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<document id="4B8510A9E4E495BADD15F1E5274865C1" ID-CLB-Dataset="30682" ID-DOI="10.11646/zootaxa.4382.3.4" ID-GBIF-Dataset="b88bfa2d-4b83-41f8-a145-a95ca196960f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1183130" ID-ZooBank="87A2B333-09FF-4726-A1EF-0699FB631808" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1519301947700" checkinUser="plazi" docAuthor="Machado, Miguel, Teixeira, Renato Augusto &amp; Lise, Arno Antonio" docDate="2018" docId="501D87B16738A43A2390FE8A5429BBDD" docLanguage="en" docName="zootaxa.4382.3.4.pdf" docOrigin="Zootaxa 4382 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epicadus camelinus" docType="treatment" docVersion="8" lastPageNumber="507" masterDocId="AC24FFC9673DA43C2307FFF85679BF26" masterDocTitle="There and back again: More on the taxonomy of the crab spiders genus Epicadus (Thomisidae: Stephanopinae)" masterLastPageNumber="530" masterPageNumber="501" pageNumber="506" updateTime="1698524041822" updateUser="ExternalLinkService">
<mods:mods id="81EAE7FC7969CC9ACFFC8B62004C5666" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3AC6F6C4BDAEAB31DFB8134FDF3571E7">
<mods:title id="5955E518A789371920BFAEEE7116CC41">There and back again: More on the taxonomy of the crab spiders genus Epicadus (Thomisidae: Stephanopinae)</mods:title>
</mods:titleInfo>
<mods:name id="45EA1DBE46CD20FA9DC81256D2147214" type="personal">
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<mods:namePart id="5CF8BD9A8FE151086D17DCF729D5C4CD">Machado, Miguel</mods:namePart>
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<mods:name id="7C3E630F2FC267583FC4D4495391B7F3" type="personal">
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<mods:namePart id="AE07936407DD459E317E6D82C322784E">Teixeira, Renato Augusto</mods:namePart>
</mods:name>
<mods:name id="A53387134915AB4C62CAB05BCD028566" type="personal">
<mods:role id="99F336B2E0B260A74B8045BAA88BBAD8">
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<mods:namePart id="D70014DE5BE2D9B36D9C3C1A68FC4DF6">Lise, Arno Antonio</mods:namePart>
</mods:name>
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<mods:title id="0EA505EDC5ECA37AA7C7850AB4D86858">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0FAA4ECE0FB7337E1390152B6BF4923B">
<mods:date id="5048D5CC5CB59B298FA3313AE5086EAA">2018</mods:date>
<mods:detail id="AC44540EE7333BCAF929D967011C204A" type="pubDate">
<mods:number id="3814AA9D9195CF0A6F47E4EE1E944E03">2018-02-22</mods:number>
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<mods:number id="63981FA96E179105F29C1B9F1D04AB2A">4382</mods:number>
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<mods:number id="D0D16762D6751E45B7E459458FA4BD71">3</mods:number>
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<mods:classification id="0C9D5DD5E23303EDB280E0D2BE4770A3">journal article</mods:classification>
<mods:identifier id="8B571D086A4502A2220E9BF73A00E650" type="CLB-Dataset">30682</mods:identifier>
<mods:identifier id="2A0CF42FDA4B7248D0C74FC63E383BD3" type="DOI">10.11646/zootaxa.4382.3.4</mods:identifier>
<mods:identifier id="37989D59BDA43395545FDC5B416F83E8" type="GBIF-Dataset">b88bfa2d-4b83-41f8-a145-a95ca196960f</mods:identifier>
<mods:identifier id="50E73073526916D16C5A3A7B0B9A601A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="501D87B16738A43A2390FE8A5429BBDD" ID-DOI="http://doi.org/10.5281/zenodo.5951302" ID-GBIF-Taxon="141118935" ID-Zenodo-Dep="5951302" LSID="urn:lsid:plazi:treatment:501D87B16738A43A2390FE8A5429BBDD" httpUri="http://treatment.plazi.org/id/501D87B16738A43A2390FE8A5429BBDD" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506">
<subSubSection id="90AE652C6738A4392390FE8A5728BE88" pageId="5" pageNumber="506" type="nomenclature">
<paragraph id="D80B36A76738A4392390FE8A5577BEAA" blockId="5.[151,782,370,430]" box="[151,782,370,396]" pageId="5" pageNumber="506">
<heading id="834381CB6738A4392390FE8A5577BEAA" bold="true" box="[151,782,370,396]" fontSize="11" level="1" pageId="5" pageNumber="506" reason="1">
<taxonomicName id="1FB44D246738A4392390FE8A5577BEAA" ID-CoL="6G3CH" authority="(O. Pickard-Cambridge, 1869)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[151,782,370,396]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56738A4392390FE8A5577BEAA" bold="true" box="[151,782,370,396]" pageId="5" pageNumber="506">
<emphasis id="EAC0EAB56738A4392390FE8A57F3BEAA" bold="true" box="[151,394,370,396]" italics="true" pageId="5" pageNumber="506">Epicadus camelinus</emphasis>
(
<bibRefCitation id="BC254B566738A439229DFE8A557FBEAA" box="[410,774,370,396]" pageId="5" pageNumber="506" refId="ref19322" refString="Pickard-Cambridge, O. (1869) Descriptions and sketches of some neW species of Araneida With characters of a neW genus. Annals and Magazine of Natural History, 3, 52 - 74. https: // doi. org / 10.1080 / 00222936908695878" type="journal article">O. Pickard-Cambridge, 1869</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D80B36A76738A4392390FE6D5728BE88" blockId="5.[151,782,370,430]" box="[151,337,405,430]" pageId="5" pageNumber="506">
<figureCitation id="408F2A226738A4392390FE6D576FBE88" box="[151,278,405,430]" captionStart="FIGURE 2" captionStartId="5.[151,250,1855,1877]" captionTargetBox="[151,1436,674,1833]" captionTargetId="figure@5.[151,1436,674,1833]" captionTargetPageId="5" captionText="FIGURE 2. Epicadus camelinus (O. Pickard-Cambridge, 1869), female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A, B), 0.2 mm(C, D)." httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="5" pageNumber="506">Figs 2A–D</figureCitation>
,
<figureCitation id="408F2A226738A4392227FE6D5728BE88" box="[288,337,405,430]" captionStart="FIGURE 16" captionStartId="26.[151,250,1869,1891]" captionTargetBox="[184,1408,193,1859]" captionTargetId="figure@26.[184,1408,193,1859]" captionTargetPageId="26" captionText="FIGURE 16. Female specimens in vivo. A Epicadus taczanowskii (RoeWer, 1951); B Epicadus dimidiaster sp. nov.; C Epicadus rubripes Mello-Leitão, 1924 (adapted from Silva-Moreira &amp; Machado 2016: fig. 1F); D Epicadus camelinus (O. Pickard-Cambridge, 1869); E Epicadus heterogaster (Guérin-Méneville, 1829); F Epicadus trituberculatus (TaczanoWski, 1872). Photo credits:César Favacho (A); Arthur Anker (B); Almir Cândido de Almeida (C); Karl Kroeker (D); Edson Roberto (E); Edelcio Muscat (F)." httpUri="https://zenodo.org/record/1183162/files/figure.png" pageId="5" pageNumber="506">16D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6738A4392390FE2552EFBD48" pageId="5" pageNumber="506" type="reference_group">
<paragraph id="D80B36A76738A4392390FE255506BED5" blockId="5.[151,1436,477,622]" box="[151,895,477,499]" pageId="5" pageNumber="506">
<treatmentCitationGroup id="F8A411896738A4392390FE255506BED5" box="[151,895,477,499]" pageId="5" pageNumber="506">
<taxonomicName id="1FB44D246738A4392390FE2554B0BED5" authority="O. Pickard-Cambridge, 1869: 64" authorityName="O. Pickard-Cambridge" authorityPageNumber="64" authorityYear="1869" box="[151,713,477,499]" class="Arachnida" family="Thomisidae" genus="Stephanopis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="camelina">
<emphasis id="EAC0EAB56738A4392390FE25570DBED5" box="[151,372,477,499]" italics="true" pageId="5" pageNumber="506">Stephanopis camelina</emphasis>
<treatmentCitation id="591510B66738A439227CFE2554B0BED5" author="" box="[379,713,477,499]" page="64" pageId="5" pageNumber="506">
<bibRefCitation id="BC254B566738A439227CFE2554DDBED5" box="[379,676,477,499]" pageId="5" pageNumber="506" refId="ref19322" refString="Pickard-Cambridge, O. (1869) Descriptions and sketches of some neW species of Araneida With characters of a neW genus. Annals and Magazine of Natural History, 3, 52 - 74. https: // doi. org / 10.1080 / 00222936908695878" type="journal article">O. Pickard-Cambridge, 1869</bibRefCitation>
: 64
</treatmentCitation>
</taxonomicName>
, pl. 5, figs 44–49.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D80B36A76738A4392390FE045568BD34" blockId="5.[151,1436,477,622]" box="[151,785,508,530]" pageId="5" pageNumber="506">
<treatmentCitationGroup id="F8A411896738A4392390FE045568BD34" box="[151,785,508,530]" pageId="5" pageNumber="506">
<taxonomicName id="1FB44D246738A4392390FE04543FBD34" ID-CoL="93RB4" authority="(O. Pickard-Cambridge)" authorityName="O. Pickard-Cambridge" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[151,582,508,530]" class="Arachnida" family="Thomisidae" genus="Tobias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56738A4392390FE04573EBD34" box="[151,327,508,530]" italics="true" pageId="5" pageNumber="506">Tobias camelinus</emphasis>
(O. Pickard-Cambridge)
</taxonomicName>
.
<treatmentCitation id="591510B66738A4392148FE045574BD34" author="" box="[591,781,508,530]" page="1053" pageId="5" pageNumber="506">
<bibRefCitation id="BC254B566738A4392148FE0454B4BD34" box="[591,717,508,530]" pageId="5" pageNumber="506" refId="ref19502" refString="Simon, E. (1895) Histoire naturelle des araignees. Librairie Encyclopedique de Roret, Paris, 1084 pp. https: // doi. org / 10.5962 / bhl. title. 47654" type="book">Simon 1895</bibRefCitation>
: 1053
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D80B36A76738A4392390FDE254DABD69" blockId="5.[151,1436,477,622]" pageId="5" pageNumber="506">
<materialsCitation id="68DC3CFA6738A4392390FDE254DABD69" ID-GBIF-Occurrence="1820860248" collectionCode="MLPA" country="Bolivia" latitude="-17.458889" location="Buenavista" longLatPrecision="21" longitude="-63.65917" pageId="5" pageNumber="506" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<taxonomicName id="1FB44D246738A4392390FDE2546EBD16" authority="Biraben, 1955: 74" authorityName="Biraben" authorityPageNumber="74" authorityYear="1955" box="[151,535,538,560]" class="Arachnida" family="Thomisidae" genus="Tobias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="EAC0EAB56738A4392390FDE2573FBD16" box="[151,326,538,560]" italics="true" pageId="5" pageNumber="506">Tobias martinezi</emphasis>
Birabén, 1955: 74
</taxonomicName>
, figs 1–4 (
<typeStatus id="070F88056738A4392195FDE25494BD16" box="[658,749,538,560]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
female from
<location id="DD6B607C6738A439208FFDE25581BD16" LSID="urn:lsid:plazi:treatment:501D87B16738A43A2390FE8A5429BBDD:DD6B607C6738A439208FFDE25581BD16" box="[904,1016,538,560]" country="Bolivia" latitude="-17.458889" longLatPrecision="21" longitude="-63.65917" name="Buenavista" pageId="5" pageNumber="506" stateProvince="Santa Cruz">Buenavista</location>
[
<geoCoordinate id="BD8050606738A439270CFDE252FBBD16" box="[1035,1154,538,560]" direction="south" orientation="latitude" pageId="5" pageNumber="506" precision="15" value="-17.458889">17°27'32&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606738A4392795FDE25376BD16" box="[1170,1295,538,560]" direction="west" orientation="longitude" pageId="5" pageNumber="506" precision="15" value="-63.65917">63°39'33&quot;W</geoCoordinate>
,
<collectingRegion id="1A70F8456738A4392627FDE253E1BD16" box="[1312,1432,538,560]" country="Bolivia" name="Santa Cruz" pageId="5" pageNumber="506">Santa Cruz</collectingRegion>
,
<collectingCountry id="A0A376376738A43923C0FDC1576FBD69" box="[199,278,569,591]" name="Bolivia" pageId="5" pageNumber="506">Bolivia</collectingCountry>
], deposited in
<collectionCode id="BEA5AE626738A43922AFFDC25795BD69" box="[424,492,570,591]" pageId="5" pageNumber="506">MLPA</collectionCode>
0 400, examined).
</materialsCitation>
</paragraph>
<paragraph id="D80B36A76738A4392390FDA052EFBD48" blockId="5.[151,1436,477,622]" box="[151,1174,600,622]" pageId="5" pageNumber="506">
<treatmentCitationGroup id="F8A411896738A4392390FDA052EFBD48" box="[151,1174,600,622]" pageId="5" pageNumber="506">
<taxonomicName id="1FB44D246738A4392390FDA054D8BD48" ID-CoL="6G3CH" authority="(O. Pickard-Cambridge, 1869)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[151,673,600,622]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56738A4392390FDA0571BBD48" box="[151,354,600,622]" italics="true" pageId="5" pageNumber="506">Epicadus camelinus</emphasis>
(
<bibRefCitation id="BC254B566738A4392276FDA054E1BD48" box="[369,664,600,622]" pageId="5" pageNumber="506" refId="ref19322" refString="Pickard-Cambridge, O. (1869) Descriptions and sketches of some neW species of Araneida With characters of a neW genus. Annals and Magazine of Natural History, 3, 52 - 74. https: // doi. org / 10.1080 / 00222936908695878" type="journal article">O. Pickard-Cambridge, 1869</bibRefCitation>
)
</taxonomicName>
.
<treatmentCitation id="591510B66738A43921ADFDA055C9BD48" author="" box="[682,944,600,622]" page="448" pageId="5" pageNumber="506">
<bibRefCitation id="BC254B566738A43921ADFDA05507BD48" box="[682,894,600,622]" pageId="5" pageNumber="506" refId="ref19045" refString="Machado, M, Teixeira, R. A. &amp; Lise, A. A. (2017) Cladistic analysis supports the monophyly of the Neotropical crab spider genus Epicadus and its senior synonymy over Tobias (Araneae: Thomisidae). Invertebrate Systematics, 31, 442 - 455. https: // doi. org / 10.1071 / IS 16074" type="journal article">
Machado
<emphasis id="EAC0EAB56738A4392008FDA1553BBD48" box="[783,834,600,622]" italics="true" pageId="5" pageNumber="506">et al.</emphasis>
2017
</bibRefCitation>
: 448
</treatmentCitation>
, figs. S4C, S6B, S15C
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="8CCB662F6738A4392390F8C75372B855" httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="5" pageNumber="506" startId="5.[151,250,1855,1877]" targetBox="[151,1436,674,1833]" targetPageId="5">
<paragraph id="D80B36A76738A4392390F8C75372B855" blockId="5.[151,1436,1855,1907]" pageId="5" pageNumber="506">
<emphasis id="EAC0EAB56738A4392390F8C7576AB873" bold="true" box="[151,275,1855,1877]" pageId="5" pageNumber="506">FIGURE 2.</emphasis>
<taxonomicName id="1FB44D246738A439221DF8C7555DB873" authority="(O. Pickard-Cambridge, 1869)" authorityName="O. Pickard-Cambridge" authorityYear="1869" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[282,804,1855,1877]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56738A439221DF8C7579CB873" box="[282,485,1855,1877]" italics="true" pageId="5" pageNumber="506">Epicadus camelinus</emphasis>
(O. Pickard-Cambridge, 1869)
</taxonomicName>
, female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal). Scale bars: 2 mm (A, B), 0.2 mm (C, D).
</paragraph>
</caption>
<subSubSection id="90AE652C6738A43A2390F86357CCBFD1" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506" type="materials_examined">
<paragraph id="D80B36A76738A4392390F863521FB892" blockId="5.[151,1436,1947,2007]" box="[151,1126,1947,1972]" pageId="5" pageNumber="506">
<emphasis id="EAC0EAB56738A4392390F86357BDB892" bold="true" box="[151,452,1947,1972]" pageId="5" pageNumber="506">Type material: Holotype:</emphasis>
female, “Amazon”, 1861, Bates (OUMNH 122, examined).
</paragraph>
<paragraph id="D80B36A76738A43A23C0F84657CCBFD1" blockId="5.[151,1436,1947,2007]" lastBlockId="6.[151,1437,151,1276]" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506">
<materialsCitation id="68DC3CFA6738A43923C0F8465362B8F1" ID-GBIF-Occurrence="1820860272" box="[199,1307,1982,2007]" collectionCode="MNHN" country="Brazil" latitude="-3.4166667" location="Other" longLatPrecision="21" longitude="-65.855835" pageId="5" pageNumber="506" specimenCode="MNHN 2617" specimenCount="5" specimenCount-female="5" stateProvince="Amazonas">
<emphasis id="EAC0EAB56738A43923C0F8465484B8F1" bold="true" box="[199,765,1982,2007]" pageId="5" pageNumber="506">
<location id="DD6B607C6738A43923C0F8465776B8F1" LSID="urn:lsid:plazi:treatment:501D87B16738A43A2390FE8A5429BBDD:DD6B607C6738A43923C0F8465776B8F1" box="[199,271,1982,2007]" country="Brazil" latitude="-3.4166667" longLatPrecision="21" longitude="-65.855835" name="Other" pageId="5" pageNumber="506" stateProvince="Amazonas">Other</location>
material examined.
<collectingCountry id="A0A376376738A4392106F8465409B8F1" box="[513,624,1982,2007]" name="Brazil" pageId="5" pageNumber="506">BRAZIL</collectingCountry>
:
<collectingRegion id="1A70F8456738A439217BF846548DB8F1" box="[636,756,1982,2007]" country="Brazil" name="Amazonas" pageId="5" pageNumber="506">
<emphasis id="EAC0EAB56738A439217BF846548DB8F1" bold="true" box="[636,756,1982,2007]" italics="true" pageId="5" pageNumber="506">Amazonas</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6738A4392003F8465557B8F1" box="[772,814,1982,2007]" pageId="5" pageNumber="506" type="female">5 ♀</specimenCount>
,
<geoCoordinate id="BD8050606738A439203CF84655D7B8F1" box="[827,942,1982,2007]" direction="south" orientation="latitude" pageId="5" pageNumber="506" precision="15" value="-3.4166667">3°25'00&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606738A43920BDF846523DB8F1" box="[954,1092,1982,2007]" direction="west" orientation="longitude" pageId="5" pageNumber="506" precision="15" value="-65.855835">65°51'21&quot;W</geoCoordinate>
, (
<specimenCode id="88129EDC6738A4392750F847528AB8F1" box="[1111,1267,1982,2007]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="506">MNHN 2617</specimenCode>
ex.)
</materialsCitation>
.
<materialsCitation id="68DC3CFA6738A43A2622F84653E0BF96" ID-GBIF-Occurrence="1820860309" collectingDate="2010-11" collectionCode="UFMG" collectorName="G. H. F. Azevedo &amp; A. J. Santos" country="Brazil" lastPageId="6" lastPageNumber="507" latitude="-10.073334" location="Senador Guiomard" longLatPrecision="21" longitude="-67.62389" pageId="5" pageNumber="506" specimenCode="UFMG 11007" specimenCount="1" specimenCount-female="1" stateProvince="Acre">
<emphasis id="EAC0EAB56738A4392622F846531DB8F1" bold="true" box="[1317,1380,1982,2007]" pageId="5" pageNumber="506">
<collectingRegion id="1A70F8456738A4392622F8465322B8F1" box="[1317,1371,1982,2007]" country="Brazil" name="Acre" pageId="5" pageNumber="506">
<emphasis id="EAC0EAB56738A4392622F8465322B8F1" bold="true" box="[1317,1371,1982,2007]" italics="true" pageId="5" pageNumber="506">Acre</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6738A439266CF84753ECB8F1" box="[1387,1429,1983,2007]" pageId="5" pageNumber="506" type="female">1 ♀</specimenCount>
,
<location id="DD6B607C673BA43A2390FF6F5708BF96" LSID="urn:lsid:plazi:treatment:501D87B16738A43A2390FE8A5429BBDD:DD6B607C673BA43A2390FF6F5708BF96" box="[151,369,151,176]" country="Brazil" latitude="-10.073334" longLatPrecision="21" longitude="-67.62389" name="Senador Guiomard" pageId="6" pageNumber="507" stateProvince="Acre">Senador Guiomard</location>
,
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,
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,
<collectingDate id="BC4EE98F673BA43A21AFFF6F551DBF96" box="[680,868,151,176]" pageId="6" pageNumber="507" value="2010-11">
<date id="AC0A1067673BA43A21AFFF6F551DBF96" box="[680,868,151,176]" pageId="6" pageNumber="507" value="2010-11">November 2010</date>
</collectingDate>
,
<collectorName id="75415371673BA43A2074FF605250BF96" box="[883,1065,151,176]" pageId="6" pageNumber="507">G.H.F. Azevedo</collectorName>
&amp;
<collectorName id="75415371673BA43A2753FF6052AEBF96" box="[1108,1239,152,176]" pageId="6" pageNumber="507">A.J. Santos</collectorName>
(
<specimenCode id="88129EDC673BA43A27EEFF6053E8BF96" box="[1257,1425,152,176]" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="6" pageNumber="507">UFMG 11007</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="68DC3CFA673BA43A2390FF4357C8BFD1" ID-GBIF-Occurrence="1820860279" collectingDate="1991-10-12" collectingDateMax="1991-10-16" collectingDateMin="1991-10-12" collectionCode="MCNZ" collectorName="A. B. Bonaldo" country="Saint Helena, Ascension and Tristan da Cunha" latitude="-25.448889" location="Refugio Biologico Bela Vista" longLatPrecision="21" longitude="-54.556667" municipality="Parana" pageId="6" pageNumber="507" specimenCode="MCNZ 21767" specimenCount="1" specimenCount-female="1">
<emphasis id="EAC0EAB5673BA43A2390FF43568CBFF2" bold="true" box="[151,245,187,212]" pageId="6" pageNumber="507">
<collectingMunicipality id="386FACDD673BA43A2390FF435695BFF2" box="[151,236,187,212]" pageId="6" pageNumber="507">
<emphasis id="EAC0EAB5673BA43A2390FF435695BFF2" bold="true" box="[151,236,187,212]" italics="true" pageId="6" pageNumber="507">Paraná</emphasis>
</collectingMunicipality>
:
</emphasis>
<specimenCount id="CEB2FD2E673BA43A23F9FF445753BFF5" box="[254,298,188,212]" pageId="6" pageNumber="507" type="female">1 ♀</specimenCount>
,
<collectingCountry id="A0A37637673BA43A223EFF4457B6BFF2" box="[313,463,187,212]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="6" pageNumber="507">Santa Helena</collectingCountry>
(
<location id="DD6B607C673BA43A22D8FF43554CBFF2" LSID="urn:lsid:plazi:treatment:501D87B16738A43A2390FE8A5429BBDD:DD6B607C673BA43A22D8FF43554CBFF2" box="[479,821,187,212]" country="Saint Helena, Ascension and Tristan da Cunha" latitude="-25.448889" longLatPrecision="21" longitude="-54.556667" municipality="Parana" name="Refugio Biologico Bela Vista" pageId="6" pageNumber="507">Refúgio Biológico Bela Vista</location>
),
<geoCoordinate id="BD805060673BA43A204EFF4355B3BFF2" box="[841,970,187,212]" direction="south" orientation="latitude" pageId="6" pageNumber="507" precision="15" value="-25.448889">25°26'56&quot;S</geoCoordinate>
,
<geoCoordinate id="BD805060673BA43A20DFFF435218BFF2" box="[984,1121,187,212]" direction="west" orientation="longitude" pageId="6" pageNumber="507" precision="15" value="-54.556667">54°33'24&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F673BA43A2768FF435323BFF2" box="[1135,1370,187,212]" pageId="6" pageNumber="507" value="1991-10-12" valueMax="1991-10-16" valueMin="1991-10-12">
<date id="AC0A1067673BA43A2768FF435323BFF2" box="[1135,1370,187,212]" pageId="6" pageNumber="507" value="1991-10-12" valueMax="1991-10-16" valueMin="1991-10-12">12–16 October 1991</date>
</collectingDate>
,
<collectorName id="75415371673BA43A2660FF44568FBFD1" pageId="6" pageNumber="507">A.B. Bonaldo</collectorName>
(
<specimenCode id="88129EDC673BA43A2203FF2757D0BFD1" box="[260,425,222,247]" collectionCode="MCNZ" country="Brazil" httpUri="http://grbio.org/cool/wyk1-wacz" name="Porto Alegre, Museu de Ciencias Naturais da Fundacao Zoo-Botanica do Rio Grande do Sul" pageId="6" pageNumber="507">MCNZ 21767</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90AE652C673BA43A23C0FEFA5693BEEA" pageId="6" pageNumber="507" type="diagnosis">
<paragraph id="D80B36A7673BA43A23C0FEFA5693BEEA" blockId="6.[151,1437,151,1276]" pageId="6" pageNumber="507">
<emphasis id="EAC0EAB5673BA43A23C0FEFA5738BE3D" bold="true" box="[199,321,258,283]" pageId="6" pageNumber="507">Diagnosis.</emphasis>
Females of
<taxonomicName id="1FB44D24673BA43A22DDFEFB5416BE3D" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[474,623,258,283]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB5673BA43A22DDFEFB5416BE3D" box="[474,623,258,283]" italics="true" pageId="6" pageNumber="507">E. camelinus</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="1FB44D24673BA43A2050FEFB5258BE3D" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[855,1057,258,283]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB5673BA43A2050FEFB5258BE3D" box="[855,1057,258,283]" italics="true" pageId="6" pageNumber="507">E. trituberculatus</emphasis>
</taxonomicName>
in the shape and disposition of opisthosomal projections but can be distinguished by large brown stains on prosoma and femur I and by the Vshaped mark on patella I (
<figureCitation id="408F2A22673BA43A22B1FEB1544DBE47" box="[438,564,329,354]" captionStart="FIGURE 2" captionStartId="5.[151,250,1855,1877]" captionTargetBox="[151,1436,674,1833]" captionTargetId="figure@5.[151,1436,674,1833]" captionTargetPageId="5" captionText="FIGURE 2. Epicadus camelinus (O. Pickard-Cambridge, 1869), female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A, B), 0.2 mm(C, D)." httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="6" pageNumber="507">Figs 2A, B</figureCitation>
,
<figureCitation id="408F2A22673BA43A2138FEB1540ABE44" box="[575,627,329,354]" captionStart="FIGURE 16" captionStartId="26.[151,250,1869,1891]" captionTargetBox="[184,1408,193,1859]" captionTargetId="figure@26.[184,1408,193,1859]" captionTargetPageId="26" captionText="FIGURE 16. Female specimens in vivo. A Epicadus taczanowskii (RoeWer, 1951); B Epicadus dimidiaster sp. nov.; C Epicadus rubripes Mello-Leitão, 1924 (adapted from Silva-Moreira &amp; Machado 2016: fig. 1F); D Epicadus camelinus (O. Pickard-Cambridge, 1869); E Epicadus heterogaster (Guérin-Méneville, 1829); F Epicadus trituberculatus (TaczanoWski, 1872). Photo credits:César Favacho (A); Arthur Anker (B); Almir Cândido de Almeida (C); Karl Kroeker (D); Edson Roberto (E); Edelcio Muscat (F)." httpUri="https://zenodo.org/record/1183162/files/figure.png" pageId="6" pageNumber="507">16D</figureCitation>
). The epigynum in ventral view is also very similar with other species of the genus (especially with
<taxonomicName id="1FB44D24673BA43A22CFFE9654F7BEA3" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[456,654,364,389]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB5673BA43A22CFFE9654F7BEA3" box="[456,654,364,389]" italics="true" pageId="6" pageNumber="507">E. trituberculatus</emphasis>
</taxonomicName>
) but is characterized by the straight lateral folds delimiting a deeper median field, by the copulatory openings being totally exposed and by the reduced secondary spermathecae (
<figureCitation id="408F2A22673BA43A266DFE6856A4BEEA" captionStart="FIGURE 2" captionStartId="5.[151,250,1855,1877]" captionTargetBox="[151,1436,674,1833]" captionTargetId="figure@5.[151,1436,674,1833]" captionTargetPageId="5" captionText="FIGURE 2. Epicadus camelinus (O. Pickard-Cambridge, 1869), female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A, B), 0.2 mm(C, D)." httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="6" pageNumber="507">Figs 2C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90AE652C673BA43A23C0FE2F536ABB48" pageId="6" pageNumber="507" type="description">
<paragraph id="D80B36A7673BA43A23C0FE2F52D4BCBF" blockId="6.[151,1437,151,1276]" pageId="6" pageNumber="507">
<emphasis id="EAC0EAB5673BA43A23C0FE2F55A7BED6" bold="true" box="[199,990,471,496]" pageId="6" pageNumber="507">Description. Female (from Senador Guiomard; UFMG 11007):</emphasis>
Anterior eye row strongly recurved, posterior eye row slightly procurved, almost straight. Prosoma as long as wide, MS and TWS present, LBL extends longitudinally to the PME (
<figureCitation id="408F2A22673BA43A22D2FDE5544BBD10" box="[469,562,541,566]" captionStart="FIGURE 2" captionStartId="5.[151,250,1855,1877]" captionTargetBox="[151,1436,674,1833]" captionTargetId="figure@5.[151,1436,674,1833]" captionTargetPageId="5" captionText="FIGURE 2. Epicadus camelinus (O. Pickard-Cambridge, 1869), female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A, B), 0.2 mm(C, D)." httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="6" pageNumber="507">Fig. 2A</figureCitation>
). Clypeus brownish, wide and bearing a black upside down V-shaped mark (
<figureCitation id="408F2A22673BA43A2398FDB95779BD7C" box="[159,256,577,602]" captionStart="FIGURE 2" captionStartId="5.[151,250,1855,1877]" captionTargetBox="[151,1436,674,1833]" captionTargetId="figure@5.[151,1436,674,1833]" captionTargetPageId="5" captionText="FIGURE 2. Epicadus camelinus (O. Pickard-Cambridge, 1869), female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A, B), 0.2 mm(C, D)." httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="6" pageNumber="507">Fig. 2B</figureCitation>
); chelicerae, sternum, endites, and labium yellowish. Opisthosoma yellowish with brown stains, presenting several black knobs and five opisthosomal projections faced backward (
<figureCitation id="408F2A22673BA43A2787FD9D529CBD5B" box="[1152,1253,613,638]" captionStart="FIGURE 2" captionStartId="5.[151,250,1855,1877]" captionTargetBox="[151,1436,674,1833]" captionTargetId="figure@5.[151,1436,674,1833]" captionTargetPageId="5" captionText="FIGURE 2. Epicadus camelinus (O. Pickard-Cambridge, 1869), female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A, B), 0.2 mm(C, D)." httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="6" pageNumber="507">Figs 2A</figureCitation>
,
<figureCitation id="408F2A22673BA43A27FFFD9D5355BD58" box="[1272,1324,613,638]" captionStart="FIGURE 16" captionStartId="26.[151,250,1869,1891]" captionTargetBox="[184,1408,193,1859]" captionTargetId="figure@26.[184,1408,193,1859]" captionTargetPageId="26" captionText="FIGURE 16. Female specimens in vivo. A Epicadus taczanowskii (RoeWer, 1951); B Epicadus dimidiaster sp. nov.; C Epicadus rubripes Mello-Leitão, 1924 (adapted from Silva-Moreira &amp; Machado 2016: fig. 1F); D Epicadus camelinus (O. Pickard-Cambridge, 1869); E Epicadus heterogaster (Guérin-Méneville, 1829); F Epicadus trituberculatus (TaczanoWski, 1872). Photo credits:César Favacho (A); Arthur Anker (B); Almir Cândido de Almeida (C); Karl Kroeker (D); Edson Roberto (E); Edelcio Muscat (F)." httpUri="https://zenodo.org/record/1183162/files/figure.png" pageId="6" pageNumber="507">16D</figureCitation>
). Median posterior opisthosomal projection larger than lateral ones, distally enlarged, ending in two roundish tips. In posterior view, between the dorsolateral and ventrolateral projections, there is a black triangular stain (
<figureCitation id="408F2A22673BA43A2630FD5453EEBDE2" box="[1335,1431,684,709]" captionStart="FIGURE 2" captionStartId="5.[151,250,1855,1877]" captionTargetBox="[151,1436,674,1833]" captionTargetId="figure@5.[151,1436,674,1833]" captionTargetPageId="5" captionText="FIGURE 2. Epicadus camelinus (O. Pickard-Cambridge, 1869), female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A, B), 0.2 mm(C, D)." httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="6" pageNumber="507">Figs 2A</figureCitation>
,
<figureCitation id="408F2A22673BA43A2390FD3756B2BDCE" box="[151,203,719,744]" captionStart="FIGURE 16" captionStartId="26.[151,250,1869,1891]" captionTargetBox="[184,1408,193,1859]" captionTargetId="figure@26.[184,1408,193,1859]" captionTargetPageId="26" captionText="FIGURE 16. Female specimens in vivo. A Epicadus taczanowskii (RoeWer, 1951); B Epicadus dimidiaster sp. nov.; C Epicadus rubripes Mello-Leitão, 1924 (adapted from Silva-Moreira &amp; Machado 2016: fig. 1F); D Epicadus camelinus (O. Pickard-Cambridge, 1869); E Epicadus heterogaster (Guérin-Méneville, 1829); F Epicadus trituberculatus (TaczanoWski, 1872). Photo credits:César Favacho (A); Arthur Anker (B); Almir Cândido de Almeida (C); Karl Kroeker (D); Edson Roberto (E); Edelcio Muscat (F)." httpUri="https://zenodo.org/record/1183162/files/figure.png" pageId="6" pageNumber="507">16D</figureCitation>
). Anterior border of opisthosoma strongly concave. Femora II with black spots on prolateral surface, one on median and another on distal position. Legs I and II with a median keel including the entire patella to the tarsal joint; anterior tibiae (I and II) with a dorsal naked gutter connected to the sensorial pit. Femora I and II with two ventral lines of setiferous tubercles. Tarsal claws curved and pectinated with teeth restricted to basal portion. Epigynum with an incomplete median septum, secondary spermathecae reduced and primary spermathecae globular; copulatory ducts long, exceeding the length of primary spermathecae) (
<figureCitation id="408F2A22673BA43A2726FC7852D9BCBF" box="[1057,1184,896,921]" captionStart="FIGURE 2" captionStartId="5.[151,250,1855,1877]" captionTargetBox="[151,1436,674,1833]" captionTargetId="figure@5.[151,1436,674,1833]" captionTargetPageId="5" captionText="FIGURE 2. Epicadus camelinus (O. Pickard-Cambridge, 1869), female (UFMG 11007). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A, B), 0.2 mm(C, D)." httpUri="https://zenodo.org/record/1183134/files/figure.png" pageId="6" pageNumber="507">Figs 2C, D</figureCitation>
).
</paragraph>
<paragraph id="D80B36A7673BA43A23C0FC5D536ABB48" blockId="6.[151,1437,151,1276]" pageId="6" pageNumber="507">Measurements: eye diameters and interdistances: AME 0.10, ALE 0.14, PME 0.10, PLE 0.08, AME–AME 0.26, AME–ALE 0.12, PME–PME 0.36, PME–PLE 0.18; MOQ length 0.58, MOQ posterior width 0.58, MOQ anterior width 0.56; leg formula: 1-2-4-3: leg I—femur 3.50/ patella 2.00/ tibia 2.35/ metatarsus 1.75/ tarsus 1.40/ total 11.00; II—3.50/ 2.00/ 2.30/ 1.60/ 1.40/ 10.80; III—1.70/ 1.15/ 1.20/ 1.40/ 0.60/ 6.05; IV—1.90/ 1.05/ 1.50/ 1.35/ 0.60/6.40. Total body length 7.75; prosoma length 3.75, width 3.75; opisthosoma length 4.00; clypeus height 0.50; sternum length 1.68, width 1.16; endites length 1.08, width 0.50; labium length 0.66, width 0.60.</paragraph>
</subSubSection>
<subSubSection id="90AE652C673BA43A23C0FB815429BBDD" pageId="6" pageNumber="507" type="distribution">
<paragraph id="D80B36A7673BA43A23C0FB81525CBBB7" blockId="6.[151,1437,151,1276]" box="[199,1061,1145,1170]" pageId="6" pageNumber="507">
<emphasis id="EAC0EAB5673BA43A23C0FB815719BBB4" bold="true" box="[199,352,1145,1170]" pageId="6" pageNumber="507">Distribution.</emphasis>
<collectingCountry id="A0A37637673BA43A2260FB8257A9BBB4" box="[359,464,1146,1170]" name="Brazil" pageId="6" pageNumber="507">BRAZIL</collectingCountry>
:
<collectingRegion id="1A70F845673BA43A22DCFB82546DBBB4" box="[475,532,1146,1170]" country="Brazil" name="Acre" pageId="6" pageNumber="507">Acre</collectingRegion>
,
<collectingRegion id="1A70F845673BA43A2118FB8254E0BBB4" box="[543,665,1146,1170]" country="Brazil" name="Amazonas" pageId="6" pageNumber="507">Amazonas</collectingRegion>
;
<collectingCountry id="A0A37637673BA43A21A3FB815565BBB7" box="[676,796,1145,1169]" name="Bolivia" pageId="6" pageNumber="507">BOLIVIA</collectingCountry>
:
<collectingRegion id="1A70F845673BA43A2021FB8155D8BBB7" box="[806,929,1145,1169]" country="Bolivia" name="Santa Cruz" pageId="6" pageNumber="507">Santa Cruz</collectingRegion>
(
<figureCitation id="408F2A22673BA43A20B6FB815261BBB7" box="[945,1048,1145,1170]" captionStart="FIGURE 17" captionStartId="27.[151,250,1766,1788]" captionTargetBox="[151,1435,193,1736]" captionTargetId="figure@27.[151,1436,193,1744]" captionTargetPageId="27" captionText="FIGURE 17. Data of distribution records of Epicadus species compiled from Machado et al. (2015, 2017); Silva-Moreira and Machado (2016); present study." httpUri="https://zenodo.org/record/1183164/files/figure.png" pageId="6" pageNumber="507">Fig. 17C</figureCitation>
).
</paragraph>
<paragraph id="D80B36A7673BA43A23C0FB645429BBDD" blockId="6.[151,1437,151,1276]" pageId="6" pageNumber="507">
<emphasis id="EAC0EAB5673BA43A23C0FB64577FBB93" bold="true" box="[199,262,1180,1205]" pageId="6" pageNumber="507">Note.</emphasis>
The male of
<taxonomicName id="1FB44D24673BA43A2298FB665448BB93" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[415,561,1180,1205]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB5673BA43A2298FB665448BB93" box="[415,561,1180,1205]" italics="true" pageId="6" pageNumber="507">E. camelinus</emphasis>
</taxonomicName>
is still unknown. The scanning microscopy of a male palp represented in the supplementary material of
<bibRefCitation id="BC254B56673BA43A22C1FB4754B8BBFE" box="[454,705,1215,1240]" pageId="6" pageNumber="507" refId="ref19045" refString="Machado, M, Teixeira, R. A. &amp; Lise, A. A. (2017) Cladistic analysis supports the monophyly of the Neotropical crab spider genus Epicadus and its senior synonymy over Tobias (Araneae: Thomisidae). Invertebrate Systematics, 31, 442 - 455. https: // doi. org / 10.1071 / IS 16074" type="journal article">
Machado
<emphasis id="EAC0EAB5673BA43A2131FB395417BBFE" box="[566,622,1215,1240]" italics="true" pageId="6" pageNumber="507">et al.</emphasis>
(2017)
</bibRefCitation>
(
<figureCitation id="408F2A22673BA43A21D6FB47553CBBFE" box="[721,837,1215,1240]" captionStart="FIGURE 12" captionStartId="22.[151,250,1876,1898]" captionTargetBox="[151,1436,733,1854]" captionTargetId="figure@22.[151,1436,733,1854]" captionTargetPageId="22" captionText="FIGURE 12. Epicadus trituberculatus (TaczanoWski, 1872), female (MPEG 022662). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A), 1 mm (B), 0.2mm (C, D)." httpUri="https://zenodo.org/record/1183154/files/figure.png" pageId="6" pageNumber="507">Fig. S12B</figureCitation>
) was a
<emphasis id="EAC0EAB5673BA43A209AFB47559DBBFE" box="[925,996,1215,1240]" italics="true" pageId="6" pageNumber="507">lapsus</emphasis>
, being in fact that copulatory structure belonging to a male of
<taxonomicName id="1FB44D24673BA43A229DFB1C5430BBDA" baseAuthorityName="Roewer" baseAuthorityYear="1951" box="[410,585,1251,1276]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="taczanowskii">
<emphasis id="EAC0EAB5673BA43A229DFB1C5430BBDA" box="[410,585,1251,1276]" italics="true" pageId="6" pageNumber="507">E. taczanowskii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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