 deep in fossa, and extends towards columella as one septum. S3-4 junctions beaded. S4 more exsert than S3, and also with sinuous axial margin. S4 dimorphic in size: those adjacent to S1 being wider and more exsert than those adjacent to S2. At calicular margin, S4 fuses to adjacent S1-2 forming high rectangular lancets. Septal faces granular. No pali. Fossa deep, containing a poorly developed papillose columella composed of ≤ seven sparsely arranged rods.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Regional: Eastern margin of South Africa, off Durban; 99 m.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Amongst the six extant
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<taxonomicName authorityName="Milne-Edwards & Haime" authorityYear="1848" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus</emphasis>
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</taxonomicName>
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species (
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<taxonomicName genus="H." lsidName="H. aequicostatus" pageId="0" pageNumber="1" rank="species" species="aequicostatus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. aequicostatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName genus="H." lsidName="H. alternatus" pageId="0" pageNumber="1" rank="species" species="alternatus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. alternatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName genus="H." lsidName="H. antoniae" pageId="0" pageNumber="1" rank="species" species="antoniae">
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<emphasis italics="true" pageId="0" pageNumber="1">H. antoniae</emphasis>
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</taxonomicName>
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Reyes, Santodomingo & Cairns, 2009,
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<taxonomicName genus="H." lsidName="H. hemisphaericus" pageId="0" pageNumber="1" rank="species" species="hemisphaericus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. hemisphaericus</emphasis>
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</taxonomicName>
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<bibRefCitation author="Gray, J" journalOrPublisher="Proceedings Zoological Society of London" pageId="0" pageNumber="1" pagination="74 - 77" refId="B116" refString="Gray, J, 1849. Description of some corals, including a new British coral discovered by W. Mac-Andrew, Esq. Proceedings Zoological Society of London 17: 74 - 77" title="Description of some corals, including a new British coral discovered by W. Mac-Andrew, Esq." volume="17" year="1849">Gray 1849</bibRefCitation>
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,
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<taxonomicName genus="H." lsidName="H. japanicus" pageId="0" pageNumber="1" rank="species" species="japanicus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. japanicus</emphasis>
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</taxonomicName>
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(Verrill, 1866), and
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<taxonomicName genus="H." lsidName="H. sulcatus" pageId="0" pageNumber="1" rank="species" species="sulcatus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. sulcatus</emphasis>
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</taxonomicName>
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) (
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<bibRefCitation author="Hoeksema, B" journalOrPublisher="Coral Reefs" pageId="0" pageNumber="1" refId="B129" refString="Hoeksema, B, Cairns, S, 2021. World List of Scleractinia. Heterocyathus Milne Edwards & Haime, 1848. Accessed through: World Register of Marine Species on 2021-07-16. http://www.marinespecies.org/aphia.php?p=taxdetails&id=206380" title="World List of Scleractinia. Heterocyathus Milne Edwards & Haime, 1848. Accessed through: World Register of Marine Species on 2021 - 07 - 16. http: // www. marinespecies. org / aphia. php? p = taxdetails & id = 206380" year="2021">Hoeksema and Cairns 2021</bibRefCitation>
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), the specimens reported herein are distinctive in the lack of pali, height of corallum, and by having a beaded axial margin at the S3-4 junctions. However, there are intermediate similarities with each of these species, for example
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<taxonomicName authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum">
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<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus monileseptatum</emphasis>
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</taxonomicName>
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sp. nov. resembles
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<taxonomicName genus="H." lsidName="H. aequicostatus" pageId="0" pageNumber="1" rank="species" species="aequicostatus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. aequicostatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName genus="H." lsidName="H. alternatus" pageId="0" pageNumber="1" rank="species" species="alternatus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. alternatus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName genus="H." lsidName="H. hemisphaericus" pageId="0" pageNumber="1" rank="species" species="hemisphaericus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. hemisphaericus</emphasis>
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</taxonomicName>
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in having septa arranged in four cycles (S1> S2> S4> S3 = 48 septa), but differs from the first two species in having no pali rather than P1-4 (in case of
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<taxonomicName genus="H." lsidName="H. aequicostatus" pageId="0" pageNumber="1" rank="species" species="aequicostatus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. aequicostatus</emphasis>
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</taxonomicName>
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) or P1, 3, 4 (in case of
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<taxonomicName genus="H." lsidName="H. alternatus" pageId="0" pageNumber="1" rank="species" species="alternatus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. alternatus</emphasis>
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</taxonomicName>
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). On the contrary, the lack of pali in
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<taxonomicName authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum">
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<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus monileseptatum</emphasis>
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</taxonomicName>
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sp. nov. is a similarity shared with
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<taxonomicName genus="H." lsidName="H. hemisphaericus" pageId="0" pageNumber="1" rank="species" species="hemisphaericus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. hemisphaericus</emphasis>
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</taxonomicName>
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but this species can be distinguished by their septa profile: all septa are thin (max 0.6 mm) and solid in
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<taxonomicName authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum">
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<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus monileseptatum</emphasis>
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</taxonomicName>
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sp. nov. compared with being thick (max 0.9 mm) and porous in
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<taxonomicName genus="H." lsidName="H. hemisphaericus" pageId="0" pageNumber="1" rank="species" species="hemisphaericus">
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<emphasis italics="true" pageId="0" pageNumber="1">H. hemisphaericus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation author="Cairns, S" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="1" pagination="361 - 417" refId="B40" refString="Cairns, S, 1998. Azooxanthellate Scleractinia (Cnidaria: Anthozoa) of western Australia. Records of the Western Australian Museum 18: 361 - 417" title="Azooxanthellate Scleractinia (Cnidaria: Anthozoa) of western Australia." volume="18" year="1998">Cairns 1998</bibRefCitation>
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). The maximum height observed in
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<taxonomicName authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum">
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<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus monileseptatum</emphasis>
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</taxonomicName>
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sp. nov. (10.9 mm) is higher than that reported in the Atlantic
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<taxonomicName genus="H." lsidName="H. antoniae" pageId="0" pageNumber="1" rank="species" species="antoniae">
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<emphasis italics="true" pageId="0" pageNumber="1">H. antoniae</emphasis>
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</taxonomicName>
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(9.0 mm) and both species also have septa arranged in a
|
||
<normalizedToken originalValue="Pourtalès">Pourtales</normalizedToken>
|
||
plan fashion. Nonetheless, differences between
|
||
<taxonomicName authorityName="Milne-Edwards & Haime" authorityYear="1848" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName genus="H." lsidName="H. antoniae" pageId="0" pageNumber="1" rank="species" species="antoniae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">H. antoniae</emphasis>
|
||
</taxonomicName>
|
||
include (i) septa being arranged according to S1> S2> S4> S3 and S1 = S2 = S4 >> S3, respectively; (ii) lack of pali in
|
||
<taxonomicName authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus monileseptatum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; (iii) columella of
|
||
<taxonomicName authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus monileseptatum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. being papillose and composed of sparsely arranged rods versus a spongy columella composed of crispate elements in
|
||
<taxonomicName genus="H." lsidName="H. antoniae" pageId="0" pageNumber="1" rank="species" species="antoniae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">H. antoniae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus japanicus" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="japanicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus japanicus</emphasis>
|
||
</taxonomicName>
|
||
is also reported to have septa arranged in a
|
||
<normalizedToken originalValue="Pourtalès">Pourtales</normalizedToken>
|
||
plan (Zibrowuis 1997). However,
|
||
<taxonomicName authorityName="Filander & Kitahara" authorityYear="2021" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus monileseptatum" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="monileseptatum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus monileseptatum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be differentiated by beaded S3-4 axial margins, a feature that is unique in relation to all
|
||
<taxonomicName authorityName="Milne-Edwards & Haime" authorityYear="1848" class="Anthozoa" family="Dendrophylliidae" genus="Heterocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterocyathus" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Heterocyathus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |