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<document id="CBF68F221F57C13997B986B5908774B3" ID-CLB-Dataset="30682" ID-DOI="10.11646/zootaxa.4382.3.4" ID-GBIF-Dataset="b88bfa2d-4b83-41f8-a145-a95ca196960f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1183130" ID-ZooBank="87A2B333-09FF-4726-A1EF-0699FB631808" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1519301947700" checkinUser="plazi" docAuthor="Machado, Miguel, Teixeira, Renato Augusto &amp; Lise, Arno Antonio" docDate="2018" docId="501D87B16733A4332390FB5454E3BD1A" docLanguage="en" docName="zootaxa.4382.3.4.pdf" docOrigin="Zootaxa 4382 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epicadus pustulosus" docType="treatment" docVersion="8" lastPageNumber="516" masterDocId="AC24FFC9673DA43C2307FFF85679BF26" masterDocTitle="There and back again: More on the taxonomy of the crab spiders genus Epicadus (Thomisidae: Stephanopinae)" masterLastPageNumber="530" masterPageNumber="501" pageNumber="515" updateTime="1698524041822" updateUser="ExternalLinkService">
<mods:mods id="5CF6082F45E6917E8106B01FF9C6FE98" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C86C7905FCA92FEABBFD2513A05B1BE5">There and back again: More on the taxonomy of the crab spiders genus Epicadus (Thomisidae: Stephanopinae)</mods:title>
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<mods:namePart id="43F7E6F4E34F535C35D638D5C526D350">Machado, Miguel</mods:namePart>
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<mods:namePart id="05BBA27EC85A458093218A858E246BBF">Teixeira, Renato Augusto</mods:namePart>
</mods:name>
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<mods:role id="CC9DC784C8F5B4A8EFCE4850642AC72D">
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<mods:namePart id="CA725614E1317DA29F8CAE7A2778E368">Lise, Arno Antonio</mods:namePart>
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<mods:title id="F8CA1DDA413EA0852CE685E2CD391F15">Zootaxa</mods:title>
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<mods:part id="F2744DC1A786B2DD6FBE05DD8D9F46AC">
<mods:date id="D4C686FBB533ADC105EE936F791D2121">2018</mods:date>
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<mods:number id="F5D873A22D4D76CDD48F8F3F4318B52B">2018-02-22</mods:number>
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<mods:number id="1DD3FC32DE8CA7ACBD172094BEEF1B49">4382</mods:number>
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<mods:identifier id="3391DA2F34C4D4D58473983D639CE5E6" type="DOI">10.11646/zootaxa.4382.3.4</mods:identifier>
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<treatment id="501D87B16733A4332390FB5454E3BD1A" ID-DOI="http://doi.org/10.5281/zenodo.5951324" ID-GBIF-Taxon="141118937" ID-Zenodo-Dep="5951324" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A" httpUri="http://treatment.plazi.org/id/501D87B16733A4332390FB5454E3BD1A" lastPageId="15" lastPageNumber="516" pageId="14" pageNumber="515">
<subSubSection id="90AE652C6733A4322390FB545725BBC1" pageId="14" pageNumber="515" type="nomenclature">
<paragraph id="D80B36A76733A4322390FB5454EDBBE0" blockId="14.[151,660,1196,1255]" box="[151,660,1196,1222]" pageId="14" pageNumber="515">
<heading id="834381CB6733A4322390FB5454EDBBE0" bold="true" box="[151,660,1196,1222]" fontSize="11" level="1" pageId="14" pageNumber="515" reason="1">
<taxonomicName id="1FB44D246733A4322390FB5454EDBBE0" ID-CoL="6G3DY" authority="(Mello-Leitao, 1929)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[151,660,1196,1222]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="EAC0EAB56733A4322390FB5454EDBBE0" bold="true" box="[151,660,1196,1222]" pageId="14" pageNumber="515">
<emphasis id="EAC0EAB56733A4322390FB5457F6BBE0" bold="true" box="[151,399,1196,1222]" italics="true" pageId="14" pageNumber="515">Epicadus pustulosus</emphasis>
(
<bibRefCitation id="BC254B566733A4322299FB5454F5BBE0" box="[414,652,1196,1222]" pageId="14" pageNumber="515" refId="ref19139" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Archivos do Museu Nacional do Rio de Janeiro, Rio de Janeiro, 360 pp." type="book">Mello-Leitão, 1929</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D80B36A76733A4322390FB365725BBC1" blockId="14.[151,660,1196,1255]" box="[151,348,1230,1255]" pageId="14" pageNumber="515">
<figureCitation id="408F2A226733A4322390FB36575ABBC1" box="[151,291,1230,1255]" captionStart="FIGURE 14" captionStartId="24.[151,250,1937,1959]" captionTargetBox="[181,1404,193,1921]" captionTargetId="figure@24.[181,1404,193,1921]" captionTargetPageId="24" captionText="FIGURE 14. Color variations of females. A–D Epicadus pustulosus (Mello-Leitão, 1929) (A, B MZSP 45484; C, D MZSP 51068); Scale bars: 2 mm (A, C), 1 mm (B, D). Color variations of females of E–F Epicadus taczanowskii (RoeWer, 1951): Photo credits:Kenji Nishida (E); Arthur Anker (F)." httpUri="https://zenodo.org/record/1183158/files/figure.png" pageId="14" pageNumber="515">Figs 14A–D</figureCitation>
,
<figureCitation id="408F2A226733A4322229FB365725BBC1" box="[302,348,1230,1255]" captionStart="FIGURE 15" captionStartId="25.[151,250,1854,1876]" captionTargetBox="[151,1436,193,1833]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURE 15. Female specimens in vivo. A–C Epicadus caudatus (Mello-Leitão, 1929) A nuptial guard, B egg-sac guarding behavior, C diagnostic orange blister-shaped setae sockets); D E. granulatus Banks 1909; E Epicadus pustulosus (Mello- Leitão, 1929), diagnostic yelloW blister-shaped setae sockets. Photo credits: Alfredo Colón Archilla (A, B, C); Morley Read (D); Paul Bertner (E)." httpUri="https://zenodo.org/record/1183160/files/figure.png" pageId="14" pageNumber="515">15E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6733A4322390FAEE55CABA4C" pageId="14" pageNumber="515" type="reference_group">
<paragraph id="D80B36A76733A4322390FAEE5546BA6D" blockId="14.[151,1117,1302,1386]" pageId="14" pageNumber="515">
<treatmentCitationGroup id="F8A411896733A4322390FAEE5546BA6D" pageId="14" pageNumber="515">
<taxonomicName id="1FB44D246733A4322390FAEE5447BA0A" authority="Mello-Leitao, 1929: 86" authorityName="Mello-Leitao" authorityPageNumber="86" authorityYear="1929" box="[151,574,1302,1324]" class="Arachnida" family="Thomisidae" genus="Tobias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="EAC0EAB56733A4322390FAEE5732BA0A" box="[151,331,1302,1324]" italics="true" pageId="14" pageNumber="515">Tobias pustulosus</emphasis>
<treatmentCitation id="591510B66733A4322256FAEE5447BA0A" author="" box="[337,574,1302,1324]" page="86" pageId="14" pageNumber="515">
<bibRefCitation id="BC254B566733A4322256FAEE5460BA0A" box="[337,537,1302,1324]" pageId="14" pageNumber="515" refId="ref19139" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Archivos do Museu Nacional do Rio de Janeiro, Rio de Janeiro, 360 pp." type="book">Mello-Leitão, 1929</bibRefCitation>
: 86
</treatmentCitation>
</taxonomicName>
, figs 183–184.
<treatmentCitation id="591510B66733A43221DCFAEE5598BA0A" author="" box="[731,993,1302,1325]" page="572" pageId="14" pageNumber="515">
<bibRefCitation id="BC254B566733A43221DCFAEE55D7BA0A" box="[731,942,1302,1325]" pageId="14" pageNumber="515" refId="ref18982" refString="Machado, M., Teixeira, R. A. &amp; Lise, A. A. (2015) Taxonomic notes on the crab spider genus Tobias Simon, 1895 (Araneae, Thomisidae, Stephanopinae). Zootaxa, 4034 (3), 565 - 576. https: // doi. org / 10.11646 / zootaxa. 4034.3.8" type="journal article">
Machado
<emphasis id="EAC0EAB56733A4322038FAE0550ABA0A" box="[831,883,1302,1325]" italics="true" pageId="14" pageNumber="515">et al.</emphasis>
2015
</bibRefCitation>
: 572
</treatmentCitation>
, figs 12–24.
<taxonomicName id="1FB44D246733A4322390FACD54D1BA6D" authority="Simon in Mello-Leitao, 1929: 87" authorityName="Simon in Mello-Leitao" authorityPageNumber="87" authorityYear="1929" box="[151,680,1333,1355]" class="Arachnida" family="Thomisidae" genus="Tobias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="monstrosus">
<emphasis id="EAC0EAB56733A4322390FACD572ABA6D" box="[151,339,1333,1355]" italics="true" pageId="14" pageNumber="515">Tobias monstrosus</emphasis>
Simon in
<treatmentCitation id="591510B66733A43222BCFACD54D1BA6D" author="" box="[443,680,1333,1355]" page="87" pageId="14" pageNumber="515">
<bibRefCitation id="BC254B566733A43222BCFACD54FABA6D" box="[443,643,1333,1355]" pageId="14" pageNumber="515" refId="ref19139" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Archivos do Museu Nacional do Rio de Janeiro, Rio de Janeiro, 360 pp." type="book">Mello-Leitão, 1929</bibRefCitation>
: 87
</treatmentCitation>
</taxonomicName>
, figs 185–186.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D80B36A76733A4322390FAAC55CABA4C" blockId="14.[151,1117,1302,1386]" box="[151,947,1364,1386]" pageId="14" pageNumber="515">
<treatmentCitationGroup id="F8A411896733A4322390FAAC55CABA4C" box="[151,947,1364,1386]" pageId="14" pageNumber="515">
<taxonomicName id="1FB44D246733A4322390FAAC547FBA4C" ID-CoL="6G3DY" authority="(Mello-Leitao)" authorityName="Mello-Leitao" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[151,518,1364,1386]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="EAC0EAB56733A4322390FAAC571FBA4C" box="[151,358,1364,1386]" italics="true" pageId="14" pageNumber="515">Epicadus pustulosus</emphasis>
(Mello-Leitão)
</taxonomicName>
.
<treatmentCitation id="591510B66733A4322117FAAC556CBA4C" author="" box="[528,789,1364,1386]" page="448" pageId="14" pageNumber="515">
<bibRefCitation id="BC254B566733A4322117FAAC549ABA4C" box="[528,739,1364,1386]" pageId="14" pageNumber="515" refId="ref19045" refString="Machado, M, Teixeira, R. A. &amp; Lise, A. A. (2017) Cladistic analysis supports the monophyly of the Neotropical crab spider genus Epicadus and its senior synonymy over Tobias (Araneae: Thomisidae). Invertebrate Systematics, 31, 442 - 455. https: // doi. org / 10.1071 / IS 16074" type="journal article">
Machado
<emphasis id="EAC0EAB56733A4322173FAAD54DEBA4C" box="[628,679,1364,1386]" italics="true" pageId="14" pageNumber="515">et al.</emphasis>
2017
</bibRefCitation>
: 448
</treatmentCitation>
, figs S7D, S
<specimenCount id="CEB2FD2E6733A4322094FAAC55CABA4C" box="[915,947,1364,1386]" pageId="14" pageNumber="515" type="generic">8F.</specimenCount>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6733A4322390FA6E525DB881" pageId="14" pageNumber="515" type="materials_examined">
<paragraph id="D80B36A76733A4322390FA6E54E0BAF5" blockId="14.[151,1436,1430,2031]" pageId="14" pageNumber="515">
<materialsCitation id="68DC3CFA6733A4322390FA6E54E0BAF5" ID-GBIF-Occurrence="1820860277" collectionCode="MNHN" country="Brazil" latitude="-2.4681" location="Fonte Boa" longLatPrecision="1" longitude="-66.141525" pageId="14" pageNumber="515" specimenCode="MNHN 4560" specimenCount="2" stateProvince="Amazonas" typeStatus="holotype">
<emphasis id="EAC0EAB56733A4322390FA6E57B7BA89" bold="true" box="[151,462,1430,1455]" pageId="14" pageNumber="515">
<typeStatus id="070F88056733A4322390FA6E56ABBA89" box="[151,210,1430,1455]" pageId="14" pageNumber="515">Type</typeStatus>
material:
<typeStatus id="070F88056733A4322266FA6E57B7BA89" box="[353,462,1430,1455]" pageId="14" pageNumber="515" type="holotype">Holotype</typeStatus>
</emphasis>
(original designation): subadult female,
<location id="DD6B607C6733A43220C3FA6F5238BA88" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A43220C3FA6F5238BA88" box="[964,1089,1431,1454]" country="Brazil" latitude="-2.4681" longLatPrecision="1" longitude="-66.141525" name="Fonte Boa" pageId="14" pageNumber="515" stateProvince="Amazonas">Fonte Boa</location>
[
<geoCoordinate id="BD8050606733A432275EFA6E529DBA89" box="[1113,1252,1430,1455]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="1" value="-2.4681">2°28'5.16&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A43227FFFA6E53E1BA89" box="[1272,1432,1430,1455]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="1" value="-66.141525">66°8'29.49&quot;W</geoCoordinate>
,
<collectingRegion id="1A70F8456733A4322390FA435769BAF5" box="[151,272,1467,1491]" country="Brazil" name="Amazonas" pageId="14" pageNumber="515">Amazonas</collectingRegion>
,
<collectingCountry id="A0A376376733A432221CFA42571BBAF5" box="[283,354,1466,1491]" name="Brazil" pageId="14" pageNumber="515">Brazil</collectingCountry>
] (
<specimenCode id="88129EDC6733A4322270FA435468BAF5" box="[375,529,1466,1491]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="515">MNHN 4560</specimenCode>
, examined).
</materialsCitation>
</paragraph>
<paragraph id="D80B36A76733A43223C0FA26525DB881" blockId="14.[151,1436,1430,2031]" pageId="14" pageNumber="515">
<materialsCitation id="68DC3CFA6733A43223C0FA26554FB93D" ID-GBIF-Occurrence="1820860237" collectionCode="IBSP" collectorName="E. F. Morato" country="Brazil" county="Other" latitude="-10.145278" location="Reserva Extrativista de Catuaba" longLatPrecision="21" longitude="-67.724724" municipality="Senador Guiomard" pageId="14" pageNumber="515" specimenCode="IBSP 135286" specimenCount="1" specimenCount-male="1" stateProvince="Acre">
<emphasis id="EAC0EAB56733A43223C0FA2654B0BAD1" bold="true" box="[199,713,1502,1527]" pageId="14" pageNumber="515">
<collectingCounty id="316A4E2B6733A43223C0FA265776BAD1" box="[199,271,1502,1527]" pageId="14" pageNumber="515">Other</collectingCounty>
material examined.
<collectingCountry id="A0A376376733A432210CFA265402BAD1" box="[523,635,1502,1527]" name="Brazil" pageId="14" pageNumber="515">BRAZIL</collectingCountry>
:
<collectingRegion id="1A70F8456733A432218DFA2654B9BAD1" box="[650,704,1502,1527]" country="Brazil" name="Acre" pageId="14" pageNumber="515">
<emphasis id="EAC0EAB56733A432218DFA2654B9BAD1" bold="true" box="[650,704,1502,1527]" italics="true" pageId="14" pageNumber="515">Acre</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6733A43221D4FA27555DBAD1" box="[723,804,1502,1527]" pageId="14" pageNumber="515" type="male">1 male</specimenCount>
,
<collectingMunicipality id="386FACDD6733A4322035FA265270BAD1" box="[818,1033,1502,1527]" pageId="14" pageNumber="515">Senador Guiomard</collectingMunicipality>
(
<location id="DD6B607C6733A432271CFA2753E9BAD1" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A432271CFA2753E9BAD1" box="[1051,1424,1502,1527]" country="Brazil" county="Other" latitude="-10.145278" longLatPrecision="21" longitude="-67.724724" municipality="Senador Guiomard" name="Reserva Extrativista de Catuaba" pageId="14" pageNumber="515" stateProvince="Acre">Reserva Extrativista de Catuaba</location>
),
<geoCoordinate id="BD8050606733A4322390F9FB5761B93D" box="[151,280,1539,1563]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="15" value="-10.145278">10°08'43&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A4322223F9FA57D4B93D" box="[292,429,1538,1563]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="15" value="-67.724724">67°43'29&quot;W</geoCoordinate>
, 2002,
<collectorName id="754153716733A43222F9F9FB54FDB93C" box="[510,644,1539,1562]" pageId="14" pageNumber="515">E.F. Morato</collectorName>
(
<specimenCode id="88129EDC6733A4322195F9FB5557B93D" box="[658,814,1538,1563]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="14" pageNumber="515">IBSP 135286</specimenCode>
)
</materialsCitation>
; 1 male (IBSP 135287); 1 male (IBSP 135289); 1 male (IBSP 135290); 1 female (IBSP 135291);
<materialsCitation id="68DC3CFA6733A4322176F9DF536FB919" ID-GBIF-Occurrence="1820860285" box="[625,1302,1574,1599]" collectionCode="IBSP" collectorName="E. F. Morato" country="Brazil" county="Other" latitude="-10.145278" location="Reserva Extrativista de Catuaba" longLatPrecision="21" longitude="-67.724724" municipality="Senador Guiomard" pageId="14" pageNumber="515" specimenCode="IBSP 135292" specimenCount="1" specimenCount-male="1" stateProvince="Acre">
<specimenCount id="CEB2FD2E6733A4322176F9DF54C4B919" box="[625,701,1574,1599]" pageId="14" pageNumber="515" type="male">1 male</specimenCount>
(
<specimenCode id="88129EDC6733A43221CAF9DF5513B919" box="[717,874,1574,1599]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="14" pageNumber="515">IBSP 135292</specimenCode>
); all same locality and collection data
</materialsCitation>
.
<materialsCitation id="68DC3CFA6733A4322626F9DE52A1B945" ID-GBIF-Occurrence="1820860305" collectingDate="2012-01-20" collectionCode="MZSP" collectorName="D. F. Candiani" country="Brazil" latitude="-8.802778" location="Porto Velho" longLatPrecision="21" longitude="-63.918056" pageId="14" pageNumber="515" specimenCode="MZSP 45458" specimenCount="1" specimenCount-female="1" stateProvince="Rondonia">
<emphasis id="EAC0EAB56733A4322626F9DE53E5B919" bold="true" box="[1313,1436,1574,1599]" pageId="14" pageNumber="515">
<collectingRegion id="1A70F8456733A4322626F9DE53EAB919" box="[1313,1427,1574,1599]" country="Brazil" name="Rondonia" pageId="14" pageNumber="515">
<emphasis id="EAC0EAB56733A4322626F9DE53EAB919" bold="true" box="[1313,1427,1574,1599]" italics="true" pageId="14" pageNumber="515">Rondônia</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6733A4322390F9B35685B945" box="[151,252,1610,1635]" pageId="14" pageNumber="515" type="female">1 female</specimenCount>
,
<location id="DD6B607C6733A432220EF9B357EBB945" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A432220EF9B357EBB945" box="[265,402,1610,1635]" country="Brazil" latitude="-8.802778" longLatPrecision="21" longitude="-63.918056" name="Porto Velho" pageId="14" pageNumber="515" stateProvince="Rondonia">Porto Velho</location>
,
<geoCoordinate id="BD8050606733A4322298F9B3546AB945" box="[415,531,1611,1635]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="15" value="-8.802778">8°48'10&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A4322127F9B254D2B945" box="[544,683,1610,1635]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="15" value="-63.918056">63°55'05&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F6733A43221B0F9B3550EB945" box="[695,887,1611,1635]" pageId="14" pageNumber="515" value="2012-01-20">
<date id="AC0A10676733A43221B0F9B3550EB945" box="[695,887,1611,1635]" pageId="14" pageNumber="515" value="2012-01-20">20 January 2012</date>
</collectingDate>
,
<collectorName id="754153716733A4322083F9B35258B945" box="[900,1057,1610,1635]" pageId="14" pageNumber="515">D.F. Candiani</collectorName>
(
<specimenCode id="88129EDC6733A4322736F9B352A9B945" box="[1073,1232,1610,1635]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="14" pageNumber="515">MZSP 45458</specimenCode>
)
</materialsCitation>
; 1 female,
<date id="AC0A10676733A4322650F9B356ABB9A1" pageId="14" pageNumber="515" value="2012-07-04">4 July 2012</date>
, A.A. Nogueira (MZSP 51068);
<materialsCitation id="68DC3CFA6733A4322139F997528BB98D" ID-GBIF-Occurrence="1820860260" collectingDate="1968-11-05" collectingDateMax="2012-07-08" collectingDateMin="1968-11-05" collectionCode="MZSP, MNRJ" country="Brazil" latitude="-13.743889" location="Indicatti" longLatPrecision="21" longitude="-39.484722" pageId="14" pageNumber="515" specimenCode="MZSP 53085, MNRJ 1351" specimenCount="2" specimenCount-female="2" stateProvince="Bahia">
<specimenCount id="CEB2FD2E6733A4322139F99754DBB9A1" box="[574,674,1646,1671]" pageId="14" pageNumber="515" type="female">1 female</specimenCount>
,
<collectingDate id="BC4EE98F6733A43221ABF9975548B9A1" box="[684,817,1646,1671]" pageId="14" pageNumber="515" value="2012-07-08">
<date id="AC0A10676733A43221ABF9975548B9A1" box="[684,817,1646,1671]" pageId="14" pageNumber="515" value="2012-07-08">8 July 2012</date>
</collectingDate>
, R.P.
<location id="DD6B607C6733A4322068F99655B5B9A1" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A4322068F99655B5B9A1" box="[879,972,1646,1671]" country="Brazil" latitude="-13.743889" longLatPrecision="21" longitude="-39.484722" name="Indicatti" pageId="14" pageNumber="515" stateProvince="Bahia">Indicatti</location>
(
<specimenCode id="88129EDC6733A43220DDF9975201B9A1" box="[986,1144,1646,1671]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="14" pageNumber="515">MZSP 53085</specimenCode>
.;
<emphasis id="EAC0EAB56733A4322781F99652ACB9A1" bold="true" box="[1158,1237,1646,1671]" pageId="14" pageNumber="515">
<collectingRegion id="1A70F8456733A4322781F99652B5B9A1" box="[1158,1228,1646,1671]" country="Brazil" name="Bahia" pageId="14" pageNumber="515">
<emphasis id="EAC0EAB56733A4322781F99652B5B9A1" bold="true" box="[1158,1228,1646,1671]" italics="true" pageId="14" pageNumber="515">Bahia</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6733A43227DBF9975339B9A1" box="[1244,1344,1646,1671]" pageId="14" pageNumber="515" type="female">1 female</specimenCount>
, Gandu, Fazenda São Rafael (cocoa forest),
<geoCoordinate id="BD8050606733A4322138F96B54B9B98D" box="[575,704,1683,1707]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="15" value="-13.743889">13°44'38&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A43221D6F96A5522B98D" box="[721,859,1682,1707]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="15" value="-39.484722">39°29'05&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F6733A432206BF96A5238B98D" box="[876,1089,1682,1707]" pageId="14" pageNumber="515" value="1968-11-05">
<date id="AC0A10676733A432206BF96A5238B98D" box="[876,1089,1682,1707]" pageId="14" pageNumber="515" value="1968-11-05">5 November 1968</date>
</collectingDate>
(
<specimenCode id="88129EDC6733A4322753F96B5290B98D" box="[1108,1257,1682,1707]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="14" pageNumber="515">MNRJ 1351</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="68DC3CFA6733A4322605F96B53E1B9E9" ID-GBIF-Occurrence="1820860246" collectingDate="2000-11-15" collectingDateMax="2000-11-28" collectingDateMin="2000-11-15" collectionCode="IBSP" collectorName="A. D. Brescovit" country="Brazil" latitude="-15.208889" location="Reserva Biologica de Una" longLatPrecision="21" longitude="-39.195835" municipality="Una" pageId="14" pageNumber="515" specimenCode="IBSP 46728" specimenCount="1" specimenCount-male="1" stateProvince="Bahia">
<specimenCount id="CEB2FD2E6733A4322605F96B532FB98D" box="[1282,1366,1682,1707]" pageId="14" pageNumber="515" type="male">1 male</specimenCount>
,
<collectingMunicipality id="386FACDD6733A4322661F96B53E1B98C" box="[1382,1432,1683,1706]" pageId="14" pageNumber="515">Una</collectingMunicipality>
,
<location id="DD6B607C6733A4322390F94F57BAB9E8" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A4322390F94F57BAB9E8" box="[151,451,1718,1743]" country="Brazil" latitude="-15.208889" longLatPrecision="21" longitude="-39.195835" municipality="Una" name="Reserva Biologica de Una" pageId="14" pageNumber="515" stateProvince="Bahia">Reserva Biológica de Una</location>
,
<geoCoordinate id="BD8050606733A43222C8F94E5429B9E9" box="[463,592,1718,1743]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="15" value="-15.208889">15°12'32&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A432215AF94E549FB9E9" box="[605,742,1718,1743]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="15" value="-39.195835">39°11'45&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F6733A43221F5F94E5583B9E9" box="[754,1018,1718,1743]" pageId="14" pageNumber="515" value="2000-11-15" valueMax="2000-11-28" valueMin="2000-11-15">
<date id="AC0A10676733A43221F5F94E5583B9E9" box="[754,1018,1718,1743]" pageId="14" pageNumber="515" value="2000-11-15" valueMax="2000-11-28" valueMin="2000-11-15">15–28 November 2000</date>
</collectingDate>
,
<collectorName id="754153716733A4322701F94F52C9B9E9" box="[1030,1200,1718,1743]" pageId="14" pageNumber="515">A.D. Brescovit</collectorName>
<emphasis id="EAC0EAB56733A43227BFF94F5288B9E9" box="[1208,1265,1718,1743]" italics="true" pageId="14" pageNumber="515">et al.</emphasis>
(
<specimenCode id="88129EDC6733A4322606F94F53E9B9E9" box="[1281,1424,1718,1743]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="14" pageNumber="515">IBSP 46728</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="68DC3CFA6733A4322390F922546CB831" ID-GBIF-Occurrence="1820860250" collectingDate="1997-07" collectionCode="IBSP" collectorName="A. J. Santos" country="Brazil" latitude="-19.151388" location="Reserva Floresta da Cia. Vale do Rio Doce" longLatPrecision="21" longitude="-40.070553" municipality="Sao Mateus" pageId="14" pageNumber="515" specimenCode="IBSP 27015" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<emphasis id="EAC0EAB56733A4322390F9225731B9D5" bold="true" box="[151,328,1754,1779]" pageId="14" pageNumber="515">
<collectingRegion id="1A70F8456733A4322390F9225746B9D5" box="[151,319,1754,1779]" country="Brazil" name="Espirito Santo" pageId="14" pageNumber="515">
<emphasis id="EAC0EAB56733A4322390F9225746B9D5" bold="true" box="[151,319,1754,1779]" italics="true" pageId="14" pageNumber="515">Espírito Santo</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6733A4322256F92357DBB9D5" box="[337,418,1754,1779]" pageId="14" pageNumber="515" type="male">1 male</specimenCount>
,
<collectingMunicipality id="386FACDD6733A43222A8F9235441B9D4" box="[431,568,1755,1779]" pageId="14" pageNumber="515">São Mateus</collectingMunicipality>
,
<location id="DD6B607C6733A4322141F9235241B9D4" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A4322141F9235241B9D4" box="[582,1080,1754,1779]" country="Brazil" latitude="-19.151388" longLatPrecision="21" longitude="-40.070553" municipality="Sao Mateus" name="Reserva Floresta da Cia. Vale do Rio Doce" pageId="14" pageNumber="515" stateProvince="Espirito Santo">Reserva Floresta da Cia. Vale do Rio Doce</location>
,
<geoCoordinate id="BD8050606733A4322741F92252BEB9D5" box="[1094,1223,1754,1779]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="15" value="-19.151388">19°09'05&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A43227D2F9235326B9D5" box="[1237,1375,1755,1779]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="15" value="-40.070553">40°04'14&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F6733A432266AF92256ABB831" pageId="14" pageNumber="515" value="1997-07">
<date id="AC0A10676733A432266AF92256ABB831" pageId="14" pageNumber="515" value="1997-07">July 1997</date>
</collectingDate>
,
<collectorName id="754153716733A43223E5F907571FB831" box="[226,358,1791,1815]" pageId="14" pageNumber="515">A.J. Santos</collectorName>
(
<specimenCode id="88129EDC6733A432227EF9075474B831" box="[377,525,1790,1815]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="14" pageNumber="515">IBSP 27015</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="68DC3CFA6733A4322123F9065567B831" ID-GBIF-Occurrence="1820860252" box="[548,798,1790,1815]" country="Brazil" location="Sao Paulo" pageId="14" pageNumber="515" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo">
<emphasis id="EAC0EAB56733A4322123F90654D3B831" bold="true" box="[548,682,1790,1815]" italics="true" pageId="14" pageNumber="515">
<collectingRegion id="1A70F8456733A4322123F90654DAB831" box="[548,675,1790,1815]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="515">São Paulo</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6733A43221B2F9075567B831" box="[693,798,1790,1815]" pageId="14" pageNumber="515" type="female">1 female</specimenCount>
</materialsCitation>
,
<materialsCitation id="68DC3CFA6733A4322028F9075417B81D" ID-GBIF-Occurrence="1820860293" collectingDate="1958-08" collectionCode="IBSP" collectorName="M. Kramer" country="Brazil" latitude="-22.706667" location="Parque Modelo" longLatPrecision="21" longitude="-46.809444" pageId="14" pageNumber="515" specimenCode="IBSP 137989" specimenCount="1" stateProvince="Sao Paulo">
<collectingRegion id="1A70F8456733A4322028F90755D1B831" box="[815,936,1790,1815]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="515">São Paulo</collectingRegion>
,
<location id="DD6B607C6733A43220B0F9075212B831" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A43220B0F9075212B831" box="[951,1131,1790,1815]" country="Brazil" latitude="-22.706667" longLatPrecision="21" longitude="-46.809444" name="Parque Modelo" pageId="14" pageNumber="515" stateProvince="Sao Paulo">Parque Modelo</location>
,
<geoCoordinate id="BD8050606733A432277CF9075284B831" box="[1147,1277,1791,1815]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="15" value="-22.706667">22°42'24&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A4322609F90653E1B831" box="[1294,1432,1790,1815]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="15" value="-46.809444">46°48'34&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F6733A4322390F8DB5755B81D" box="[151,300,1826,1851]" pageId="14" pageNumber="515" value="1958-08">
<date id="AC0A10676733A4322390F8DB5755B81D" box="[151,300,1826,1851]" pageId="14" pageNumber="515" value="1958-08">August 1958</date>
</collectingDate>
,
<collectorName id="754153716733A432223EF8DB57CEB81C" box="[313,439,1827,1850]" pageId="14" pageNumber="515">M. Kramer</collectorName>
(
<specimenCode id="88129EDC6733A43222CFF8DB541FB81D" box="[456,614,1827,1851]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="14" pageNumber="515">IBSP 137989</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="68DC3CFA6733A432217BF8DB55A9B879" ID-GBIF-Occurrence="1820860294" collectingDate="2001-05" collectionCode="IBSP" collectorName="E. K. Kashimata &amp; R. Martins" country="Brazil" county="Mogi" latitude="-23.489445" location="Manuel Ferreira" longLatPrecision="21" longitude="-46.16639" municipality="Cruzes" pageId="14" pageNumber="515" specimenCode="IBSP 56568" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo">
<specimenCount id="CEB2FD2E6733A432217BF8DB54B5B81D" box="[636,716,1826,1851]" pageId="14" pageNumber="515" type="male">1 male</specimenCount>
,
<collectingCounty id="316A4E2B6733A43221DEF8DA556FB81D" box="[729,790,1826,1851]" pageId="14" pageNumber="515">Mogi</collectingCounty>
das
<collectingMunicipality id="386FACDD6733A432204AF8DB55E7B81D" box="[845,926,1827,1851]" pageId="14" pageNumber="515">Cruzes</collectingMunicipality>
,
<location id="DD6B607C6733A43220ACF8DA521CB81D" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A43220ACF8DA521CB81D" box="[939,1125,1826,1851]" country="Brazil" county="Mogi" latitude="-23.489445" longLatPrecision="21" longitude="-46.16639" municipality="Cruzes" name="Manuel Ferreira" pageId="14" pageNumber="515" stateProvince="Sao Paulo">Manuel Ferreira</location>
,
<location id="DD6B607C6733A4322775F8DA5370B81D" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A4322775F8DA5370B81D" box="[1138,1289,1826,1851]" country="Brazil" county="Mogi" latitude="-23.489445" longLatPrecision="21" longitude="-46.16639" municipality="Cruzes" name="Birituba-Acu" pageId="14" pageNumber="515" stateProvince="Sao Paulo">Birituba-Açu</location>
,
<geoCoordinate id="BD8050606733A4322611F8DB53EEB81D" box="[1302,1431,1827,1851]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="15" value="-23.489445">23°29'22&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A4322390F8BE5758B879" box="[151,289,1862,1887]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="15" value="-46.16639">46°09'59&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F6733A4322235F8BF57D5B879" box="[306,428,1863,1887]" pageId="14" pageNumber="515" value="2001-05">
<date id="AC0A10676733A4322235F8BF57D5B879" box="[306,428,1863,1887]" pageId="14" pageNumber="515" value="2001-05">May 2001</date>
</collectingDate>
,
<collectorName id="754153716733A43222BAF8BF540CB879" box="[445,629,1862,1887]" pageId="14" pageNumber="515">E.K. Kashimata</collectorName>
&amp;
<collectorName id="754153716733A43221A4F8BF5559B879" box="[675,800,1862,1887]" pageId="14" pageNumber="515">R. Martins</collectorName>
(
<specimenCode id="88129EDC6733A4322033F8BF55B1B879" box="[820,968,1862,1887]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="14" pageNumber="515">IBSP 56568</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="68DC3CFA6733A43220E7F8BF5216B8A5" ID-GBIF-Occurrence="1820860289" collectingDate="1999-08-23" collectingDateMax="1999-08-25" collectingDateMin="1999-08-23" collectionCode="MZSP" collectorName="G. Machado" country="Brazil" latitude="-25.133617" location="Parque Estadual da Ilha do Cardoso" longLatPrecision="1" longitude="-47.96165" pageId="14" pageNumber="515" specimenCode="MZSP 21592" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo">
<specimenCount id="CEB2FD2E6733A43220E7F8BF5233B879" box="[992,1098,1862,1887]" pageId="14" pageNumber="515" type="female">1 female</specimenCount>
,
<location id="DD6B607C6733A432275DF8BF568EB8A5" LSID="urn:lsid:plazi:treatment:501D87B16733A4332390FB5454E3BD1A:DD6B607C6733A432275DF8BF568EB8A5" country="Brazil" latitude="-25.133617" longLatPrecision="1" longitude="-47.96165" name="Parque Estadual da Ilha do Cardoso" pageId="14" pageNumber="515" stateProvince="Sao Paulo">Parque Estadual da Ilha do Cardoso</location>
,
<geoCoordinate id="BD8050606733A4322205F89257F3B8A5" box="[258,394,1898,1923]" direction="south" orientation="latitude" pageId="14" pageNumber="515" precision="1" value="-25.133617">25°8'1.02&quot;S</geoCoordinate>
,
<geoCoordinate id="BD8050606733A4322292F892543BB8A5" box="[405,578,1898,1923]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="1" value="-47.96165">47°57'41.94&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F6733A432214AF8925555B8A5" box="[589,812,1898,1923]" pageId="14" pageNumber="515" value="1999-08-23" valueMax="1999-08-25" valueMin="1999-08-23">
<date id="AC0A10676733A432214AF8925555B8A5" box="[589,812,1898,1923]" pageId="14" pageNumber="515" value="1999-08-23" valueMax="1999-08-25" valueMin="1999-08-23">23–25 August 1999</date>
</collectingDate>
,
<collectorName id="754153716733A4322031F89355C4B8A5" box="[822,957,1898,1923]" pageId="14" pageNumber="515">G. Machado</collectorName>
(
<specimenCode id="88129EDC6733A43220CCF893521EB8A5" box="[971,1127,1898,1923]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="14" pageNumber="515">MZSP 21592</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="68DC3CFA6733A432277EF892525DB881" ID-GBIF-Occurrence="1820860306" collectingDate="1972-01" collectionCode="MACN" collectorName="Bordon" country="Ecuador" latitude="-0.99611115" location="Rio Napo" longLatPrecision="21" longitude="-78.42166" pageId="14" pageNumber="515" specimenCode="MACN 18958" specimenCount="1" specimenCount-male="1" stateProvince="Cotopaxi">
<emphasis id="EAC0EAB56733A432277EF89253FEB8A5" bold="true" box="[1145,1415,1898,1923]" pageId="14" pageNumber="515">
<collectingCountry id="A0A376376733A432277EF8925372B8A5" box="[1145,1291,1898,1923]" name="Ecuador" pageId="14" pageNumber="515">ECUADOR</collectingCountry>
:
<collectingRegion id="1A70F8456733A4322611F8925307B8A5" box="[1302,1406,1898,1923]" country="Ecuador" name="Cotopaxi" pageId="14" pageNumber="515">
<emphasis id="EAC0EAB56733A4322611F8925307B8A5" bold="true" box="[1302,1406,1898,1923]" italics="true" pageId="14" pageNumber="515">Cotopaxi</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6733A4322689F89356A9B881" pageId="14" pageNumber="515" type="male">1 male</specimenCount>
,
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Rio
<collectingRegion id="1A70F8456733A432220CF8775732B880" box="[267,331,1935,1958]" country="Ecuador" name="Napo" pageId="14" pageNumber="515">Napo</collectingRegion>
</location>
,
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,
<geoCoordinate id="BD8050606733A43222D2F8765427B881" box="[469,606,1934,1959]" direction="west" orientation="longitude" pageId="14" pageNumber="515" precision="15" value="-78.42166">78°25'18&quot;W</geoCoordinate>
,
<collectingDate id="BC4EE98F6733A432216DF877557AB881" box="[618,771,1935,1959]" pageId="14" pageNumber="515" value="1972-01">
<date id="AC0A10676733A432216DF877557AB881" box="[618,771,1935,1959]" pageId="14" pageNumber="515" value="1972-01">January 1972</date>
</collectingDate>
,
<collectorName id="754153716733A4322009F876551BB881" box="[782,866,1934,1959]" pageId="14" pageNumber="515">Bordon</collectorName>
(
<specimenCode id="88129EDC6733A4322076F8775260B881" box="[881,1049,1934,1959]" collectionCode="MACN" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="14" pageNumber="515">MACN 18958</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6733A43323C0F84A54A1BE66" lastPageId="15" lastPageNumber="516" pageId="14" pageNumber="515" type="diagnosis">
<paragraph id="D80B36A76733A43323C0F84A54A1BE66" blockId="14.[151,1436,1430,2031]" lastBlockId="15.[151,1437,151,573]" lastPageId="15" lastPageNumber="516" pageId="14" pageNumber="515">
<emphasis id="EAC0EAB56733A43223C0F84A5738B8ED" bold="true" box="[199,321,1970,1995]" pageId="14" pageNumber="515">Diagnosis.</emphasis>
Females of
<taxonomicName id="1FB44D246733A43222D2F84C5417B8ED" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[469,622,1970,1995]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="EAC0EAB56733A43222D2F84C5417B8ED" box="[469,622,1970,1995]" italics="true" pageId="14" pageNumber="515">E. pustulosus</emphasis>
</taxonomicName>
are very similar to those of
<taxonomicName id="1FB44D246733A43220C4F84C5232B8ED" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[963,1099,1970,1995]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="caudatus">
<emphasis id="EAC0EAB56733A43220C4F84C5232B8ED" box="[963,1099,1970,1995]" italics="true" pageId="14" pageNumber="515">E. caudatus</emphasis>
</taxonomicName>
in their cryptic bark-dweller habitus with rugose tegument and predominant dark-brown coloration. Both species also share squared leg segments and protuberant anterior femora suffused with many conical tubercles. However,
<taxonomicName id="1FB44D246732A43327B7FF61533EBF96" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[1200,1351,151,176]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="EAC0EAB56732A43327B7FF61533EBF96" box="[1200,1351,151,176]" italics="true" pageId="15" pageNumber="516">E. pustulosus</emphasis>
</taxonomicName>
can be distinguished by the yellow blister-shaped sockets on the ventral surface of femora I and II and by the anterior projection on the epigynal plate, which covers the copulatory openings entirely. Males can be easily recognized by their equal-sized, short and spiniform opisthosomal projections, by a basal indentation on the DTA and by its curvature (pointing to the dorsum of the cymbium).
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6732A43323C0FEB454F4BE43" box="[199,653,332,357]" pageId="15" pageNumber="516" type="description">
<paragraph id="D80B36A76732A43323C0FEB454F4BE43" blockId="15.[151,1437,151,573]" box="[199,653,332,357]" pageId="15" pageNumber="516">
<emphasis id="EAC0EAB56732A43323C0FEB45720BE43" bold="true" box="[199,345,332,357]" pageId="15" pageNumber="516">Description.</emphasis>
See
<bibRefCitation id="BC254B566732A4332288FEB454F0BE43" box="[399,649,332,357]" pageId="15" pageNumber="516" refId="ref18982" refString="Machado, M., Teixeira, R. A. &amp; Lise, A. A. (2015) Taxonomic notes on the crab spider genus Tobias Simon, 1895 (Araneae, Thomisidae, Stephanopinae). Zootaxa, 4034 (3), 565 - 576. https: // doi. org / 10.11646 / zootaxa. 4034.3.8" type="journal article">Machado et al. (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6732A43323C0FE975719BE8A" pageId="15" pageNumber="516" type="distribution">
<paragraph id="D80B36A76732A43323C0FE975719BE8A" blockId="15.[151,1437,151,573]" pageId="15" pageNumber="516">
<emphasis id="EAC0EAB56732A43323C0FE975719BEAE" bold="true" box="[199,352,367,392]" pageId="15" pageNumber="516">Distribution.</emphasis>
<collectingCountry id="A0A376376732A433226EFE8857AABEAE" box="[361,467,368,392]" name="Brazil" pageId="15" pageNumber="516">BRAZIL</collectingCountry>
:
<collectingRegion id="1A70F8456732A43322D8FE88542CBEAE" box="[479,597,367,392]" country="Brazil" name="Sao Paulo" pageId="15" pageNumber="516">São Paulo</collectingRegion>
, Espiírito Santo,
<collectingRegion id="1A70F8456732A433201CFE975519BEAE" box="[795,864,367,392]" country="Brazil" name="Bahia" pageId="15" pageNumber="516">Bahia</collectingRegion>
;
<collectingCountry id="A0A376376732A433206AFE885580BEAE" box="[877,1017,368,392]" name="Ecuador" pageId="15" pageNumber="516">ECUADOR</collectingCountry>
:
<collectingRegion id="1A70F8456732A4332701FE975209BEAE" box="[1030,1136,367,392]" country="Ecuador" name="Cotopaxi" pageId="15" pageNumber="516">Cotopaxi</collectingRegion>
;
<collectingRegion id="1A70F8456732A433277AFE885281BEAE" box="[1149,1272,368,392]" country="Panama" name="Panama" pageId="15" pageNumber="516">
<collectingCountry id="A0A376376732A433277AFE885281BEAE" box="[1149,1272,368,392]" name="Panama" pageId="15" pageNumber="516">PANAMA</collectingCountry>
</collectingRegion>
:
<collectingRegion id="1A70F8456732A4332602FE8853E1BEAE" box="[1285,1432,367,392]" country="Panama" name="Panama" pageId="15" pageNumber="516">
<collectingCountry id="A0A376376732A4332602FE885327BEA1" box="[1285,1374,368,391]" name="Panama" pageId="15" pageNumber="516">Panama</collectingCountry>
City
</collectingRegion>
,
<collectingRegion id="1A70F8456732A4332390FE6C56A2BE8B" box="[151,219,404,429]" country="Panama" name="Colon" pageId="15" pageNumber="516">Colón</collectingRegion>
(
<figureCitation id="408F2A226732A43323ECFE6C572BBE8A" box="[235,338,404,429]" captionStart="FIGURE 17" captionStartId="27.[151,250,1766,1788]" captionTargetBox="[151,1435,193,1736]" captionTargetId="figure@27.[151,1436,193,1744]" captionTargetPageId="27" captionText="FIGURE 17. Data of distribution records of Epicadus species compiled from Machado et al. (2015, 2017); Silva-Moreira and Machado (2016); present study." httpUri="https://zenodo.org/record/1183164/files/figure.png" pageId="15" pageNumber="516">Fig. 17A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6732A43323C0FE4F54E3BD1A" pageId="15" pageNumber="516" type="description">
<paragraph id="D80B36A76732A43323C0FE4F54E3BD1A" blockId="15.[151,1437,151,573]" pageId="15" pageNumber="516">
<emphasis id="EAC0EAB56732A43323C0FE4F5745BEF6" bold="true" box="[199,316,439,464]" pageId="15" pageNumber="516">Variation.</emphasis>
Most of the specimens examined by
<bibRefCitation id="BC254B566732A4332008FE4F5264BEF6" box="[783,1053,439,464]" pageId="15" pageNumber="516" refId="ref18982" refString="Machado, M., Teixeira, R. A. &amp; Lise, A. A. (2015) Taxonomic notes on the crab spider genus Tobias Simon, 1895 (Araneae, Thomisidae, Stephanopinae). Zootaxa, 4034 (3), 565 - 576. https: // doi. org / 10.11646 / zootaxa. 4034.3.8" type="journal article">
Machado
<emphasis id="EAC0EAB56732A4332082FE4155BDBEF6" box="[901,964,439,464]" italics="true" pageId="15" pageNumber="516">et al.</emphasis>
(2015)
</bibRefCitation>
present a predominantly brown coloration. Here, we present two additional color variations for the females of
<taxonomicName id="1FB44D246732A433271FFE2552D7BED3" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[1048,1198,476,501]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="EAC0EAB56732A433271FFE2552D7BED3" box="[1048,1198,476,501]" italics="true" pageId="15" pageNumber="516">E. pustulosus</emphasis>
</taxonomicName>
: dorsum of prosoma and opisthosoma black with front and legs yellow (
<figureCitation id="408F2A226732A43321E0FE07550FBD3E" box="[743,886,511,536]" captionStart="FIGURE 14" captionStartId="24.[151,250,1937,1959]" captionTargetBox="[181,1404,193,1921]" captionTargetId="figure@24.[181,1404,193,1921]" captionTargetPageId="24" captionText="FIGURE 14. Color variations of females. A–D Epicadus pustulosus (Mello-Leitão, 1929) (A, B MZSP 45484; C, D MZSP 51068); Scale bars: 2 mm (A, C), 1 mm (B, D). Color variations of females of E–F Epicadus taczanowskii (RoeWer, 1951): Photo credits:Kenji Nishida (E); Arthur Anker (F)." httpUri="https://zenodo.org/record/1183158/files/figure.png" pageId="15" pageNumber="516">Figs 14A, B</figureCitation>
) and brown with distal portion of femora I and dorsum of opisthosoma yellow (
<figureCitation id="408F2A226732A4332107FDDC54F4BD1A" box="[512,653,548,573]" captionStart="FIGURE 14" captionStartId="24.[151,250,1937,1959]" captionTargetBox="[181,1404,193,1921]" captionTargetId="figure@24.[181,1404,193,1921]" captionTargetPageId="24" captionText="FIGURE 14. Color variations of females. A–D Epicadus pustulosus (Mello-Leitão, 1929) (A, B MZSP 45484; C, D MZSP 51068); Scale bars: 2 mm (A, C), 1 mm (B, D). Color variations of females of E–F Epicadus taczanowskii (RoeWer, 1951): Photo credits:Kenji Nishida (E); Arthur Anker (F)." httpUri="https://zenodo.org/record/1183158/files/figure.png" pageId="15" pageNumber="516">Figs 14C, D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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