treatments-xml/data/50/1D/87/501D87B16728A4202390FD77540EBD3E.xml

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<mods:title id="2B2796C042E40207250ECEC45C569A82">There and back again: More on the taxonomy of the crab spiders genus Epicadus (Thomisidae: Stephanopinae)</mods:title>
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<mods:namePart id="95F76728116D561D06455E1EF27B9415">Teixeira, Renato Augusto</mods:namePart>
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<mods:namePart id="297DC1345FD85EFFA83D1DBFA60D61DD">Lise, Arno Antonio</mods:namePart>
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<treatment id="501D87B16728A4202390FD77540EBD3E" ID-DOI="http://doi.org/10.5281/zenodo.5951340" ID-GBIF-Taxon="141118943" ID-Zenodo-Dep="5951340" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E" httpUri="http://treatment.plazi.org/id/501D87B16728A4202390FD77540EBD3E" lastPageId="28" lastPageNumber="529" pageId="21" pageNumber="522">
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<taxonomicName id="1FB44D246728A4292390FD7754BCBD8C" ID-CoL="6FRDL" authority="(Taczanowski, 1872)" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[151,709,655,682]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A4292390FD7754BCBD8C" bold="true" box="[151,709,655,682]" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A4292390FD7757B8BD8F" bold="true" box="[151,449,655,681]" italics="true" pageId="21" pageNumber="522">Epicadus trituberculatus</emphasis>
(
<bibRefCitation id="BC254B566728A42922D6FD6854C4BD8C" box="[465,701,656,682]" pageId="21" pageNumber="522" refId="ref19570" refString="TaczanoWski, L. (1872) Les araneides de la Guyane francaise. Horae Societatis Entomologicae Rossicae, 9, 64 - 112." type="journal article">Taczanowski, 1872</bibRefCitation>
)
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</taxonomicName>
</heading>
</paragraph>
<paragraph id="D80B36A76728A4292390FD4A57C3BDED" blockId="21.[151,709,655,715]" box="[151,442,690,715]" pageId="21" pageNumber="522">
<figureCitation id="408F2A226728A4292390FD4A575ABDED" box="[151,291,690,715]" captionStart="FIGURE 12" captionStartId="22.[151,250,1876,1898]" captionTargetBox="[151,1436,733,1854]" captionTargetId="figure@22.[151,1436,733,1854]" captionTargetPageId="22" captionText="FIGURE 12. Epicadus trituberculatus (TaczanoWski, 1872), female (MPEG 022662). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A), 1 mm (B), 0.2mm (C, D)." httpUri="https://zenodo.org/record/1183154/files/figure.png" pageId="21" pageNumber="522">Figs 12A–D</figureCitation>
, 
<figureCitation id="408F2A226728A4292229FD4B57FABDED" box="[302,387,691,715]" captionStart="FIGURE 13" captionStartId="23.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1564]" captionTargetId="figure@23.[151,1436,193,1564]" captionTargetPageId="23" captionText="FIGURE 13. Epicadus trituberculatus (TaczanoWski, 1872), male (MPEG 022623). A, B habitus (A dorsal, B frontal); C, D left palp (C frontal, D retrolateral).A dorsal habitus; B front; C frontal vieW of left palp; D retrolateral vieW of left palp. Scale bars:1 mm (A), 0.5 mm (B), 0.1 mm (C, D)." httpUri="https://zenodo.org/record/1183156/files/figure.png" pageId="21" pageNumber="522">13A–D</figureCitation>
, 
<figureCitation id="408F2A226728A4292288FD4A57C3BDED" box="[399,442,690,715]" captionStart="FIGURE 16" captionStartId="26.[151,250,1869,1891]" captionTargetBox="[184,1408,193,1859]" captionTargetId="figure@26.[184,1408,193,1859]" captionTargetPageId="26" captionText="FIGURE 16. Female specimens in vivo. A Epicadus taczanowskii (RoeWer, 1951); B Epicadus dimidiaster sp. nov.; C Epicadus rubripes Mello-Leitão, 1924 (adapted from Silva-Moreira &amp; Machado 2016: fig. 1F); D Epicadus camelinus (O. Pickard-Cambridge, 1869); E Epicadus heterogaster (Guérin-Méneville, 1829); F Epicadus trituberculatus (TaczanoWski, 1872). Photo credits:César Favacho (A); Arthur Anker (B); Almir Cândido de Almeida (C); Karl Kroeker (D); Edson Roberto (E); Edelcio Muscat (F)." httpUri="https://zenodo.org/record/1183162/files/figure.png" pageId="21" pageNumber="522">16F</figureCitation>
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<taxonomicName id="1FB44D246728A4292390FD0254FFBC36" authority="TaczanoWski, 1872: 92" authorityName="TaczanoWski" authorityPageNumber="92" authorityYear="1872" box="[151,646,762,784]" class="Arachnida" family="Thomisidae" genus="Thomisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A4292390FD0257EDBC36" box="[151,404,762,784]" italics="true" pageId="21" pageNumber="522">Thomisus trituberculatus</emphasis>
<treatmentCitation id="591510B66728A429229DFD0254FFBC36" author="" box="[410,646,762,784]" page="92" pageId="21" pageNumber="522">
<bibRefCitation id="BC254B566728A429229DFD025418BC36" box="[410,609,762,784]" pageId="21" pageNumber="522" refId="ref19570" refString="TaczanoWski, L. (1872) Les araneides de la Guyane francaise. Horae Societatis Entomologicae Rossicae, 9, 64 - 112." type="journal article">TaczanoWski, 1872</bibRefCitation>
: 92
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, pl. 4, fig. 7.
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<taxonomicName id="1FB44D246728A4292390FCE1546DBC09" ID-CoL="93RB7" authority="(TaczanoWski)" authorityName="TaczanoWski" box="[151,532,793,815]" class="Arachnida" family="Thomisidae" genus="Tobias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A4292390FCE1570CBC09" box="[151,373,793,815]" italics="true" pageId="21" pageNumber="522">Tobias trituberculatus</emphasis>
(TaczanoWski)
</taxonomicName>
. 
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<bibRefCitation id="BC254B566728A429211AFCE154E2BC09" box="[541,667,793,815]" pageId="21" pageNumber="522" refId="ref19502" refString="Simon, E. (1895) Histoire naturelle des araignees. Librairie Encyclopedique de Roret, Paris, 1084 pp. https: // doi. org / 10.5962 / bhl. title. 47654" type="book">Simon 1895</bibRefCitation>
: 1047
</treatmentCitation>
, fig. 1029.
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<taxonomicName id="1FB44D246728A4292390FCC0541CBC68" authority="Mello-Leitao, 1929: 91" authorityName="Mello-Leitao" authorityPageNumber="91" authorityYear="1929" box="[151,613,824,846]" class="Arachnida" family="Thomisidae" genus="Tobias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="paraguayensis">
<emphasis id="EAC0EAB56728A4292390FCC0570ABC68" box="[151,371,824,846]" italics="true" pageId="21" pageNumber="522">Tobias paraguayensis</emphasis>
<treatmentCitation id="591510B66728A429227EFCC0541CBC68" author="" box="[377,613,824,846]" page="91" pageId="21" pageNumber="522">
<bibRefCitation id="BC254B566728A429227EFCC05438BC68" box="[377,577,824,846]" pageId="21" pageNumber="522" refId="ref19139" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Archivos do Museu Nacional do Rio de Janeiro, Rio de Janeiro, 360 pp." type="book">Mello-Leitão, 1929</bibRefCitation>
: 91
</treatmentCitation>
</taxonomicName>
, figs 191–195 (syntypes from “Paraguay”, deposited in MNHN 8790, examined).
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</paragraph>
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<treatmentCitationGroup id="F8A411896728A4292390FC8D526EBC8C" infered="true" pageId="21" pageNumber="522">
<taxonomicName id="1FB44D246728A4292390FC8D544DBCAD" ID-CoL="6G3DX" authority="Mello-Leitao, 1932: 73" authorityName="Mello-Leitao" authorityPageNumber="73" authorityYear="1932" box="[151,564,885,907]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="EAC0EAB56728A4292390FC8D5739BCAD" box="[151,320,885,907]" italics="true" pageId="21" pageNumber="522">Epicadus planus</emphasis>
<treatmentCitation id="591510B66728A4292240FC8D544DBCAD" author="" box="[327,564,885,907]" page="73" pageId="21" pageNumber="522">
<bibRefCitation id="BC254B566728A4292240FC8D5469BCAD" box="[327,528,885,907]" pageId="21" pageNumber="522" refId="ref19170" refString="Mello-Leitao, C. F. (1932) Pequenas notas arachnologicas. Boletim do Museu Nacional do Rio de Janeiro, 8, 67 - 75." type="journal article">Mello-Leitão, 1932</bibRefCitation>
: 73
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</taxonomicName>
(holotype female from Ilhéus [14°48.985'S, 39°1.996'W, Bahia, Brazil], deposited in MNRJ 14205, examined). 
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<bibRefCitation id="BC254B566728A42922D3FC6C5564BC8C" box="[468,797,916,938]" pageId="21" pageNumber="522" refId="ref19438" refString="Silva-Moreira, T. &amp; Machado, M. (2016) Taxonomic revision of the crab spider genus Epicadus Simon, 1895 (Arachnida: Araneae: Thomisidae) With notes on related genera of Stephanopinae Simon, 1895. Zootaxa, 4147 (3), 281 - 310. https: // doi. org / 10.11646 / zootaxa. 4147.3.4" type="journal article">Silva-Moreira &amp; Machado 2016</bibRefCitation>
: 297
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, figs 11, 12, 13, 14.
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<taxonomicName id="1FB44D246728A4292390FC4A54F8BCEE" ID-CoL="6FRDS" authority="Mello-Leitao, 1940: 213" authorityName="Mello-Leitao" authorityPageNumber="213" authorityYear="1940" box="[151,641,946,968]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="nigronotatus">
<emphasis id="EAC0EAB56728A4292390FC4A5704BCEE" box="[151,381,946,968]" italics="true" pageId="21" pageNumber="522">Epicadus nigronotatus</emphasis>
<treatmentCitation id="591510B66728A4292282FC4A54F8BCEE" author="" box="[389,641,946,968]" page="213" pageId="21" pageNumber="522">
<bibRefCitation id="BC254B566728A4292282FC4A5429BCEE" box="[389,592,946,968]" pageId="21" pageNumber="522" refId="ref19198" refString="Mello-Leitao, C. F. (1940) Aranhas do Espirito Santo coligidas por Mario Rosa, em 1936 e 1937. Arquivos de Zoologia do Estado de Sao Paulo, 2, 199 - 214." type="journal article">Mello-Leitão, 1940</bibRefCitation>
: 213
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</taxonomicName>
(
<typeStatus id="070F88056728A4292197FC4A5493BCEE" box="[656,746,946,968]" pageId="21" pageNumber="522" type="holotype">holotype</typeStatus>
female from 
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[
<geoCoordinate id="BD8050606728A42920F1FC4A52F5BCEE" box="[1014,1164,946,968]" direction="south" orientation="latitude" pageId="21" pageNumber="522" precision="1" value="-19.396667">19°23'48.00&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A429279EFC4A5348BCEE" box="[1177,1329,946,968]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="1" value="-40.061665">40° 3'42.00&quot;W</geoCoordinate>
, 
<collectingMunicipality id="386FACDD6728A429263AFC4A53E1BCEE" box="[1341,1432,946,968]" pageId="21" pageNumber="522">Linhares</collectingMunicipality>
, 
<collectingCountry id="A0A376376728A42923C0FC295773BCC1" box="[199,266,977,999]" name="Brazil" pageId="21" pageNumber="522">Brazil</collectingCountry>
], deposited in 
<specimenCode id="88129EDC6728A429229CFC2A545EBCC1" box="[411,551,977,999]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="21" pageNumber="522">MNRJ 58252</specimenCode>
, examined).
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<treatmentCitationGroup id="F8A411896728A4292390FC0852E5BB20" box="[151,1180,1008,1030]" pageId="21" pageNumber="522">
<taxonomicName id="1FB44D246728A4292390FC085456BB20" ID-CoL="6FRDL" authority="(TaczanoWski)" authorityName="TaczanoWski" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[151,559,1008,1030]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A4292390FC0857E8BB20" box="[151,401,1008,1030]" italics="true" pageId="21" pageNumber="522">Epicadus trituberculatus</emphasis>
(TaczanoWski)
</taxonomicName>
. 
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<bibRefCitation id="BC254B566728A429213FFC085574BB20" box="[568,781,1008,1030]" pageId="21" pageNumber="522" refId="ref19045" refString="Machado, M, Teixeira, R. A. &amp; Lise, A. A. (2017) Cladistic analysis supports the monophyly of the Neotropical crab spider genus Epicadus and its senior synonymy over Tobias (Araneae: Thomisidae). Invertebrate Systematics, 31, 442 - 455. https: // doi. org / 10.1071 / IS 16074" type="journal article">
Machado 
<emphasis id="EAC0EAB56728A429219AFC0954A9BB20" box="[669,720,1008,1030]" italics="true" pageId="21" pageNumber="522">et al.</emphasis>
2017
</bibRefCitation>
: 448
</treatmentCitation>
, figs 2A, S3A, S4B, S8E, S9A, D.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6728A42B2390FBCA5353B858" lastPageId="23" lastPageNumber="524" pageId="21" pageNumber="522" type="materials_examined">
<paragraph id="D80B36A76728A4292390FBCA55AFBB49" blockId="21.[151,1437,1074,2035]" pageId="21" pageNumber="522">
<materialsCitation id="68DC3CFA6728A4292390FBCA55AFBB49" ID-GBIF-Occurrence="1820860142" collectionCode="MZPW" collectorName="Iles du Salut &amp; Saint Laurent du Maroni" country="French Guiana" latitude="5.2833333" location="Saint Laurent du Maroni" longLatPrecision="1" longitude="-52.583332" municipality="Uassa" pageId="21" pageNumber="522" specimenCount="2" typeStatus="syntype">
<emphasis id="EAC0EAB56728A4292390FBCA57BBBB6D" bold="true" box="[151,450,1074,1099]" pageId="21" pageNumber="522">
<typeStatus id="070F88056728A4292390FBCA56ABBB6D" box="[151,210,1074,1099]" pageId="21" pageNumber="522">Type</typeStatus>
material: 
<typeStatus id="070F88056728A4292250FBCA57BBBB6D" box="[343,450,1074,1099]" pageId="21" pageNumber="522" type="syntype">Syntypes</typeStatus>
</emphasis>
(quantity not specified): 
<collectorName id="754153716728A42921F7FBCA55F1BB6D" box="[752,904,1074,1099]" pageId="21" pageNumber="522">Iles du Salut</collectorName>
, [
<geoCoordinate id="BD8050606728A4292098FBCA525ABB6D" box="[927,1059,1074,1099]" direction="north" orientation="latitude" pageId="21" pageNumber="522" precision="1" value="5.2833333">5°17.000'N</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A4292733FBCA52B1BB6D" box="[1076,1224,1074,1099]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="1" value="-52.583332">52°35.000'W</geoCoordinate>
, 
<collectorName id="754153716728A42927D0FBCA5693BB49" pageId="21" pageNumber="522">
<location id="DD6B607C6728A42927D0FBCA5693BB49" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A42927D0FBCA5693BB49" country="French Guiana" latitude="5.2833333" longLatPrecision="1" longitude="-52.583332" municipality="Uassa" name="Saint Laurent du Maroni" pageId="21" pageNumber="522">Saint Laurent du Maroni</location>
</collectorName>
and 
<collectingMunicipality id="386FACDD6728A4292227FBAF571FBB48" box="[288,358,1111,1134]" pageId="21" pageNumber="522">Uassa</collectingMunicipality>
, 
<collectingCountry id="A0A376376728A4292276FBAE5465BB49" box="[369,540,1110,1135]" name="French Guiana" pageId="21" pageNumber="522">French Guiana</collectingCountry>
], deposited in 
<collectionCode id="BEA5AE626728A42921BAFBAF5568BB48" box="[701,785,1111,1134]" country="Poland" httpUri="http://grbio.org/cool/ynr4-t34e" name="Polish Academy of Science, Museum of the Institute of Zoology" pageId="21" pageNumber="522">MZPW</collectionCode>
, presumably lost.
</materialsCitation>
</paragraph>
<paragraph id="D80B36A76728A42923C0FB8255F3B964" blockId="21.[151,1437,1074,2035]" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A42923C0FB82577FBBB5" bold="true" box="[199,262,1146,1171]" pageId="21" pageNumber="522">Note.</emphasis>
<typeStatus id="070F88056728A4292217FB835731BBB5" box="[272,328,1147,1171]" pageId="21" pageNumber="522">Type</typeStatus>
material of 
<taxonomicName id="1FB44D246728A42922DEFB8454D9BBB5" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[473,672,1146,1171]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A42922DEFB8454D9BBB5" box="[473,672,1146,1171]" italics="true" pageId="21" pageNumber="522">E. trituberculatus</emphasis>
</taxonomicName>
could not be traced despite the extensive search in European and South American arachnological and entomological collections. However, 
<taxonomicName id="1FB44D246728A42920E3FB6752D2BB91" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[996,1195,1182,1207]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A42920E3FB6752D2BB91" box="[996,1195,1182,1207]" italics="true" pageId="21" pageNumber="522">E. trituberculatus</emphasis>
</taxonomicName>
could be recognized through the examination of specimens deposited in 
<collectionCode id="BEA5AE626728A4292012FB3B5513BBFC" box="[789,874,1219,1242]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="522">MNHN</collectionCode>
and determined by 
<bibRefCitation id="BC254B566728A429276BFB3A5324BBFD" box="[1132,1373,1218,1243]" pageId="21" pageNumber="522" refId="ref19139" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Archivos do Museu Nacional do Rio de Janeiro, Rio de Janeiro, 360 pp." type="book">Mello-Leitão (1929)</bibRefCitation>
. The specimens in these vials match perfectly with the original description of the species, and besides that, 
<taxonomicName id="1FB44D246728A4292683FB1F5745BA05" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A4292683FB1F5745BA05" italics="true" pageId="21" pageNumber="522">E. trituberculatus</emphasis>
</taxonomicName>
presents a remarkable set of features, especially the median posterior opisthosomal projection ending in three equal-sized small tips (which give name to the species), that helps to identify it unequivocally. Moreover, we propose synonymy of 
<taxonomicName id="1FB44D246728A4292133FAAC54E7BA4D" authorityName="Mello-Leitao" authorityYear="1932" box="[564,670,1362,1387]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="EAC0EAB56728A4292133FAAC54E7BA4D" box="[564,670,1362,1387]" italics="true" pageId="21" pageNumber="522">E. planus</emphasis>
</taxonomicName>
based on the vouchers deposited in 
<collectionCode id="BEA5AE626728A4292733FAAB52F0BA4C" box="[1076,1161,1363,1386]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="522">MNHN</collectionCode>
that were examined and redescribed by 
<bibRefCitation id="BC254B566728A4292244FA8E5455BAA9" box="[323,556,1398,1423]" pageId="21" pageNumber="522" refId="ref19139" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Archivos do Museu Nacional do Rio de Janeiro, Rio de Janeiro, 360 pp." type="book">Mello-Leitão (1929)</bibRefCitation>
. The description of the general body coloration, and the shape and disposition of the opisthosomal projections of these specimens perfectly match the description made by 
<bibRefCitation id="BC254B566728A42927B6FA6253E1BA95" box="[1201,1432,1434,1459]" pageId="21" pageNumber="522" refId="ref19570" refString="TaczanoWski, L. (1872) Les araneides de la Guyane francaise. Horae Societatis Entomologicae Rossicae, 9, 64 - 112." type="journal article">Taczanowski (1872)</bibRefCitation>
. The trifid median posterior opisthosomal projection (the main diagnosis of 
<taxonomicName id="1FB44D246728A4292743FA47536BBAF1" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[1092,1298,1470,1495]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A4292743FA47536BBAF1" box="[1092,1298,1470,1495]" italics="true" pageId="21" pageNumber="522">E. trituberculatus</emphasis>
</taxonomicName>
) is clearly recognizable. The only species that presents striking similarities with 
<taxonomicName id="1FB44D246728A42920C6FA1C5206BADD" box="[961,1151,1506,1531]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="tritubeculatus">
<emphasis id="EAC0EAB56728A42920C6FA1C5206BADD" box="[961,1151,1506,1531]" italics="true" pageId="21" pageNumber="522">E. tritubeculatus</emphasis>
</taxonomicName>
is 
<taxonomicName id="1FB44D246728A42927A1FA1C5342BADD" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[1190,1339,1506,1531]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56728A42927A1FA1C5342BADD" box="[1190,1339,1506,1531]" italics="true" pageId="21" pageNumber="522">E. camelinus</emphasis>
</taxonomicName>
, but the large brown stains on prosoma and anterior legs of this species are constant, diagnostic and useful characters to distinguish it from 
<taxonomicName id="1FB44D246728A429226BF9D45448B965" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[364,561,1578,1603]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56728A429226BF9D45448B965" box="[364,561,1578,1603]" italics="true" pageId="21" pageNumber="522">E. trituberculatus</emphasis>
</taxonomicName>
, besides the genital structures.
</paragraph>
<paragraph id="D80B36A76728A42A23C0F9B653EABD8E" blockId="21.[151,1437,1074,2035]" lastBlockId="22.[151,1437,151,680]" lastPageId="22" lastPageNumber="523" pageId="21" pageNumber="522">
<materialsCitation id="68DC3CFA6728A42923C0F9B65401B9AD" ID-GBIF-Occurrence="1820860144" collectingDate="1992-02-01" collectingDateMax="1992-02-14" collectingDateMin="1992-02-01" collectionCode="MCTP" collectorName="A. A. Lise" country="Brazil" latitude="3.4166667" location="Ilha de Maraca" longLatPrecision="21" longitude="-61.666668" municipality="Other" pageId="21" pageNumber="522" specimenCode="MCTP 1943" specimenCount="1" specimenCount-male="1" stateProvince="Amapa">
<emphasis id="EAC0EAB56728A42923C0F9B65481B941" bold="true" box="[199,760,1614,1639]" pageId="21" pageNumber="522">
<collectingMunicipality id="386FACDD6728A42923C0F9B65776B941" box="[199,271,1614,1639]" pageId="21" pageNumber="522">Other</collectingMunicipality>
material examined. 
<collectingCountry id="A0A376376728A429211EF9B654F0B941" box="[537,649,1614,1639]" name="Brazil" pageId="21" pageNumber="522">BRAZIL</collectingCountry>
: 
<collectingRegion id="1A70F8456728A429219BF9B65496B941" box="[668,751,1614,1639]" country="Brazil" name="Amapa" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A429219BF9B65496B941" bold="true" box="[668,751,1614,1639]" italics="true" pageId="21" pageNumber="522">Amapá</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6728A4292001F9B75525B941" box="[774,860,1614,1639]" pageId="21" pageNumber="522" type="male">1 male</specimenCount>
, 
<location id="DD6B607C6728A4292068F9B65250B941" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A4292068F9B65250B941" box="[879,1065,1614,1639]" country="Brazil" latitude="3.4166667" longLatPrecision="21" longitude="-61.666668" municipality="Other" name="Ilha de Maraca" pageId="21" pageNumber="522" stateProvince="Amapa">Ilha de Maracá</location>
, 
<geoCoordinate id="BD8050606728A429273BF9B652CDB941" box="[1084,1204,1614,1639]" direction="north" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="3.4166667">3°25'00&quot;N</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A42927C0F9B65328B941" box="[1223,1361,1614,1639]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-61.666668">61°40'00&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F6728A4292663F9B7573FB9AD" pageId="21" pageNumber="522" value="1992-02-01" valueMax="1992-02-14" valueMin="1992-02-01">
<date id="AC0A10676728A4292663F9B7573FB9AD" pageId="21" pageNumber="522" value="1992-02-01" valueMax="1992-02-14" valueMin="1992-02-01">1–14 February 1992</date>
</collectingDate>
, 
<collectorName id="754153716728A429225DF98B57B6B9AD" box="[346,463,1650,1675]" pageId="21" pageNumber="522">A.A. Lise</collectorName>
(
<specimenCode id="88129EDC6728A42922D0F98A5409B9AD" box="[471,624,1650,1675]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="21" pageNumber="522">MCTP 1943</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA6728A429218BF98A5519B989" ID-GBIF-Occurrence="1820860154" collectingDate="2009-03-13" collectingDateMax="2009-04-14" collectingDateMin="2009-03-13" collectionCode="ISBP" collectorName="C. Kirsh" country="Brazil" latitude="3.2027779" location="Aldeia Hali Kato" longLatPrecision="21" longitude="-63.39167" municipality="Alto Alegre" pageId="21" pageNumber="522" specimenCode="ISBP 149628" specimenCount="1" specimenCount-female="1" stateProvince="Roraima">
<emphasis id="EAC0EAB56728A429218BF98A5483B9AD" bold="true" box="[652,762,1650,1675]" pageId="21" pageNumber="522">
<collectingRegion id="1A70F8456728A429218BF98A5488B9AD" box="[652,753,1650,1675]" country="Brazil" name="Roraima" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A429218BF98A5488B9AD" bold="true" box="[652,753,1650,1675]" italics="true" pageId="21" pageNumber="522">Roraima</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6728A429200DF98B550FB9AD" box="[778,886,1650,1675]" pageId="21" pageNumber="522" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="386FACDD6728A429208DF98A526EB9AD" box="[906,1047,1650,1675]" pageId="21" pageNumber="522">Alto Alegre</collectingMunicipality>
(
<location id="DD6B607C6728A4292729F98A537AB9AD" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A4292729F98A537AB9AD" box="[1070,1283,1650,1675]" country="Brazil" latitude="3.2027779" longLatPrecision="21" longitude="-63.39167" municipality="Alto Alegre" name="Aldeia Hali Kato" pageId="21" pageNumber="522" stateProvince="Roraima">Aldeia Hali Kato</location>
), 
<geoCoordinate id="BD8050606728A4292618F98B53EEB9AD" box="[1311,1431,1651,1675]" direction="north" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="3.2027779">3°12'10&quot;N</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A4292390F96E5758B989" box="[151,289,1686,1711]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-63.39167">63°23'30&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F6728A429222BF96F543CB989" box="[300,581,1686,1711]" pageId="21" pageNumber="522" value="2009-03-13" valueMax="2009-04-14" valueMin="2009-03-13">
<date id="AC0A10676728A429222BF96F543CB989" box="[300,581,1686,1711]" pageId="21" pageNumber="522" value="2009-03-13" valueMax="2009-04-14" valueMin="2009-03-13">13 March–14 April 2009</date>
</collectingDate>
, 
<collectorName id="754153716728A4292148F96F54D4B989" box="[591,685,1686,1711]" pageId="21" pageNumber="522">C. Kirsh</collectorName>
(
<specimenCode id="88129EDC6728A42921BBF96F5521B989" box="[700,856,1686,1711]" collectionCode="ISBP" pageId="21" pageNumber="522">ISBP 149628</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA6728A429206DF96E55CFB9F5" ID-GBIF-Occurrence="1820860231" collectingDate="1969-10-25" collectionCode="MZSP" country="Brazil" latitude="-2.4519446" location="Boca do Cuminaimiri" longLatPrecision="21" longitude="-54.6925" municipality="Santarem" pageId="21" pageNumber="522" specimenCode="MZSP 11605" specimenCount="1" specimenCount-female="1" stateProvince="Para">
<emphasis id="EAC0EAB56728A429206DF96E55D5B989" bold="true" box="[874,940,1686,1711]" pageId="21" pageNumber="522">
<collectingRegion id="1A70F8456728A429206DF96E55DBB989" box="[874,930,1686,1711]" country="Brazil" name="Para" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A429206DF96E55DBB989" bold="true" box="[874,930,1686,1711]" italics="true" pageId="21" pageNumber="522">Pará</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6728A42920B5F96F526CB989" box="[946,1045,1686,1711]" pageId="21" pageNumber="522" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="386FACDD6728A4292727F96F52F3B989" box="[1056,1162,1687,1711]" pageId="21" pageNumber="522">Santarém</collectingMunicipality>
(
<location id="DD6B607C6728A429279FF96F53E8B989" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A429279FF96F53E8B989" box="[1176,1425,1686,1711]" country="Brazil" latitude="-2.4519446" longLatPrecision="21" longitude="-54.6925" municipality="Santarem" name="Boca do Cuminaimiri" pageId="21" pageNumber="522" stateProvince="Para">Boca do Cuminaímiri</location>
), 
<geoCoordinate id="BD8050606728A4292390F9435773B9F5" box="[151,266,1723,1747]" direction="south" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="-2.4519446">2°27'07&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A429221BF94257DFB9F5" box="[284,422,1722,1747]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-54.6925">54°41'33&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F6728A42922B1F94254F9B9F5" box="[438,640,1722,1747]" pageId="21" pageNumber="522" value="1969-10-25">
<date id="AC0A10676728A42922B1F94254F9B9F5" box="[438,640,1722,1747]" pageId="21" pageNumber="522" value="1969-10-25">25 October 1969</date>
</collectingDate>
, leg. EPA (
<specimenCode id="88129EDC6728A429200AF94355D7B9F5" box="[781,942,1722,1747]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="522">MZSP 11605</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA6728A42920C1F9425480B9D1" ID-GBIF-Occurrence="1820860233" collectingDate="1992-01-08" collectionCode="MCTP" collectorName="A. A. Lise" country="Brazil" latitude="-3.1188889" location="Manaus" longLatPrecision="21" longitude="-60.02139" pageId="21" pageNumber="522" specimenCode="MCTP 2001" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<emphasis id="EAC0EAB56728A42920C1F942523FB9F5" bold="true" box="[966,1094,1722,1747]" pageId="21" pageNumber="522">
<collectingRegion id="1A70F8456728A42920C1F9425247B9F5" box="[966,1086,1722,1747]" country="Peru" name="Amazonas" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A42920C1F9425247B9F5" bold="true" box="[966,1086,1722,1747]" italics="true" pageId="21" pageNumber="522">Amazonas</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6728A4292754F94352DFB9F5" box="[1107,1190,1722,1747]" pageId="21" pageNumber="522" type="male">1 male</specimenCount>
, 
<location id="DD6B607C6728A42927B0F943536AB9F4" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A42927B0F943536AB9F4" box="[1207,1299,1723,1746]" country="Brazil" latitude="-3.1188889" longLatPrecision="21" longitude="-60.02139" name="Manaus" pageId="21" pageNumber="522" stateProvince="Amazonas">Manaus</location>
, 
<geoCoordinate id="BD8050606728A4292624F94353E1B9F5" box="[1315,1432,1723,1747]" direction="south" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="-3.1188889">3°07'08&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A4292390F9265758B9D1" box="[151,289,1758,1783]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-60.02139">60°01'17&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F6728A429222AF92757A4B9D1" box="[301,477,1759,1783]" pageId="21" pageNumber="522" value="1992-01-08">
<date id="AC0A10676728A429222AF92757A4B9D1" box="[301,477,1759,1783]" pageId="21" pageNumber="522" value="1992-01-08">8 January 1992</date>
</collectingDate>
, 
<collectorName id="754153716728A42922EEF927542EB9D1" box="[489,599,1758,1783]" pageId="21" pageNumber="522">A.A. Lise</collectorName>
(
<specimenCode id="88129EDC6728A4292158F9265488B9D1" box="[607,753,1758,1783]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="21" pageNumber="522">MCTP 2001</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA6728A4292002F926540DB83D" ID-GBIF-Occurrence="1820860238" collectingDate="2010-08-27" collectionCode="MZSP" collectorName="R. Lemos" country="Brazil" latitude="-8.802778" location="Porto Velho" longLatPrecision="21" longitude="-63.918056" pageId="21" pageNumber="522" specimenCode="MZSP 34731" specimenCount="1" specimenCount-female="1" stateProvince="Rondonia">
<emphasis id="EAC0EAB56728A4292002F9265506B9D1" bold="true" box="[773,895,1758,1783]" pageId="21" pageNumber="522">
<collectingRegion id="1A70F8456728A4292002F926550EB9D1" box="[773,887,1758,1783]" country="Brazil" name="Rondonia" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A4292002F926550EB9D1" bold="true" box="[773,887,1758,1783]" italics="true" pageId="21" pageNumber="522">Rondônia</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6728A429208FF9275595B9D1" box="[904,1004,1758,1783]" pageId="21" pageNumber="522" type="female">1 female</specimenCount>
, 
<location id="DD6B607C6728A42920FEF92752F8B9D1" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A42920FEF92752F8B9D1" box="[1017,1153,1758,1783]" country="Brazil" latitude="-8.802778" longLatPrecision="21" longitude="-63.918056" name="Porto Velho" pageId="21" pageNumber="522" stateProvince="Rondonia">Porto Velho</location>
, 
<geoCoordinate id="BD8050606728A4292789F9275378B9D1" box="[1166,1281,1759,1783]" direction="south" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="-8.802778">8°48'10&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A4292609F92653E1B9D1" box="[1294,1432,1758,1783]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-63.918056">63°55'05&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F6728A4292390F8FB5732B83D" box="[151,331,1795,1819]" pageId="21" pageNumber="522" value="2010-08-27">
<date id="AC0A10676728A4292390F8FB5732B83D" box="[151,331,1795,1819]" pageId="21" pageNumber="522" value="2010-08-27">27 August 2010</date>
</collectingDate>
, 
<collectorName id="754153716728A4292251F8FB57BBB83C" box="[342,450,1795,1818]" pageId="21" pageNumber="522">R. Lemos</collectorName>
(
<specimenCode id="88129EDC6728A42922D7F8FB5415B83D" box="[464,620,1795,1819]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="522">MZSP 34731</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA6728A4292179F8FB5690B819" ID-GBIF-Occurrence="1820860229" collectionCode="MNHN" collectorName="Mato Grosso" country="Brazil" location="Mato Grosso" pageId="21" pageNumber="522" specimenCode="MNHN 10358" specimenCount="12" specimenCount-female="4" specimenCount-male="8" stateProvince="Mato Grosso">
<emphasis id="EAC0EAB56728A4292179F8FB5562B83D" bold="true" box="[638,795,1794,1819]" pageId="21" pageNumber="522">
<collectingRegion id="1A70F8456728A4292179F8FB556BB83D" box="[638,786,1794,1819]" country="Brazil" name="Mato Grosso" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A4292179F8FB556BB83D" bold="true" box="[638,786,1794,1819]" italics="true" pageId="21" pageNumber="522">Mato Grosso</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6728A4292025F8FB5503B83D" box="[802,890,1794,1819]" pageId="21" pageNumber="522" type="male">8 males</specimenCount>
, 
<specimenCount id="CEB2FD2E6728A4292082F8FB558AB83D" box="[901,1011,1794,1819]" pageId="21" pageNumber="522" type="female">4 females</specimenCount>
, [labelled as “ 
<collectorName id="754153716728A4292789F8FB535FB83D" box="[1166,1318,1795,1819]" pageId="21" pageNumber="522">
<collectingRegion id="1A70F8456728A4292789F8FB535FB83D" box="[1166,1318,1795,1819]" country="Brazil" name="Mato Grosso" pageId="21" pageNumber="522">Mato Grosso</collectingRegion>
</collectorName>
”] (
<specimenCode id="88129EDC6728A4292642F8FB5698B819" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="522">MNHN 10358</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="68DC3CFA6728A42923F0F8DF573EB845" ID-GBIF-Occurrence="1820860226" collectingDate="2006-05-27" collectionCode="MPEG" collectorName="D. F. Candiani" country="Brazil" county="Canarana" latitude="-12.831944" location="Fazenda Tanguro" longLatPrecision="21" longitude="-52.339165" municipality="Querencia" pageId="21" pageNumber="522" specimenCode="MPEG 22621" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso">
<specimenCount id="CEB2FD2E6728A42923F0F8DF573EB819" box="[247,327,1830,1855]" pageId="21" pageNumber="522" type="male">1 male</specimenCount>
, 
<collectingCounty id="316A4E2B6728A4292253F8DF57C4B819" box="[340,445,1831,1855]" pageId="21" pageNumber="522">Canarana</collectingCounty>
/ 
<collectingMunicipality id="386FACDD6728A42922C9F8DE5438B819" box="[462,577,1830,1855]" pageId="21" pageNumber="522">Querência</collectingMunicipality>
(
<location id="DD6B607C6728A4292155F8DE5562B819" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A4292155F8DE5562B819" box="[594,795,1830,1855]" country="Brazil" county="Canarana" latitude="-12.831944" longLatPrecision="21" longitude="-52.339165" municipality="Querencia" name="Fazenda Tanguro" pageId="21" pageNumber="522" stateProvince="Mato Grosso">Fazenda Tanguro</location>
), 
<geoCoordinate id="BD8050606728A4292037F8DE55C9B819" box="[816,944,1830,1855]" direction="south" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="-12.831944">12°49'55''S</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A42920B9F8DE5230B819" box="[958,1097,1830,1855]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-52.339165">52°20'21&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F6728A4292752F8DF528BB819" box="[1109,1266,1830,1855]" pageId="21" pageNumber="522" value="2006-05-27">
<date id="AC0A10676728A4292752F8DF528BB819" box="[1109,1266,1830,1855]" pageId="21" pageNumber="522" value="2006-05-27">27 May 2006</date>
</collectingDate>
, 
<collectorName id="754153716728A42927F8F8DF53E5B819" box="[1279,1436,1830,1855]" pageId="21" pageNumber="522">D.F. Candiani</collectorName>
(
<specimenCode id="88129EDC6728A4292399F8B35739B845" box="[158,320,1866,1891]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="522">MPEG 22621</specimenCode>
)
</materialsCitation>
; 1 male, 
<date id="AC0A10676728A42922ABF8B3544AB845" box="[428,563,1866,1891]" pageId="21" pageNumber="522" value="2006-06-08">8 June 2006</date>
(MPEG 22623); 1 male, 
<date id="AC0A10676728A4292048F8B3559DB845" box="[847,996,1866,1891]" pageId="21" pageNumber="522" value="2006-06-11">11 June 2006</date>
(MPEG 22625); 1 male, 
<date id="AC0A10676728A4292607F8B253E1B845" box="[1280,1432,1866,1891]" pageId="21" pageNumber="522" value="2006-06-15">15 June 2006</date>
, N.F. Lo-Man-Hung (MPEG 22627); 1 female, 
<date id="AC0A10676728A42921B5F8965535B8A1" box="[690,844,1902,1927]" pageId="21" pageNumber="522" value="2006-06-16">16 June 2006</date>
(MPEG 22662); 
<materialsCitation id="68DC3CFA6728A4292710F8975266B88D" ID-GBIF-Occurrence="1820860220" collectingDate="1992-07-15" collectingDateMax="1992-07-26" collectingDateMin="1992-07-15" collectionCode="MCTP" collectorName="A. A. Lise &amp; A. Braul" country="Brazil" latitude="-15.358056" location="Guimaraes" longLatPrecision="21" longitude="-55.89778" municipality="Chapada" pageId="21" pageNumber="522" specimenCode="MCTP 2192" specimenCount="4" specimenCount-male="4" stateProvince="Mato Grosso">
<specimenCount id="CEB2FD2E6728A4292710F897520BB8A1" box="[1047,1138,1902,1927]" pageId="21" pageNumber="522" type="male">4 males</specimenCount>
, 
<collectingMunicipality id="386FACDD6728A4292787F896529BB8A1" box="[1152,1250,1902,1927]" pageId="21" pageNumber="522">Chapada</collectingMunicipality>
dos 
<location id="DD6B607C6728A429261BF89653E1B8A1" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A429261BF89653E1B8A1" box="[1308,1432,1902,1927]" country="Brazil" latitude="-15.358056" longLatPrecision="21" longitude="-55.89778" municipality="Chapada" name="Guimaraes" pageId="21" pageNumber="522" stateProvince="Mato Grosso">Guimarães</location>
, 
<geoCoordinate id="BD8050606728A4292390F86A5760B88D" box="[151,281,1938,1963]" direction="south" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="-15.358056">15°21'29&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A4292222F86A57D6B88D" box="[293,431,1938,1963]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-55.89778">55°53'52&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F6728A42922BCF86A5402B88D" box="[443,635,1938,1963]" pageId="21" pageNumber="522" value="1992-07-15" valueMax="1992-07-26" valueMin="1992-07-15">
<date id="AC0A10676728A42922BCF86A5402B88D" box="[443,635,1938,1963]" pageId="21" pageNumber="522" value="1992-07-15" valueMax="1992-07-26" valueMin="1992-07-15">15–26 July 1992</date>
</collectingDate>
, 
<collectorName id="754153716728A4292180F86B548CB88D" box="[647,757,1938,1963]" pageId="21" pageNumber="522">A.A. Lise</collectorName>
&amp; 
<collectorName id="754153716728A429201CF86B5505B88D" box="[795,892,1938,1963]" pageId="21" pageNumber="522">A. Braul</collectorName>
(
<specimenCode id="88129EDC6728A4292083F86A526EB88D" box="[900,1047,1938,1963]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="21" pageNumber="522">MCTP 2192</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="68DC3CFA6728A429272CF86B5509B8E9" ID-GBIF-Occurrence="1820860166" collectingDate="1992-03-01" collectionCode="MCTP" collectorName="A. Braul" country="Brazil" latitude="-16.266388" location="Porto Cercado" longLatPrecision="21" longitude="-56.62639" municipality="Pocone" pageId="21" pageNumber="522" specimenCode="MCTP 2482" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso">
<specimenCount id="CEB2FD2E6728A429272CF86B5203B88D" box="[1067,1146,1938,1963]" pageId="21" pageNumber="522" type="male">1 male</specimenCount>
, 
<collectingMunicipality id="386FACDD6728A4292781F86B52A1B88D" box="[1158,1240,1939,1963]" pageId="21" pageNumber="522">Poconé</collectingMunicipality>
(
<location id="DD6B607C6728A42927EFF86B53E8B88D" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C6728A42927EFF86B53E8B88D" box="[1256,1425,1938,1963]" country="Brazil" latitude="-16.266388" longLatPrecision="21" longitude="-56.62639" municipality="Pocone" name="Porto Cercado" pageId="21" pageNumber="522" stateProvince="Mato Grosso">Porto Cercado</location>
), 
<geoCoordinate id="BD8050606728A4292390F84E5760B8E9" box="[151,281,1974,1999]" direction="south" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="-16.266388">16°15'59&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A4292223F84E57D6B8E9" box="[292,431,1974,1999]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-56.62639">56°37'35&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F6728A42922BDF84F5422B8E9" box="[442,603,1974,1999]" pageId="21" pageNumber="522" value="1992-03-01">
<date id="AC0A10676728A42922BDF84F5422B8E9" box="[442,603,1974,1999]" pageId="21" pageNumber="522" value="1992-03-01">1 March 1992</date>
</collectingDate>
, 
<collectorName id="754153716728A4292161F84F54BEB8E9" box="[614,711,1974,1999]" pageId="21" pageNumber="522">A. Braul</collectorName>
(
<specimenCode id="88129EDC6728A42921D1F84F5510B8E9" box="[726,873,1975,1999]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="21" pageNumber="522">MCTP 2482</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA6728A42A207CF84F574CBF96" ID-GBIF-Occurrence="1820860143" collectionCode="IBSP" collectorName="Equipe" country="Brazil" lastPageId="22" lastPageNumber="523" latitude="-23.4675" location="Mato Grosso do Sul" longLatPrecision="21" longitude="-46.699444" pageId="21" pageNumber="522" specimenCode="IBSP 30355, IBSP 35634" specimenCount="2" specimenCount-male="2" stateProvince="Mato Grosso do Sul">
<emphasis id="EAC0EAB56728A429207CF84F5212B8E9" bold="true" box="[891,1131,1974,1999]" pageId="21" pageNumber="522">
<collectingRegion id="1A70F8456728A429207CF84F521BB8E9" box="[891,1122,1974,1999]" country="Brazil" name="Mato Grosso do Sul" pageId="21" pageNumber="522">
<emphasis id="EAC0EAB56728A429207CF84F521BB8E9" bold="true" box="[891,1122,1974,1999]" italics="true" pageId="21" pageNumber="522">
<collectingRegion id="1A70F8456728A429207CF84F5269B8E9" box="[891,1040,1974,1999]" inferredBySuffix="true" pageId="21" pageNumber="522">Mato Grosso</collectingRegion>
do Sul
</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E6728A4292774F84F52B8B8E9" box="[1139,1217,1974,1999]" pageId="21" pageNumber="522" type="male">1 male</specimenCount>
, Brasilândia, Usina Hidrelétrica Engenheiro Sérgio Mota, 
<geoCoordinate id="BD8050606728A4292151F82354AEB8D5" box="[598,727,2011,2035]" direction="south" orientation="latitude" pageId="21" pageNumber="522" precision="15" value="-23.4675">23°28'03&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD8050606728A42921E0F822550BB8D5" box="[743,882,2010,2035]" direction="west" orientation="longitude" pageId="21" pageNumber="522" precision="15" value="-46.699444">46°41'58&quot;W</geoCoordinate>
, 2000, leg. 
<collectorName id="754153716728A4292707F8225236B8D5" box="[1024,1103,2010,2035]" pageId="21" pageNumber="522">Equipe</collectorName>
<collectionCode id="BEA5AE626728A429275CF82352EFB8D5" box="[1115,1174,2011,2035]" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="21" pageNumber="522">IBSP</collectionCode>
(
<specimenCode id="88129EDC6728A42927ADF8235340B8D5" box="[1194,1337,2010,2035]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="21" pageNumber="522">IBSP 30355</specimenCode>
) 
<specimenCount id="CEB2FD2E6728A429264BF82353E5B8D5" box="[1356,1436,2010,2035]" pageId="21" pageNumber="522" type="male">1 male</specimenCount>
(
<specimenCode id="88129EDC672BA42A2399FF605754BF96" box="[158,301,151,176]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="523">IBSP 35634</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA672BA42A2238FF6F53E1BF96" ID-GBIF-Occurrence="1820860145" box="[319,1432,151,176]" collectingDate="1998-04-12" collectionCode="IBSP" collectorName="A. D. Brescovit" country="Brazil" latitude="-14.82" location="Ilheus" longLatPrecision="21" longitude="-39.332775" pageId="22" pageNumber="523" specimenCode="IBSP 19058" specimenCount="1" specimenCount-male="1" stateProvince="Bahia">
<emphasis id="EAC0EAB5672BA42A2238FF6F57F7BF96" bold="true" box="[319,398,151,176]" pageId="22" pageNumber="523">
<collectingRegion id="1A70F845672BA42A2238FF6F57FCBF96" box="[319,389,151,176]" country="Brazil" name="Bahia" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A2238FF6F57FCBF96" bold="true" box="[319,389,151,176]" italics="true" pageId="22" pageNumber="523">Bahia</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E672BA42A2292FF60579ABF96" box="[405,483,151,176]" pageId="22" pageNumber="523" type="male">1 male</specimenCount>
, 
<location id="DD6B607C672BA42A22E9FF6F544CBF96" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A22E9FF6F544CBF96" box="[494,565,151,176]" country="Brazil" latitude="-14.82" longLatPrecision="21" longitude="-39.332775" name="Ilheus" pageId="22" pageNumber="523" stateProvince="Bahia">Ilhéus</location>
, 
<geoCoordinate id="BD805060672BA42A2147FF6054B8BF96" box="[576,705,152,176]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-14.82">14°49'12&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A21CAFF6F552EBF96" box="[717,855,151,176]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-39.332775">39°19'58&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A2065FF60527BBF96" box="[866,1026,151,176]" pageId="22" pageNumber="523" value="1998-04-12">
<date id="AC0A1067672BA42A2065FF60527BBF96" box="[866,1026,151,176]" pageId="22" pageNumber="523" value="1998-04-12">12 April 1998</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A270AFF6052CCBF96" box="[1037,1205,151,176]" pageId="22" pageNumber="523">A.D. Brescovit</collectorName>
<emphasis id="EAC0EAB5672BA42A27BBFF61528DBF96" box="[1212,1268,151,176]" italics="true" pageId="22" pageNumber="523">et al.</emphasis>
(
<specimenCode id="88129EDC672BA42A2605FF6053E9BF96" box="[1282,1424,151,176]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="523">IBSP 19058</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA672BA42A2390FF445206BFF3" ID-GBIF-Occurrence="1820860149" box="[151,1151,188,213]" collectingDate="1942-06-18" collectionCode="MZSP" collectorName="F. Jaime" country="Brazil" latitude="-16.686666" location="Goiania" longLatPrecision="21" longitude="-49.26472" pageId="22" pageNumber="523" specimenCode="MZSP 386" specimenCount="1" specimenCount-male="1" stateProvince="Goias">
<emphasis id="EAC0EAB5672BA42A2390FF44569ABFF3" bold="true" box="[151,227,188,213]" pageId="22" pageNumber="523">
<collectingRegion id="1A70F845672BA42A2390FF4456A3BFF3" box="[151,218,188,213]" country="Brazil" name="Goias" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A2390FF4456A3BFF3" bold="true" box="[151,218,188,213]" italics="true" pageId="22" pageNumber="523">Goiás</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E672BA42A23EBFF445744BFF3" box="[236,317,188,213]" pageId="22" pageNumber="523" type="male">1 male</specimenCount>
, 
<location id="DD6B607C672BA42A224DFF4457DFBFF3" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A224DFF4457DFBFF3" box="[330,422,188,213]" country="Brazil" latitude="-16.686666" longLatPrecision="21" longitude="-49.26472" name="Goiania" pageId="22" pageNumber="523" stateProvince="Goias">Goiânia</location>
, 
<geoCoordinate id="BD805060672BA42A22B4FF44544CBFF3" box="[435,565,188,213]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-16.686666">16°41'12&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A2144FF4454B4BFF3" box="[579,717,188,213]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-49.26472">49°15'53&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A21DCFF445501BFF2" box="[731,888,188,212]" pageId="22" pageNumber="523" value="1942-06-18">
<date id="AC0A1067672BA42A21DCFF445501BFF2" box="[731,888,188,212]" pageId="22" pageNumber="523" value="1942-06-18">18 June 1942</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A2082FF45559DBFF3" box="[901,996,188,213]" pageId="22" pageNumber="523">F. Jaime</collectorName>
(
<specimenCode id="88129EDC672BA42A20F2FF44520EBFF3" box="[1013,1143,188,213]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="523">MZSP 386</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA672BA42A278BFF44579FBE3A" ID-GBIF-Occurrence="1820860159" collectingDate="1997-07-19" collectingDateMax="1997-07-25" collectingDateMin="1997-07-19" collectionCode="IBSP" collectorName="A. D. Brescovit" country="Brazil" latitude="-19.151388" location="Reserva Floresta da Cia. Vale do Rio Doce" longLatPrecision="21" longitude="-40.070553" municipality="Sao Mateus" pageId="22" pageNumber="523" specimenCode="IBSP 12893" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<emphasis id="EAC0EAB5672BA42A278BFF445344BFF3" bold="true" box="[1164,1341,188,213]" pageId="22" pageNumber="523">
<collectingRegion id="1A70F845672BA42A278BFF44534CBFF3" box="[1164,1333,188,213]" country="Brazil" name="Espirito Santo" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A278BFF44534CBFF3" bold="true" box="[1164,1333,188,213]" italics="true" pageId="22" pageNumber="523">Espírito Santo</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E672BA42A2640FF4453EEBFF3" box="[1351,1431,188,213]" pageId="22" pageNumber="523" type="male">1 male</specimenCount>
, 
<collectingMunicipality id="386FACDD672BA42A2390FF18575ABFD1" box="[151,291,224,248]" pageId="22" pageNumber="523">São Mateus</collectingMunicipality>
, 
<location id="DD6B607C672BA42A2231FF18553EBFD1" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A2231FF18553EBFD1" box="[310,839,223,248]" country="Brazil" latitude="-19.151388" longLatPrecision="21" longitude="-40.070553" municipality="Sao Mateus" name="Reserva Floresta da Cia. Vale do Rio Doce" pageId="22" pageNumber="523" stateProvince="Espirito Santo">Reserva Floresta da Cia. Vale do Rio Doce</location>
, 
<geoCoordinate id="BD805060672BA42A205EFF2755A3BFDE" box="[857,986,223,248]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-19.151388">19°09'05&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A20EAFF18520FBFDE" box="[1005,1142,224,248]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-40.070553">40°04'14&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A278EFF27532DBFDE" box="[1161,1364,223,248]" pageId="22" pageNumber="523" value="1997-07-19" valueMax="1997-07-25" valueMin="1997-07-19">
<date id="AC0A1067672BA42A278EFF27532DBFDE" box="[1161,1364,223,248]" pageId="22" pageNumber="523" value="1997-07-19" valueMax="1997-07-25" valueMin="1997-07-19">19–25 July 1997</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A2661FF18577ABE3B" pageId="22" pageNumber="523">A.D. Brescovit</collectorName>
<emphasis id="EAC0EAB5672BA42A220EFEFD5738BE3B" box="[265,321,260,285]" italics="true" pageId="22" pageNumber="523">et al.</emphasis>
(
<specimenCode id="88129EDC672BA42A2248FEFC57A4BE3A" box="[335,477,260,284]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="523">IBSP 12893</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="68DC3CFA672BA42A22F6FEFC5287BE3A" ID-GBIF-Occurrence="1820860146" box="[497,1278,260,285]" collectingDate="1998-01-05" collectingDateMax="1998-01-12" collectingDateMin="1998-01-05" collectionCode="IBSP" collectorName="A. D. Brescovit" country="Brazil" latitude="-19.151388" location="Reserva Floresta da Cia. Vale do Rio Doce" longLatPrecision="21" longitude="-40.070553" municipality="Sao Mateus" pageId="22" pageNumber="523" specimenCode="IBSP 24328" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="CEB2FD2E672BA42A22F6FEFC5446BE3B" box="[497,575,260,285]" pageId="22" pageNumber="523" type="male">1 male</specimenCount>
, same locality and collector, 
<collectingDate id="BC4EE98F672BA42A2083FEFC5220BE3A" box="[900,1113,260,285]" pageId="22" pageNumber="523" value="1998-01-05" valueMax="1998-01-12" valueMin="1998-01-05">
<date id="AC0A1067672BA42A2083FEFC5220BE3A" box="[900,1113,260,285]" pageId="22" pageNumber="523" value="1998-01-05" valueMax="1998-01-12" valueMin="1998-01-05">5–12 January 1998</date>
</collectingDate>
(
<specimenCode id="88129EDC672BA42A276FFEFC528FBE3A" box="[1128,1270,260,284]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="523">IBSP 24328</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="68DC3CFA672BA42A260EFEFC5568BE66" ID-GBIF-Occurrence="1820860242" collectingDate="2000-02-07" collectingDateMax="2000-02-14" collectingDateMin="2000-02-07" collectionCode="IBSP" collectorName="G. Schwartz" country="Brazil" latitude="-19.151388" location="Reserva Floresta da Cia. Vale do Rio Doce" longLatPrecision="21" longitude="-40.070553" municipality="Sao Mateus" pageId="22" pageNumber="523" specimenCode="IBSP 37035" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="CEB2FD2E672BA42A260EFEFC532EBE3B" box="[1289,1367,260,285]" pageId="22" pageNumber="523" type="male">1 male</specimenCount>
, same locality, 
<collectingDate id="BC4EE98F672BA42A23F1FED057A5BE66" box="[246,476,295,320]" pageId="22" pageNumber="523" value="2000-02-07" valueMax="2000-02-14" valueMin="2000-02-07">
<date id="AC0A1067672BA42A23F1FED057A5BE66" box="[246,476,295,320]" pageId="22" pageNumber="523" value="2000-02-07" valueMax="2000-02-14" valueMin="2000-02-07">7–14 February 2000</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A22E1FED05414BE66" box="[486,621,295,320]" pageId="22" pageNumber="523">G. Schwartz</collectorName>
(
<specimenCode id="88129EDC672BA42A217CFED05570BE66" box="[635,777,295,320]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="523">IBSP 37035</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="68DC3CFA672BA42A201BFED0540FBE43" ID-GBIF-Occurrence="1820860251" collectingDate="1942-09-22" collectionCode="MZSP" collectorName="Svans" country="Brazil" latitude="-19.314999" location="Rio Sao Jose" longLatPrecision="21" longitude="-40.117775" municipality="Linhares" pageId="22" pageNumber="523" specimenCode="MZSP 7654" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="CEB2FD2E672BA42A201BFED05510BE66" box="[796,873,295,320]" pageId="22" pageNumber="523" type="male">1 male</specimenCount>
, 
<collectingMunicipality id="386FACDD672BA42A2073FEDF55A0BE66" box="[884,985,295,320]" pageId="22" pageNumber="523">Linhares</collectingMunicipality>
, 
<location id="DD6B607C672BA42A20E3FEDF520FBE66" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A20E3FEDF520FBE66" box="[996,1142,295,320]" country="Brazil" latitude="-19.314999" longLatPrecision="21" longitude="-40.117775" municipality="Linhares" name="Rio Sao Jose" pageId="22" pageNumber="523" stateProvince="Espirito Santo">Rio São José</location>
, 
<geoCoordinate id="BD805060672BA42A2786FEDF537ABE66" box="[1153,1283,295,320]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-19.314999">19°18'54&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A2609FED053E1BE66" box="[1294,1432,296,320]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-40.117775">40°07'04&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A2390FEB45703BE42" box="[151,378,332,357]" pageId="22" pageNumber="523" value="1942-09-22">
<date id="AC0A1067672BA42A2390FEB45703BE42" box="[151,378,332,357]" pageId="22" pageNumber="523" value="1942-09-22">22 September 1942</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A228FFEB457B2BE42" box="[392,459,332,356]" pageId="22" pageNumber="523">Svans</collectorName>
(
<specimenCode id="88129EDC672BA42A22DBFEB45416BE43" box="[476,623,332,357]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="523">MZSP 7654</specimenCode>
)
</materialsCitation>
; 1 male, 
<date id="AC0A1067672BA42A21E2FEB455BCBE42" box="[741,965,332,357]" pageId="22" pageNumber="523" value="1942-09-21">21 September 1942</date>
(MZSP 8594). 
<materialsCitation id="68DC3CFA672BA42A277AFEB45690BE8A" ID-GBIF-Occurrence="1820860239" collectingDate="2011-07-11" collectingDateMax="2011-07-25" collectingDateMin="2011-07-11" collectionCode="UFMG" collectorName="G. H. F. Azevedo" country="Brazil" latitude="-19.716112" location="Parque Estadual do Rio Doce" longLatPrecision="21" longitude="-42.733612" municipality="Marlieria" pageId="22" pageNumber="523" specimenCode="UFMG 10873" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais">
<emphasis id="EAC0EAB5672BA42A277AFEB4535FBE43" bold="true" box="[1149,1318,332,357]" pageId="22" pageNumber="523">
<collectingRegion id="1A70F845672BA42A277AFEB45364BE43" box="[1149,1309,332,357]" country="Brazil" name="Minas Gerais" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A277AFEB45364BE43" bold="true" box="[1149,1309,332,357]" italics="true" pageId="22" pageNumber="523">Minas Gerais</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E672BA42A2636FEB453E1BE43" box="[1329,1432,332,357]" pageId="22" pageNumber="523" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="386FACDD672BA42A2390FE97577BBEAE" box="[151,258,367,392]" pageId="22" pageNumber="523">Marliéria</collectingMunicipality>
, 
<location id="DD6B607C672BA42A2215FE885409BEA1" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A2215FE885409BEA1" box="[274,624,367,392]" country="Brazil" latitude="-19.716112" longLatPrecision="21" longitude="-42.733612" municipality="Marlieria" name="Parque Estadual do Rio Doce" pageId="22" pageNumber="523" stateProvince="Minas Gerais">Parque Estadual do Rio Doce</location>
, 
<geoCoordinate id="BD805060672BA42A2187FE975578BEAE" box="[640,769,367,392]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-19.716112">19°42'58&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A2015FE8855E2BEAE" box="[786,923,368,392]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-42.733612">42°44'01&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A20ABFE975209BEAE" box="[940,1136,367,392]" pageId="22" pageNumber="523" value="2011-07-11" valueMax="2011-07-25" valueMin="2011-07-11">
<date id="AC0A1067672BA42A20ABFE975209BEAE" box="[940,1136,367,392]" pageId="22" pageNumber="523" value="2011-07-11" valueMax="2011-07-25" valueMin="2011-07-11">11–25 July 2011</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A2787FE88534EBEAE" box="[1152,1335,367,392]" pageId="22" pageNumber="523">G.H.F. Azevedo</collectorName>
(
<specimenCode id="88129EDC672BA42A264EFE88569BBE8A" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="22" pageNumber="523">UFMG 10873</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA672BA42A23FBFE6C525BBEF6" ID-GBIF-Occurrence="1820860234" collectingDate="1999-01-16" collectingDateMax="1999-02-13" collectingDateMin="1999-01-16" collectionCode="IBSP" collectorName="Equipe" country="Brazil" latitude="-21.764723" location="Sao Paulo" longLatPrecision="21" longitude="-52.105278" pageId="22" pageNumber="523" specimenCode="IBSP 23240" specimenCount="2" specimenCount-male="2" stateProvince="Sao Paulo">
<emphasis id="EAC0EAB5672BA42A23FBFE6C57FABE8B" bold="true" box="[252,387,404,429]" pageId="22" pageNumber="523">
<collectingRegion id="1A70F845672BA42A23FBFE6C5700BE8B" box="[252,377,404,429]" country="Brazil" name="Sao Paulo" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A23FBFE6C5700BE8B" bold="true" box="[252,377,404,429]" italics="true" pageId="22" pageNumber="523">São Paulo</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E672BA42A2295FE6C578BBE8B" box="[402,498,404,429]" pageId="22" pageNumber="523" type="male">2 males</specimenCount>
, Presidente Epitácio, Usina Hidrelétrica Engenheiro Sérgio Mota, 
<geoCoordinate id="BD805060672BA42A2611FE6C53EEBE8B" box="[1302,1431,404,429]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-21.764723">21°45'53&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A2390FE4F5758BEF6" box="[151,289,439,464]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-52.105278">52°06'19&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A2235FE4F54EDBEF6" box="[306,660,439,464]" pageId="22" pageNumber="523" value="1999-01-16" valueMax="1999-02-13" valueMin="1999-01-16">
<date id="AC0A1067672BA42A2235FE4F54EDBEF6" box="[306,660,439,464]" pageId="22" pageNumber="523" value="1999-01-16" valueMax="1999-02-13" valueMin="1999-01-16">16 January–13 February 1999</date>
</collectingDate>
, leg. 
<collectorName id="75415371672BA42A21DDFE4F5550BEF6" box="[730,809,439,464]" pageId="22" pageNumber="523">Equipe</collectorName>
<collectionCode id="BEA5AE62672BA42A2031FE405508BEF6" box="[822,881,440,464]" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="523">IBSP</collectionCode>
(
<specimenCode id="88129EDC672BA42A2081FE405263BEF6" box="[902,1050,440,464]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="523">IBSP 23240</specimenCode>
)
</materialsCitation>
; 1 male (IBSP 23327); 1 male, Piracicaba, Campus ESALQ, 22°43'47&quot;S, 47°39'29&quot;W, 
<date id="AC0A1067672BA42A201CFE2455A0BED3" box="[795,985,476,501]" pageId="22" pageNumber="523" value="1946-08-19">19 August 1946</date>
, A. Correa (IBSP 138001). 
<materialsCitation id="68DC3CFA672BA42A2623FE24569FBD3E" ID-GBIF-Occurrence="1820860222" country="Brazil" location="Parana" pageId="22" pageNumber="523" specimenCount="1" specimenCount-female="1" stateProvince="Parana">
<emphasis id="EAC0EAB5672BA42A2623FE2453FABED3" bold="true" box="[1316,1411,476,501]" pageId="22" pageNumber="523">
<collectingRegion id="1A70F845672BA42A2623FE245300BED3" box="[1316,1401,476,501]" country="Brazil" name="Parana" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A2623FE245300BED3" bold="true" box="[1316,1401,476,501]" italics="true" pageId="22" pageNumber="523">Paraná</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E672BA42A2689FE24569FBD3E" pageId="22" pageNumber="523" type="female">1 female</specimenCount>
</materialsCitation>
, 
<materialsCitation id="68DC3CFA672BA42A23F6FE07537DBD3E" ID-GBIF-Occurrence="1820860241" box="[241,1284,511,536]" collectingDate="1987-11-23" collectionCode="MCTP" collectorName="A. D. Brescovit" country="Brazil" latitude="-23.470278" location="Jundiai do Sul" longLatPrecision="21" longitude="-50.224167" pageId="22" pageNumber="523" specimenCode="MCTP 38706" specimenCount="1" stateProvince="Jundiai do Sul">
<collectingRegion id="1A70F845672BA42A23F6FE0757EABD3E" box="[241,403,511,536]" inferredBySuffix="true" pageId="22" pageNumber="523">Jundiaí do Sul</collectingRegion>
, 
<geoCoordinate id="BD805060672BA42A2299FDF85459BD3E" box="[414,544,512,536]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-23.470278">23°28'13&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A212CFE0754CCBD3E" box="[555,693,511,536]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-50.224167">50°13'27&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A21C7FDF855E2BD3E" box="[704,923,511,536]" pageId="22" pageNumber="523" value="1987-11-23">
<date id="AC0A1067672BA42A21C7FDF855E2BD3E" box="[704,923,511,536]" pageId="22" pageNumber="523" value="1987-11-23">23 November 1987</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A20A2FDF85234BD3E" box="[933,1101,511,536]" pageId="22" pageNumber="523">A.D. Brescovit</collectorName>
(
<specimenCode id="88129EDC672BA42A275BFDF85285BD3E" box="[1116,1276,511,536]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="22" pageNumber="523">MCTP 38706</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="68DC3CFA672BA42A2609FDF853E0BD1A" ID-GBIF-Occurrence="1820860267" collectingDate="1993-03-29" collectionCode="MCTP" collectorName="A. D. Brescovit" country="Brazil" latitude="-25.46028" location="Parque Nacional do Iguacu" longLatPrecision="21" longitude="-53.817776" municipality="Foz do Iguacu" pageId="22" pageNumber="523" specimenCode="MCTP 4322" specimenCount="2" specimenCount-male="2" stateProvince="Jundiai do Sul">
<specimenCount id="CEB2FD2E672BA42A2609FDF8531EBD3E" box="[1294,1383,511,536]" pageId="22" pageNumber="523" type="male">2 males</specimenCount>
, 
<collectingMunicipality id="386FACDD672BA42A2675FDF85772BD1B" pageId="22" pageNumber="523">Foz do Iguaçu</collectingMunicipality>
, 
<location id="DD6B607C672BA42A221FFDDD5429BD1B" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A221FFDDD5429BD1B" box="[280,592,548,573]" country="Brazil" latitude="-25.46028" longLatPrecision="21" longitude="-53.817776" municipality="Foz do Iguacu" name="Parque Nacional do Iguacu" pageId="22" pageNumber="523" stateProvince="Jundiai do Sul">Parque Nacional do Iguaçu</location>
, 
<geoCoordinate id="BD805060672BA42A215AFDDC54A7BD1B" box="[605,734,548,573]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-25.46028">25°27'37&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A21EBFDDC550FBD1B" box="[748,886,548,573]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-53.817776">53°49'04&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A2084FDDC524CBD1A" box="[899,1077,548,573]" pageId="22" pageNumber="523" value="1993-03-29">
<date id="AC0A1067672BA42A2084FDDC524CBD1A" box="[899,1077,548,573]" pageId="22" pageNumber="523" value="1993-03-29">29 March 1993</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A2745FDDC5294BD1B" box="[1090,1261,548,573]" pageId="22" pageNumber="523">A.D. Brescovit</collectorName>
(
<specimenCode id="88129EDC672BA42A27FAFDDC53E8BD1A" box="[1277,1425,548,572]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="22" pageNumber="523">MCTP 4322</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA672BA42A2390FDBF531CBD46" ID-GBIF-Occurrence="1820860175" box="[151,1381,583,608]" collectingDate="1998-08-08" collectionCode="MCTP" collectorName="P. P. Marques" country="Brazil" latitude="-27.10389" location="Mondai" longLatPrecision="20" longitude="-53.444447" pageId="22" pageNumber="523" specimenCode="MCTP 10313" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina">
<emphasis id="EAC0EAB5672BA42A2390FDBF572DBD46" bold="true" box="[151,340,583,608]" pageId="22" pageNumber="523">
<collectingRegion id="1A70F845672BA42A2390FDBF5733BD46" box="[151,330,583,608]" country="Brazil" name="Santa Catarina" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A2390FDBF5733BD46" bold="true" box="[151,330,583,608]" italics="true" pageId="22" pageNumber="523">Santa Catarina</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E672BA42A2258FDB057C8BD46" box="[351,433,583,608]" pageId="22" pageNumber="523" type="male">1 male</specimenCount>
, 
<location id="DD6B607C672BA42A22C7FDBF5463BD46" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A22C7FDBF5463BD46" box="[448,538,583,608]" country="Brazil" latitude="-27.10389" longLatPrecision="20" longitude="-53.444447" name="Mondai" pageId="22" pageNumber="523" stateProvince="Santa Catarina">Mondaí</location>
, 
<geoCoordinate id="BD805060672BA42A212EFDBF54D2BD46" box="[553,683,583,608]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-27.10389">27°06'14&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A21BCFDBF553DBD46" box="[699,836,583,608]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-53.444447">53°26'40&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A2053FDB0527ABD46" box="[852,1027,584,608]" pageId="22" pageNumber="523" value="1998-08-08">
<date id="AC0A1067672BA42A2053FDB0527ABD46" box="[852,1027,584,608]" pageId="22" pageNumber="523" value="1998-08-08">8 August 1998</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A2714FDB052DEBD79" box="[1043,1191,584,607]" pageId="22" pageNumber="523">P.P. Marques</collectorName>
(
<specimenCode id="88129EDC672BA42A27BEFDB05325BD46" box="[1209,1372,584,608]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="22" pageNumber="523">MCTP 10313</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA672BA42A2673FDBF57A5BD8E" ID-GBIF-Occurrence="1820860228" collectingDate="2012-01-31" collectionCode="MCTP" collectorName="M. Machado" country="Brazil" latitude="-29.046389" location="Reserva Biologica de Sao Donato" longLatPrecision="20" longitude="-56.08361" municipality="Sao Borja" pageId="22" pageNumber="523" specimenCode="MCTP 34653" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul">
<emphasis id="EAC0EAB5672BA42A2673FDBF573FBDA3" bold="true" pageId="22" pageNumber="523">
<collectingRegion id="1A70F845672BA42A2673FDBF5747BDA3" country="Brazil" name="Rio Grande do Sul" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A2673FDBF5747BDA3" bold="true" italics="true" pageId="22" pageNumber="523">
Rio 
<collectingRegion id="1A70F845672BA42A2390FD945747BDA3" box="[151,318,620,645]" inferredBySuffix="true" pageId="22" pageNumber="523">Grande do Sul</collectingRegion>
</emphasis>
</collectingRegion>
:
</emphasis>
<specimenCount id="CEB2FD2E672BA42A224AFD9457C8BDA3" box="[333,433,620,645]" pageId="22" pageNumber="523" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="386FACDD672BA42A22BCFD945454BDA3" box="[443,557,620,645]" pageId="22" pageNumber="523">São Borja</collectingMunicipality>
, 
<location id="DD6B607C672BA42A2130FD9555C8BDA2" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A2130FD9555C8BDA2" box="[567,945,620,645]" country="Brazil" latitude="-29.046389" longLatPrecision="20" longitude="-56.08361" municipality="Sao Borja" name="Reserva Biologica de Sao Donato" pageId="22" pageNumber="523" stateProvince="Rio Grande do Sul">Reserva Biológica de São Donato</location>
, 
<geoCoordinate id="BD805060672BA42A20BBFD945247BDA2" box="[956,1086,620,644]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-29.046389">29°02'47&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A274EFD9452AABDA3" box="[1097,1235,620,645]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-56.08361">56°05'01&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A27DAFD9453E1BDA2" box="[1245,1432,620,645]" pageId="22" pageNumber="523" value="2012-01-31">
<date id="AC0A1067672BA42A27DAFD9453E1BDA2" box="[1245,1432,620,645]" pageId="22" pageNumber="523" value="2012-01-31">31 January 2012</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A2390FD68575FBD8E" box="[151,294,655,680]" pageId="22" pageNumber="523">M. Machado</collectorName>
(
<specimenCode id="88129EDC672BA42A2233FD6857ADBD8E" box="[308,468,655,680]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="22" pageNumber="523">MCTP 34653</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="68DC3CFA672BA42A22E0FD6853F6BD8E" ID-GBIF-Occurrence="1820860221" box="[487,1423,655,680]" collectingDate="1994-04-24" collectionCode="MCTP" collectorName="Projeto Garabi" country="Brazil" latitude="-28.044167" location="Pirapo" longLatPrecision="20" longitude="-55.200558" pageId="22" pageNumber="523" specimenCode="MCTP 55" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
<specimenCount id="CEB2FD2E672BA42A22E0FD68544FBD8E" box="[487,566,655,680]" pageId="22" pageNumber="523" type="male">1 male</specimenCount>
, 
<location id="DD6B607C672BA42A2146FD7754F4BD8E" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672BA42A2146FD7754F4BD8E" box="[577,653,655,680]" country="Brazil" latitude="-28.044167" longLatPrecision="20" longitude="-55.200558" name="Pirapo" pageId="22" pageNumber="523" stateProvince="Rio Grande do Sul">Pirapó</location>
, 
<geoCoordinate id="BD805060672BA42A219FFD685563BD8E" box="[664,794,656,680]" direction="south" orientation="latitude" pageId="22" pageNumber="523" precision="15" value="-28.044167">28°02'39&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672BA42A2022FD7755C9BD8E" box="[805,944,655,680]" direction="west" orientation="longitude" pageId="22" pageNumber="523" precision="15" value="-55.200558">55°12'02&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672BA42A20BCFD685222BD8E" box="[955,1115,655,680]" pageId="22" pageNumber="523" value="1994-04-24">
<date id="AC0A1067672BA42A20BCFD685222BD8E" box="[955,1115,655,680]" pageId="22" pageNumber="523" value="1994-04-24">24 April 1994</date>
</collectingDate>
, 
<collectorName id="75415371672BA42A2761FD775372BD8E" box="[1126,1291,655,680]" pageId="22" pageNumber="523">Projeto Garabi</collectorName>
(
<specimenCode id="88129EDC672BA42A2615FD7753FEBD8E" box="[1298,1415,655,680]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="22" pageNumber="523">MCTP 55</specimenCode>
)
</materialsCitation>
.
</paragraph>
<caption id="8CCB662F672BA42A2390F8AC5325B8AE" httpUri="https://zenodo.org/record/1183154/files/figure.png" pageId="22" pageNumber="523" startId="22.[151,250,1876,1898]" targetBox="[151,1436,733,1854]" targetPageId="22">
<paragraph id="D80B36A7672BA42A2390F8AC5325B8AE" blockId="22.[151,1436,1876,1928]" pageId="22" pageNumber="523">
<emphasis id="EAC0EAB5672BA42A2390F8AC5758B84C" bold="true" box="[151,289,1876,1898]" pageId="22" pageNumber="523">FIGURE 12.</emphasis>
<taxonomicName id="1FB44D24672BA42A222DF8AC557FB84C" authority="(TaczanoWski, 1872)" authorityName="TaczanoWski" authorityYear="1872" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[298,774,1876,1898]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="523" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB5672BA42A222DF8AC545FB84C" box="[298,550,1876,1898]" italics="true" pageId="22" pageNumber="523">Epicadus trituberculatus</emphasis>
(TaczanoWski, 1872)
</taxonomicName>
, female (MPEG 022662). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal). Scale bars: 2 mm (A), 1 mm (B), 0.2 mm (C, D).
</paragraph>
</caption>
<caption id="8CCB662F672AA42B2390F9C95425B9A2" httpUri="https://zenodo.org/record/1183156/files/figure.png" pageId="23" pageNumber="524" startId="23.[151,250,1585,1607]" targetBox="[151,1436,193,1564]" targetPageId="23">
<paragraph id="D80B36A7672AA42B2390F9C95425B9A2" blockId="23.[151,1436,1585,1668]" pageId="23" pageNumber="524">
<emphasis id="EAC0EAB5672AA42B2390F9C95758B961" bold="true" box="[151,289,1585,1607]" pageId="23" pageNumber="524">FIGURE 13.</emphasis>
<taxonomicName id="1FB44D24672AA42B222FF9C9557BB961" authority="(TaczanoWski, 1872)" authorityName="TaczanoWski" authorityYear="1872" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[296,770,1585,1607]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB5672AA42B222FF9C9545AB961" box="[296,547,1585,1607]" italics="true" pageId="23" pageNumber="524">Epicadus trituberculatus</emphasis>
(TaczanoWski, 1872)
</taxonomicName>
, male (MPEG 022623). A, B habitus (A dorsal, B frontal); C, D left palp (C frontal, D retrolateral). A dorsal habitus; B front; C frontal vieW of left palp; D retrolateral vieW of left palp. Scale bars: 1 mm (A), 0.5 mm (B), 0.1 mm (C, D).
</paragraph>
</caption>
<paragraph id="D80B36A7672AA42B2390F92D5353B858" blockId="23.[151,1436,1749,1918]" pageId="23" pageNumber="524">
<materialsCitation id="68DC3CFA672AA42B2390F92D52A2B9C8" ID-GBIF-Occurrence="1820860225" box="[151,1243,1749,1774]" collectingDate="1928-02" collectionCode="PERU, AMNH" collectorName="H. Bassler" country="Peru" latitude="-10.963056" location="El" longLatPrecision="21" longitude="-73.954445" municipality="rio Tambo" pageId="23" pageNumber="524" specimenCount="1" specimenCount-female="1">
<emphasis id="EAC0EAB5672AA42B2390F92D5695B9C8" bold="true" box="[151,236,1749,1774]" pageId="23" pageNumber="524">
<collectionCode id="BEA5AE62672AA42B2390F92D569EB9C8" box="[151,231,1749,1774]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14398" name="Universit<69> di Perugia" pageId="23" pageNumber="524">
<collectingCountry id="A0A37637672AA42B2390F92D569EB9C8" box="[151,231,1749,1774]" name="Peru" pageId="23" pageNumber="524">PERU</collectingCountry>
</collectionCode>
:
</emphasis>
<specimenCount id="CEB2FD2E672AA42B23F2F92E5725B9C8" box="[245,348,1749,1774]" pageId="23" pageNumber="524" type="female">1 female</specimenCount>
, 
<location id="DD6B607C672AA42B226EF92D57FBB9C8" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672AA42B226EF92D57FBB9C8" box="[361,386,1749,1774]" country="Peru" latitude="-10.963056" longLatPrecision="21" longitude="-73.954445" municipality="rio Tambo" name="El" pageId="23" pageNumber="524">El</location>
<collectingMunicipality id="386FACDD672AA42B228CF92E547DB9C8" box="[395,516,1749,1774]" pageId="23" pageNumber="524">río Tambo</collectingMunicipality>
, 
<geoCoordinate id="BD805060672AA42B2116F92D54EAB9C8" box="[529,659,1749,1774]" direction="south" orientation="latitude" pageId="23" pageNumber="524" precision="15" value="-10.963056">10°57'47&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672AA42B21A6F92D5552B9C8" box="[673,811,1749,1774]" direction="west" orientation="longitude" pageId="23" pageNumber="524" precision="15" value="-73.954445">73°57'16&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672AA42B203FF92D559BB9C8" box="[824,994,1749,1774]" pageId="23" pageNumber="524" value="1928-02">
<date id="AC0A1067672AA42B203FF92D559BB9C8" box="[824,994,1749,1774]" pageId="23" pageNumber="524" value="1928-02">February 1928</date>
</collectingDate>
, 
<collectorName id="75415371672AA42B20E8F92E521FB9C8" box="[1007,1126,1749,1774]" pageId="23" pageNumber="524">H. Bassler</collectorName>
(
<collectionCode id="BEA5AE62672AA42B2770F92E52AAB9C8" box="[1143,1235,1750,1774]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="524">AMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="68DC3CFA672AA42B27E0F92D5690B810" ID-GBIF-Occurrence="1820860224" collectingDate="1989-11" collectionCode="MACN" collectorName="M. Ramirez" country="Argentina" latitude="-25.597223" location="Missiones" longLatPrecision="21" longitude="-54.578056" municipality="Parque Nacional Iguazu" pageId="23" pageNumber="524" specimenCode="MACN 18683" specimenCount="1" specimenCount-female="1">
<emphasis id="EAC0EAB5672AA42B27E0F92D576AB834" bold="true" pageId="23" pageNumber="524">
<collectingCountry id="A0A37637672AA42B27E0F92D53EEB9C8" box="[1255,1431,1749,1774]" name="Argentina" pageId="23" pageNumber="524">ARGENTINA</collectingCountry>
: 
<location id="DD6B607C672AA42B2390F9015773B834" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672AA42B2390F9015773B834" box="[151,266,1785,1810]" country="Argentina" latitude="-25.597223" longLatPrecision="21" longitude="-54.578056" municipality="Parque Nacional Iguazu" name="Missiones" pageId="23" pageNumber="524">
<emphasis id="EAC0EAB5672AA42B2390F9015773B834" bold="true" box="[151,266,1785,1810]" italics="true" pageId="23" pageNumber="524">Missiones</emphasis>
</location>
:
</emphasis>
<specimenCount id="CEB2FD2E672AA42B2219F90257FCB834" box="[286,389,1785,1810]" pageId="23" pageNumber="524" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="386FACDD672AA42B2293F90254D3B834" box="[404,682,1785,1810]" pageId="23" pageNumber="524">Parque Nacional Iguazú</collectingMunicipality>
, 
<geoCoordinate id="BD805060672AA42B21BEF9015540B834" box="[697,825,1785,1810]" direction="south" orientation="latitude" pageId="23" pageNumber="524" precision="15" value="-25.597223">25°35'50''S</geoCoordinate>
, 
<geoCoordinate id="BD805060672AA42B204FF90155AAB834" box="[840,979,1785,1810]" direction="west" orientation="longitude" pageId="23" pageNumber="524" precision="15" value="-54.578056">54°34'41&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672AA42B20E6F90152E4B834" box="[993,1181,1785,1810]" pageId="23" pageNumber="524" value="1989-11">
<date id="AC0A1067672AA42B20E6F90152E4B834" box="[993,1181,1785,1810]" pageId="23" pageNumber="524" value="1989-11">November 1989</date>
</collectingDate>
, 
<collectorName id="75415371672AA42B27ACF902534CB834" box="[1195,1333,1786,1810]" pageId="23" pageNumber="524">M. Ramírez</collectorName>
(
<specimenCode id="88129EDC672AA42B2641F9025698B810" collectionCode="MACN" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="524">MACN 18683</specimenCode>
)
</materialsCitation>
; 1 male, (MACN 18953); 1 female, 
<date id="AC0A1067672AA42B21B5F8E555C6B810" box="[690,959,1821,1846]" pageId="23" pageNumber="524" value="1999-12-11" valueMax="1999-12-16" valueMin="1999-12-11">11–16 December 1999</date>
(MACN 18687); 
<materialsCitation id="68DC3CFA672AA42B2799F8E65599B87C" ID-GBIF-Occurrence="1820860253" collectingDate="1989-06-21" collectingDateMax="1989-06-30" collectingDateMin="1989-06-21" collectionCode="MCTP" collectorName="Projeto Garabi" country="Argentina" latitude="-27.871944" location="San Javier" longLatPrecision="20" longitude="-55.13889" pageId="23" pageNumber="524" specimenCode="MCTP 567" specimenCount="1" specimenCount-female="1">
<specimenCount id="CEB2FD2E672AA42B2799F8E65370B810" box="[1182,1289,1821,1846]" pageId="23" pageNumber="524" type="female">1 female</specimenCount>
, 
<location id="DD6B607C672AA42B261CF8E653E1B810" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672AA42B261CF8E653E1B810" box="[1307,1432,1821,1846]" country="Argentina" latitude="-27.871944" longLatPrecision="20" longitude="-55.13889" name="San Javier" pageId="23" pageNumber="524">San Javier</location>
, 
<geoCoordinate id="BD805060672AA42B2390F8B95761B87C" box="[151,280,1857,1882]" direction="south" orientation="latitude" pageId="23" pageNumber="524" precision="15" value="-27.871944">27°52'19&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672AA42B2220F8B957C8B87C" box="[295,433,1857,1882]" direction="west" orientation="longitude" pageId="23" pageNumber="524" precision="15" value="-55.13889">55°08'20&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672AA42B22C7F8BA54F0B87C" box="[448,649,1858,1882]" pageId="23" pageNumber="524" value="1989-06-21" valueMax="1989-06-30" valueMin="1989-06-21">
<date id="AC0A1067672AA42B22C7F8BA54F0B87C" box="[448,649,1858,1882]" pageId="23" pageNumber="524" value="1989-06-21" valueMax="1989-06-30" valueMin="1989-06-21">21–30 June 1989</date>
</collectingDate>
, 
<collectorName id="75415371672AA42B219FF8B95546B87C" box="[664,831,1857,1882]" pageId="23" pageNumber="524">Projeto Garabi</collectorName>
(
<specimenCode id="88129EDC672AA42B2056F8BA55AEB87C" box="[849,983,1857,1882]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="23" pageNumber="524">MCTP 567</specimenCode>
)
</materialsCitation>
; 1 male, 
<date id="AC0A1067672AA42B2748F8BA5345B87C" box="[1103,1340,1857,1882]" pageId="23" pageNumber="524" value="1988-10-22" valueMax="1988-10-30" valueMin="1988-10-22">22–30 October 1988</date>
(MCTP 1661); 
<materialsCitation id="68DC3CFA672AA42B23E0F89E535FB858" ID-GBIF-Occurrence="1820860268" box="[231,1318,1893,1918]" collectingDate="1954-11" collectionCode="MACN" collectorName="R. D. Schiapelli" country="Argentina" latitude="-27.9" location="Santa Maria" longLatPrecision="20" longitude="-55.383335" pageId="23" pageNumber="524" specimenCode="MACN 18959" specimenCount="1" specimenCount-male="1">
<specimenCount id="CEB2FD2E672AA42B23E0F89E574CB858" box="[231,309,1893,1918]" pageId="23" pageNumber="524" type="male">1 male</specimenCount>
, 
<location id="DD6B607C672AA42B2247F89E57B5B858" LSID="urn:lsid:plazi:treatment:501D87B16728A4202390FD77540EBD3E:DD6B607C672AA42B2247F89E57B5B858" box="[320,460,1893,1918]" country="Argentina" latitude="-27.9" longLatPrecision="20" longitude="-55.383335" name="Santa Maria" pageId="23" pageNumber="524">Santa Maria</location>
, 
<geoCoordinate id="BD805060672AA42B22D0F89D5421B858" box="[471,600,1893,1918]" direction="south" orientation="latitude" pageId="23" pageNumber="524" precision="15" value="-27.9">27°54'00&quot;S</geoCoordinate>
, 
<geoCoordinate id="BD805060672AA42B2163F89D5494B858" box="[612,749,1893,1918]" direction="west" orientation="longitude" pageId="23" pageNumber="524" precision="15" value="-55.383335">55°23'00&quot;W</geoCoordinate>
, 
<collectingDate id="BC4EE98F672AA42B21FEF89D55C8B858" box="[761,945,1893,1918]" pageId="23" pageNumber="524" value="1954-11">
<date id="AC0A1067672AA42B21FEF89D55C8B858" box="[761,945,1893,1918]" pageId="23" pageNumber="524" value="1954-11">November 1954</date>
</collectingDate>
, 
<collectorName id="75415371672AA42B20BBF89E5211B858" box="[956,1128,1893,1918]" pageId="23" pageNumber="524">R.D. Schiapelli</collectorName>
(
<specimenCode id="88129EDC672AA42B2770F89E5367B858" box="[1143,1310,1893,1918]" collectionCode="MACN" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="524">MACN 18959</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="8CCB662F6725A4242390F86954E6B8C2" httpUri="https://zenodo.org/record/1183158/files/figure.png" pageId="24" pageNumber="525" startId="24.[151,250,1937,1959]" targetBox="[181,1404,193,1921]" targetPageId="24">
<paragraph id="D80B36A76725A4242390F86954E6B8C2" blockId="24.[151,1436,1937,2020]" pageId="24" pageNumber="525">
<emphasis id="EAC0EAB56725A4242390F8695758B881" bold="true" box="[151,289,1937,1959]" pageId="24" pageNumber="525">FIGURE 14.</emphasis>
Color variations of females. A–D 
<taxonomicName id="1FB44D246725A424218CF8695245B881" authority="(Mello-Leitao, 1929)" authorityName="Mello-Leitao" authorityYear="1929" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[651,1084,1937,1959]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="EAC0EAB56725A424218CF8695490B881" box="[651,745,1937,1959]" italics="true" pageId="24" pageNumber="525">Epicadus</emphasis>
<emphasis id="EAC0EAB56725A42421F5F8695524B881" box="[754,861,1937,1959]" italics="true" pageId="24" pageNumber="525">pustulosus</emphasis>
(Mello-Leitão, 1929)
</taxonomicName>
(A, B MZSP 45484; C, D MZSP 51068); Scale bars: 2 mm (A, C), 1 mm (B, D). Color variations of females of E–F 
<taxonomicName id="1FB44D246725A4242705F85753EAB8E3" authority="(RoeWer, 1951)" authorityName="RoeWer" authorityYear="1951" baseAuthorityName="Roewer" baseAuthorityYear="1951" box="[1026,1427,1967,1989]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="taczanowskii">
<emphasis id="EAC0EAB56725A4242705F8575219B8E3" box="[1026,1120,1967,1989]" italics="true" pageId="24" pageNumber="525">Epicadus</emphasis>
<emphasis id="EAC0EAB56725A424276EF8575295B8E3" box="[1129,1260,1967,1989]" italics="true" pageId="24" pageNumber="525">taczanowskii</emphasis>
(RoeWer, 1951)
</taxonomicName>
: Photo credits: Kenji Nishida (E); Arthur Anker (F).
</paragraph>
</caption>
<caption id="8CCB662F6724A4252390F8C6570DB896" httpUri="https://zenodo.org/record/1183160/files/figure.png" pageId="25" pageNumber="526" startId="25.[151,250,1854,1876]" targetBox="[151,1436,193,1833]" targetPageId="25">
<paragraph id="D80B36A76724A4252390F8C6570DB896" blockId="25.[151,1436,1854,1968]" pageId="25" pageNumber="526">
<emphasis id="EAC0EAB56724A4252390F8C65758B872" bold="true" box="[151,289,1854,1876]" pageId="25" pageNumber="526">FIGURE 15.</emphasis>
Female specimens 
<emphasis id="EAC0EAB56724A42522EAF8C7544AB872" box="[493,563,1855,1876]" italics="true" pageId="25" pageNumber="526">in vivo</emphasis>
. A–C 
<taxonomicName id="1FB44D246724A425217EF8C65263B872" authority="(Mello-Leitao, 1929)" authorityName="Mello-Leitao" authorityYear="1929" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[633,1050,1854,1876]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="caudatus">
<emphasis id="EAC0EAB56724A425217EF8C654AEB872" box="[633,727,1854,1876]" italics="true" pageId="25" pageNumber="526">Epicadus</emphasis>
<emphasis id="EAC0EAB56724A42521E7F8C65545B872" box="[736,828,1854,1876]" italics="true" pageId="25" pageNumber="526">caudatus</emphasis>
(Mello-Leitão, 1929)
</taxonomicName>
A nuptial guard, B egg-sac guarding behavior, C diagnostic orange blister-shaped setae sockets); D 
<taxonomicName id="1FB44D246724A4252036F8A65230B855" authority="Banks 1909" authorityName="Banks" authorityYear="1909" box="[817,1097,1885,1907]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="EAC0EAB56724A4252036F8A65531B855" box="[817,840,1886,1907]" italics="true" pageId="25" pageNumber="526">E.</emphasis>
<emphasis id="EAC0EAB56724A4252055F8A555C6B855" box="[850,959,1885,1907]" italics="true" pageId="25" pageNumber="526">granulatus</emphasis>
Banks 1909
</taxonomicName>
; E 
<taxonomicName id="1FB44D246724A4252776F8A5533DB855" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1929" box="[1137,1348,1885,1907]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="EAC0EAB56724A4252776F8A5533DB855" box="[1137,1348,1885,1907]" italics="true" pageId="25" pageNumber="526">Epicadus pustulosus</emphasis>
</taxonomicName>
(Mello- Leitão, 1929), diagnostic yelloW blister-shaped setae sockets. Photo credits: Alfredo Colón Archilla (A, B, C); Morley Read (D); Paul Bertner (E).
</paragraph>
</caption>
<caption id="8CCB662F6727A4262390F8B557F6B8F8" httpUri="https://zenodo.org/record/1183162/files/figure.png" pageId="26" pageNumber="527" startId="26.[151,250,1869,1891]" targetBox="[184,1408,193,1859]" targetPageId="26">
<paragraph id="D80B36A76727A4262390F8B557F6B8F8" blockId="26.[151,1436,1869,2014]" pageId="26" pageNumber="527">
<emphasis id="EAC0EAB56727A4262390F8B5575DB845" bold="true" box="[151,292,1869,1891]" pageId="26" pageNumber="527">FIGURE 16.</emphasis>
Female specimens 
<emphasis id="EAC0EAB56727A42622FDF8B7543AB842" box="[506,579,1871,1892]" italics="true" pageId="26" pageNumber="527">in vivo</emphasis>
. A 
<taxonomicName id="1FB44D246727A4262175F8B55276B845" authority="(RoeWer, 1951)" authorityName="RoeWer" authorityYear="1951" baseAuthorityName="Roewer" baseAuthorityYear="1951" box="[626,1039,1869,1891]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="taczanowskii">
<emphasis id="EAC0EAB56727A4262175F8B554A9B845" box="[626,720,1869,1891]" italics="true" pageId="26" pageNumber="527">Epicadus</emphasis>
<emphasis id="EAC0EAB56727A42621DCF8B55527B845" box="[731,862,1869,1891]" italics="true" pageId="26" pageNumber="527">taczanowskii</emphasis>
(RoeWer, 1951)
</taxonomicName>
; B 
<taxonomicName id="1FB44D246727A426273CF8B55361B845" authority="Machado &amp; Teixeira &amp; Lise, 2018" authorityName="Machado &amp; Teixeira &amp; Lise" authorityYear="2018" box="[1083,1304,1869,1891]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="dimidiaster" status="sp. nov.">
<emphasis id="EAC0EAB56727A426273CF8B55361B845" box="[1083,1304,1869,1891]" italics="true" pageId="26" pageNumber="527">Epicadus dimidiaster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1F357CE6727A4262624F8B55301B845" box="[1315,1400,1869,1891]" pageId="26" pageNumber="527" rank="species">
<emphasis id="EAC0EAB56727A4262624F8B55301B845" bold="true" box="[1315,1400,1869,1891]" pageId="26" pageNumber="527">sp. nov.</emphasis>
</taxonomicNameLabel>
; C 
<taxonomicName id="1FB44D246727A4262390F894545FB8A4" authority="Mello-Leitao, 1924" authorityName="Mello-Leitao" authorityYear="1924" box="[151,550,1900,1922]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="EAC0EAB56727A4262390F894572DB8A4" box="[151,340,1900,1922]" italics="true" pageId="26" pageNumber="527">Epicadus rubripes</emphasis>
Mello-Leitão, 1924
</taxonomicName>
(adapted from Silva-Moreira &amp; Machado 2016: fig. 1F); D 
<taxonomicName id="1FB44D246727A42627A4F89457D4B887" authority="(O. Pickard-Cambridge, 1869)" authorityName="O. Pickard-Cambridge" authorityYear="1869" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56727A42627A4F8945378B8A4" box="[1187,1281,1900,1922]" italics="true" pageId="26" pageNumber="527">Epicadus</emphasis>
<emphasis id="EAC0EAB56727A426260CF894530BB8A4" box="[1291,1394,1900,1922]" italics="true" pageId="26" pageNumber="527">camelinus</emphasis>
(O. Pickard-Cambridge, 1869)
</taxonomicName>
; E 
<taxonomicName id="1FB44D246727A42622D1F87355AEB887" authority="(Guerin-Meneville, 1829)" authorityName="Guerin-Meneville" authorityYear="1829" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1829" box="[470,983,1931,1953]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="heterogaster">
<emphasis id="EAC0EAB56727A42622D1F87354C7B887" box="[470,702,1931,1953]" italics="true" pageId="26" pageNumber="527">Epicadus heterogaster</emphasis>
(Guérin-Méneville, 1829)
</taxonomicName>
; F 
<taxonomicName id="1FB44D246727A42620F8F87356AFB8E6" authority="(TaczanoWski, 1872)" authorityName="TaczanoWski" authorityYear="1872" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56727A42620F8F8735286B887" box="[1023,1279,1931,1953]" italics="true" pageId="26" pageNumber="527">Epicadus trituberculatus</emphasis>
(TaczanoWski, 1872)
</taxonomicName>
. Photo credits: César Favacho (A); Arthur Anker (B); Almir Cândido de Almeida (C); Karl Kroeker (D); Edson Roberto (E); Edelcio Muscat (F).
</paragraph>
</caption>
<caption id="8CCB662F6726A4272390F91E57A1B839" httpUri="https://zenodo.org/record/1183164/files/figure.png" pageId="27" pageNumber="528" startId="27.[151,250,1766,1788]" targetBox="[151,1435,193,1736]" targetPageId="27">
<paragraph id="D80B36A76726A4272390F91E57A1B839" blockId="27.[151,1435,1766,1823]" pageId="27" pageNumber="528">
<emphasis id="EAC0EAB56726A4272390F91E5759B9DA" bold="true" box="[151,288,1766,1788]" pageId="27" pageNumber="528">FIGURE 17.</emphasis>
Data of distribution records of 
<taxonomicName id="1FB44D246726A4272166F91E54C6B9DA" authorityName="Simon, 1895:" authorityYear="1052" box="[609,703,1766,1788]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="EAC0EAB56726A4272166F91E54C6B9DA" box="[609,703,1766,1788]" italics="true" pageId="27" pageNumber="528">Epicadus</emphasis>
</taxonomicName>
species compiled from Machado 
<emphasis id="EAC0EAB56726A427271FF91F5232B9DA" box="[1048,1099,1766,1788]" italics="true" pageId="27" pageNumber="528">et al.</emphasis>
(2015, 2017); Silva-Moreira and Machado (2016); present study.
</paragraph>
</caption>
<subSubSection id="90AE652C6726A42023C0F8B7524ABE66" lastPageId="28" lastPageNumber="529" pageId="27" pageNumber="528" type="diagnosis">
<paragraph id="D80B36A76726A42023C0F8B7524ABE66" blockId="27.[151,1436,1871,2004]" lastBlockId="28.[151,1436,151,536]" lastPageId="28" lastPageNumber="529" pageId="27" pageNumber="528">
<emphasis id="EAC0EAB56726A42723C0F8B75738B84E" bold="true" box="[199,321,1871,1896]" pageId="27" pageNumber="528">Diagnosis.</emphasis>
Females of 
<taxonomicName id="1FB44D246726A42722D9F8A854D1B84E" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[478,680,1871,1896]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56726A42722D9F8A854D1B84E" box="[478,680,1871,1896]" italics="true" pageId="27" pageNumber="528">E. trituberculatus</emphasis>
</taxonomicName>
have a distinct morphology, being similar only to those of 
<taxonomicName id="1FB44D246726A4272683F8A85770B8AA" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56726A4272683F8A85770B8AA" italics="true" pageId="27" pageNumber="528">E. camelinus</emphasis>
</taxonomicName>
in the truncated shape of the median posterior opisthosomal projection (
<figureCitation id="408F2A226726A4272728F88B52EEB8AA" box="[1071,1175,1907,1932]" captionStart="FIGURE 12" captionStartId="22.[151,250,1876,1898]" captionTargetBox="[151,1436,733,1854]" captionTargetId="figure@22.[151,1436,733,1854]" captionTargetPageId="22" captionText="FIGURE 12. Epicadus trituberculatus (TaczanoWski, 1872), female (MPEG 022662). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A), 1 mm (B), 0.2mm (C, D)." httpUri="https://zenodo.org/record/1183154/files/figure.png" pageId="27" pageNumber="528">Fig. 12A</figureCitation>
). However, they can be distinguished by the simple coloration pattern, whereas 
<taxonomicName id="1FB44D246726A427202DF86055B9B896" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[810,960,1943,1968]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56726A427202DF86055B9B896" box="[810,960,1943,1968]" italics="true" pageId="27" pageNumber="528">E. camelinus</emphasis>
</taxonomicName>
females have large brown stains on the prosoma, anterior femora and tibiae. Unlike 
<taxonomicName id="1FB44D246726A4272189F8445559B8F2" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1869" box="[654,800,1979,2004]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="camelinus">
<emphasis id="EAC0EAB56726A4272189F8445559B8F2" box="[654,800,1979,2004]" italics="true" pageId="27" pageNumber="528">E. camelinus</emphasis>
</taxonomicName>
, the females of 
<taxonomicName id="1FB44D246726A42720D0F84452E5B8F2" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[983,1180,1979,2004]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56726A42720D0F84452E5B8F2" box="[983,1180,1979,2004]" italics="true" pageId="27" pageNumber="528">E. trituberculatus</emphasis>
</taxonomicName>
bear three equal-sized tips at the end of the median posterior opisthosomal projection, their copulatory openings are entirely covered by the posterior folds of the epigynal plate and their secondary spermathecae are enlarged (
<figureCitation id="408F2A226721A4202788FF445358BFF2" box="[1167,1313,188,213]" captionStart="FIGURE 12" captionStartId="22.[151,250,1876,1898]" captionTargetBox="[151,1436,733,1854]" captionTargetId="figure@22.[151,1436,733,1854]" captionTargetPageId="22" captionText="FIGURE 12. Epicadus trituberculatus (TaczanoWski, 1872), female (MPEG 022662). A, B habitus (A dorsal, B frontal); C epigynal plate (ventral); D spermathecae and copulatory ducts (dorsal).Scale bars: 2 mm (A), 1 mm (B), 0.2mm (C, D)." httpUri="https://zenodo.org/record/1183154/files/figure.png" pageId="28" pageNumber="529">Figs 12C, D</figureCitation>
). In dorsal view (
<figureCitation id="408F2A226721A42023D9FF27573FBFDE" box="[222,326,223,248]" captionStart="FIGURE 13" captionStartId="23.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1564]" captionTargetId="figure@23.[151,1436,193,1564]" captionTargetPageId="23" captionText="FIGURE 13. Epicadus trituberculatus (TaczanoWski, 1872), male (MPEG 022623). A, B habitus (A dorsal, B frontal); C, D left palp (C frontal, D retrolateral).A dorsal habitus; B front; C frontal vieW of left palp; D retrolateral vieW of left palp. Scale bars:1 mm (A), 0.5 mm (B), 0.1 mm (C, D)." httpUri="https://zenodo.org/record/1183156/files/figure.png" pageId="28" pageNumber="529">Fig. 13A</figureCitation>
), males of 
<taxonomicName id="1FB44D246721A42022C3FF1954F0BFDE" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[452,649,223,248]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56721A42022C3FF1954F0BFDE" box="[452,649,223,248]" italics="true" pageId="28" pageNumber="529">E. trituberculatus</emphasis>
</taxonomicName>
are almost indiscernible from those of 
<taxonomicName id="1FB44D246721A420274EFF195290BFDE" authority="Machado &amp; Teixeira &amp; Lise, 2018" authorityName="Machado &amp; Teixeira &amp; Lise" authorityYear="2018" box="[1097,1257,223,248]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="dimidiaster" status="sp. nov.">
<emphasis id="EAC0EAB56721A420274EFF195290BFDE" box="[1097,1257,223,248]" italics="true" pageId="28" pageNumber="529">E. dimidiaster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1F357CE6721A42027F6FF275332BFDE" box="[1265,1355,223,248]" pageId="28" pageNumber="529" rank="species">
<emphasis id="EAC0EAB56721A42027F6FF275332BFDE" bold="true" box="[1265,1355,223,248]" pageId="28" pageNumber="529">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="1FB44D246721A4202683FF195751BE3B" baseAuthorityName="Roewer" baseAuthorityYear="1951" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="taczanowskii">
<emphasis id="EAC0EAB56721A4202683FF195751BE3B" italics="true" pageId="28" pageNumber="529">E. taczanowskii</emphasis>
</taxonomicName>
, but they can be distinguished by the slightly higher clypeus (
<figureCitation id="408F2A226721A4202713FEFC5205BE3A" box="[1044,1148,260,285]" captionStart="FIGURE 13" captionStartId="23.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1564]" captionTargetId="figure@23.[151,1436,193,1564]" captionTargetPageId="23" captionText="FIGURE 13. Epicadus trituberculatus (TaczanoWski, 1872), male (MPEG 022623). A, B habitus (A dorsal, B frontal); C, D left palp (C frontal, D retrolateral).A dorsal habitus; B front; C frontal vieW of left palp; D retrolateral vieW of left palp. Scale bars:1 mm (A), 0.5 mm (B), 0.1 mm (C, D)." httpUri="https://zenodo.org/record/1183156/files/figure.png" pageId="28" pageNumber="529">Fig. 13B</figureCitation>
) and especially by their tegulum with TF and embolus emerging from eight o’clock position (
<figureCitation id="408F2A226721A42020A7FEDF5252BE66" box="[928,1067,295,320]" captionStart="FIGURE 13" captionStartId="23.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1564]" captionTargetId="figure@23.[151,1436,193,1564]" captionTargetPageId="23" captionText="FIGURE 13. Epicadus trituberculatus (TaczanoWski, 1872), male (MPEG 022623). A, B habitus (A dorsal, B frontal); C, D left palp (C frontal, D retrolateral).A dorsal habitus; B front; C frontal vieW of left palp; D retrolateral vieW of left palp. Scale bars:1 mm (A), 0.5 mm (B), 0.1 mm (C, D)." httpUri="https://zenodo.org/record/1183156/files/figure.png" pageId="28" pageNumber="529">Figs 13C, D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6721A42023C0FEB45323BE43" box="[199,1370,332,357]" pageId="28" pageNumber="529" type="description">
<paragraph id="D80B36A76721A42023C0FEB45323BE43" blockId="28.[151,1436,151,536]" box="[199,1370,332,357]" pageId="28" pageNumber="529">
<emphasis id="EAC0EAB56721A42023C0FEB45720BE43" bold="true" box="[199,345,332,357]" pageId="28" pageNumber="529">Description.</emphasis>
See description of 
<taxonomicName id="1FB44D246721A4202137FEB45492BE43" authorityName="Mello-Leitao" authorityYear="1932" box="[560,747,332,357]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="EAC0EAB56721A4202137FEB45492BE43" box="[560,747,332,357]" italics="true" pageId="28" pageNumber="529">Epicadus planus</emphasis>
</taxonomicName>
(junior synonym) in 
<bibRefCitation id="BC254B566721A42020DFFEB4532FBE43" box="[984,1366,332,357]" pageId="28" pageNumber="529" refId="ref19438" refString="Silva-Moreira, T. &amp; Machado, M. (2016) Taxonomic revision of the crab spider genus Epicadus Simon, 1895 (Arachnida: Araneae: Thomisidae) With notes on related genera of Stephanopinae Simon, 1895. Zootaxa, 4147 (3), 281 - 310. https: // doi. org / 10.11646 / zootaxa. 4147.3.4" type="journal article">Silva-Moreira &amp; Machado (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6721A42023C0FE975564BE8A" pageId="28" pageNumber="529" type="distribution">
<paragraph id="D80B36A76721A42023C0FE975564BE8A" blockId="28.[151,1436,151,536]" pageId="28" pageNumber="529">
<emphasis id="EAC0EAB56721A42023C0FE975719BEAE" bold="true" box="[199,352,367,392]" pageId="28" pageNumber="529">Distribution.</emphasis>
<collectingCountry id="A0A376376721A420226FFE885469BEAE" box="[360,528,368,392]" name="Argentina" pageId="28" pageNumber="529">ARGENTINA</collectingCountry>
: 
<collectingRegion id="1A70F8456721A420211DFE9754FDBEAE" box="[538,644,367,392]" country="Paraguay" name="Misiones" pageId="28" pageNumber="529">Misiones</collectingRegion>
; 
<collectingCountry id="A0A376376721A4202197FE885483BEAE" box="[656,762,368,392]" name="Brazil" pageId="28" pageNumber="529">BRAZIL</collectingCountry>
: 
<collectingRegion id="1A70F8456721A4202002FE9755A2BEAE" box="[773,987,367,392]" country="Brazil" name="Rio Grande do Sul" pageId="28" pageNumber="529">Rio Grande do Sul</collectingRegion>
, 
<collectingRegion id="1A70F8456721A42020E0FE8852F6BEAE" box="[999,1167,367,392]" country="Brazil" name="Santa Catarina" pageId="28" pageNumber="529">Santa Catarina</collectingRegion>
, 
<collectingRegion id="1A70F8456721A420279DFE885290BEAE" box="[1178,1257,368,392]" country="Brazil" name="Parana" pageId="28" pageNumber="529">Paraná</collectingRegion>
, 
<collectingRegion id="1A70F8456721A42027F2FE9753EEBEAE" box="[1269,1431,367,392]" country="Brazil" name="Espirito Santo" pageId="28" pageNumber="529">Espírito Santo</collectingRegion>
, 
<collectingRegion id="1A70F8456721A4202390FE6C56A2BE8B" box="[151,219,404,429]" country="Brazil" name="Goias" pageId="28" pageNumber="529">Goiás</collectingRegion>
, 
<collectingRegion id="1A70F8456721A42023E1FE6D5703BE8A" box="[230,378,404,428]" country="Brazil" name="Mato Grosso" pageId="28" pageNumber="529">Mato Grosso</collectingRegion>
, 
<collectingRegion id="1A70F8456721A4202282FE6C5787BE8A" box="[389,510,404,428]" country="Peru" name="Amazonas" pageId="28" pageNumber="529">Amazonas</collectingRegion>
, 
<collectingRegion id="1A70F8456721A420210DFE6C5446BE8A" box="[522,575,404,428]" country="Brazil" name="Para" pageId="28" pageNumber="529">Pará</collectingRegion>
, 
<collectingRegion id="1A70F8456721A420214EFE6C54E3BE8A" box="[585,666,404,428]" country="Brazil" name="Amapa" pageId="28" pageNumber="529">Amapá</collectingRegion>
(
<figureCitation id="408F2A226721A42021ADFE6C5569BE8A" box="[682,784,404,429]" captionStart="FIGURE 17" captionStartId="27.[151,250,1766,1788]" captionTargetBox="[151,1435,193,1736]" captionTargetId="figure@27.[151,1436,193,1744]" captionTargetPageId="27" captionText="FIGURE 17. Data of distribution records of Epicadus species compiled from Machado et al. (2015, 2017); Silva-Moreira and Machado (2016); present study." httpUri="https://zenodo.org/record/1183164/files/figure.png" pageId="28" pageNumber="529">Fig. 17B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90AE652C6721A42023C0FE4F540EBD3E" pageId="28" pageNumber="529" type="description">
<paragraph id="D80B36A76721A42023C0FE4F540EBD3E" blockId="28.[151,1436,151,536]" pageId="28" pageNumber="529">
<emphasis id="EAC0EAB56721A42023C0FE4F5745BEF6" bold="true" box="[199,316,439,464]" pageId="28" pageNumber="529">Variation.</emphasis>
Females of 
<taxonomicName id="1FB44D246721A42022CBFE4154EBBEF6" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[460,658,439,464]" class="Arachnida" family="Thomisidae" genus="Epicadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="trituberculatus">
<emphasis id="EAC0EAB56721A42022CBFE4154EBBEF6" box="[460,658,439,464]" italics="true" pageId="28" pageNumber="529">E. trituberculatus</emphasis>
</taxonomicName>
may present the yellowish coloration 
<emphasis id="EAC0EAB56721A4202741FE4152EABEF6" box="[1094,1171,441,464]" italics="true" pageId="28" pageNumber="529">in vivo</emphasis>
, but specimens may as well be totally whitish or show a whitish opisthosoma with prosoma and legs in magenta (
<figureCitation id="408F2A226721A42027B0FE245364BED3" box="[1207,1309,476,501]" captionStart="FIGURE 16" captionStartId="26.[151,250,1869,1891]" captionTargetBox="[184,1408,193,1859]" captionTargetId="figure@26.[184,1408,193,1859]" captionTargetPageId="26" captionText="FIGURE 16. Female specimens in vivo. A Epicadus taczanowskii (RoeWer, 1951); B Epicadus dimidiaster sp. nov.; C Epicadus rubripes Mello-Leitão, 1924 (adapted from Silva-Moreira &amp; Machado 2016: fig. 1F); D Epicadus camelinus (O. Pickard-Cambridge, 1869); E Epicadus heterogaster (Guérin-Méneville, 1829); F Epicadus trituberculatus (TaczanoWski, 1872). Photo credits:César Favacho (A); Arthur Anker (B); Almir Cândido de Almeida (C); Karl Kroeker (D); Edson Roberto (E); Edelcio Muscat (F)." httpUri="https://zenodo.org/record/1183162/files/figure.png" pageId="28" pageNumber="529">Fig. 16F</figureCitation>
; see Silva- 
<bibRefCitation id="BC254B566721A4202390FE0757AABD3E" box="[151,467,511,536]" pageId="28" pageNumber="529" refId="ref19438" refString="Silva-Moreira, T. &amp; Machado, M. (2016) Taxonomic revision of the crab spider genus Epicadus Simon, 1895 (Arachnida: Araneae: Thomisidae) With notes on related genera of Stephanopinae Simon, 1895. Zootaxa, 4147 (3), 281 - 310. https: // doi. org / 10.11646 / zootaxa. 4147.3.4" type="journal article">Moreira and Machado 2016</bibRefCitation>
: figs 1G, H, I).
</paragraph>
</subSubSection>
</treatment>
</document>