231 lines
24 KiB
XML
231 lines
24 KiB
XML
<document id="7CD3BB2ED3755619CF4A6EB4E013D207" ID-DOI="10.5281/zenodo.175214" ID-GBIF-Dataset="98a01672-a763-414f-8e60-be30f03f54eb" ID-ISSN="11755326" ID-Zenodo-Dep="175214" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459802938358" checkinUser="plazi" docAuthor="Zahniser, James N. & Hicks, Andrew" docDate="2007" docId="480487D2081BFF91FF3F71201003D5DF" docLanguage="en" docName="zt01390p039.pdf" docOrigin="Zootaxa 1390" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Athysanella (Amphipyga) Osborn" docType="treatment" docVersion="10" lastPageNumber="32" masterDocId="B43DFFAA081EFF94FFA87302125FD360" masterDocTitle="Three new species of Icaia Linnavuori (Hemiptera: Cicadellidae: Deltocephalinae) from Peru and distinction of the genus from Athysanella Baker" masterLastPageNumber="39" masterPageNumber="27" pageNumber="32" updateTime="1698220144257" updateUser="plazi">
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<mods:titleInfo id="115D1F03F9EB8EA459DE852C6706273E">
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<mods:title id="6CBBC1F147AF205B203412DE5D57FDC7">Three new species of Icaia Linnavuori (Hemiptera: Cicadellidae: Deltocephalinae) from Peru and distinction of the genus from Athysanella Baker</mods:title>
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<mods:name id="FE5AC57068B85AECD24AC9F69664CDB5" type="personal">
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<mods:namePart id="39FA836008153010122D7F3F02CC47CC">Zahniser, James N.</mods:namePart>
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<mods:name id="EA746AD94D338BB87D783F7528A288A4" type="personal">
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<mods:roleTerm id="6B1199DB61B38916CE55EBF210764E45">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="900F9C17E49D84D4CECF6F85D6519537">Hicks, Andrew</mods:namePart>
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<mods:typeOfResource id="CD89D8661EF92ABBE7603927BC5F6D2E">text</mods:typeOfResource>
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<mods:title id="35B24A4CFABA5F7FAB87239D0128BF72">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CE589E8E7C172FC62F830D360A761D11">
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<mods:date id="079BCA396070B6EE6493B59FF0471FD5">2007</mods:date>
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<mods:detail id="9EEAF2653B73F338FAC96DAC4D0FE725" type="volume">
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<mods:number id="AEAC6E132E43B05EE0E8CBE3E62D015D">1390</mods:number>
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<mods:start id="662A7B1BCE21583E5A53322CAFFC6322">27</mods:start>
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<mods:end id="7F73DC0E37BD5CB1BE607F39A016BDF2">39</mods:end>
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<mods:classification id="3CB2C23706E299547F283D102BE73995">journal article</mods:classification>
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<mods:identifier id="C8C539169677CBCE6E4289D035F8F153" type="DOI">10.5281/zenodo.175214</mods:identifier>
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<treatment id="480487D2081BFF91FF3F71201003D5DF" ID-DOI="http://doi.org/10.5281/zenodo.5671223" ID-GBIF-Taxon="119352185" ID-Zenodo-Dep="5671223" LSID="urn:lsid:plazi:treatment:480487D2081BFF91FF3F71201003D5DF" httpUri="http://treatment.plazi.org/id/480487D2081BFF91FF3F71201003D5DF" lastPageNumber="32" pageId="5" pageNumber="32">
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<subSubSection id="88B7654F081BFF91FF3F712011B8D1E7" pageId="5" pageNumber="32" type="nomenclature">
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<paragraph id="C01236C4081BFF91FF3F7120106ED15C" blockId="5.[151,1019,546,647]" box="[151,561,546,572]" pageId="5" pageNumber="32">
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<heading id="9B5A81A8081BFF91FF3F7120106ED15C" bold="true" box="[151,561,546,572]" fontSize="11" level="1" pageId="5" pageNumber="32" reason="1">
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<taxonomicName id="07AD4D47081BFF91FF3F7120106ED15C" ID-CoL="8JMFX" authority="Osborn" authorityName="Osborn" box="[151,561,546,572]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="subGenus" subGenus="Amphipyga">
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<emphasis id="F2D9EAD6081BFF91FF3F7120106ED15C" bold="true" box="[151,561,546,572]" pageId="5" pageNumber="32">
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<emphasis id="F2D9EAD6081BFF91FF3F71201379D15C" bold="true" box="[151,294,546,572]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
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(
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<emphasis id="F2D9EAD6081BFF91FE9F7120139ED15C" bold="true" box="[311,449,546,572]" italics="true" pageId="5" pageNumber="32">Amphipyga</emphasis>
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) Osborn
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="C01236C4081BFF91FF6E714711B8D1E7" blockId="5.[151,1019,546,647]" pageId="5" pageNumber="32">
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<taxonomicName id="07AD4D47081BFF91FF6E714713A8D13F" authority="Ball & Beamer" authorityName="Ball & Beamer" box="[198,503,581,607]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="rubicunda">
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<emphasis id="F2D9EAD6081BFF91FF6E71471360D13E" box="[198,319,581,606]" italics="true" pageId="5" pageNumber="32">rubicunda</emphasis>
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Ball & Beamer
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</taxonomicName>
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<emphasis id="F2D9EAD6081BFF91FE577147115BD13F" bold="true" box="[511,772,581,607]" pageId="5" pageNumber="32">placement reinstated</emphasis>
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from
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<taxonomicName id="07AD4D47081BFF91FCE3714713BBD1E7" authority="Ball & Beamer" authorityName="Ball & Beamer" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="triodana" subGenus="Diphipyga">
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<emphasis id="F2D9EAD6081BFF91FCE371471102D13E" box="[843,861,581,606]" italics="true" pageId="5" pageNumber="32">A</emphasis>
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. (
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<emphasis id="F2D9EAD6081BFF91FCDD714711AFD13E" box="[885,1008,581,606]" italics="true" pageId="5" pageNumber="32">Diphipyga</emphasis>
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)
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<emphasis id="F2D9EAD6081BFF91FF6E716F1374D1E6" box="[198,299,621,646]" italics="true" pageId="5" pageNumber="32">triodana</emphasis>
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Ball & Beamer
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</taxonomicName>
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<emphasis id="F2D9EAD6081BFF91FE43716F10AFD1E7" bold="true" box="[491,752,621,647]" pageId="5" pageNumber="32">placement reinstated</emphasis>
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from
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<emphasis id="F2D9EAD6081BFF91FC9F716F1116D1E6" box="[823,841,621,646]" italics="true" pageId="5" pageNumber="32">A</emphasis>
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. (
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<emphasis id="F2D9EAD6081BFF91FCCA716F1182D1E6" box="[866,989,621,646]" italics="true" pageId="5" pageNumber="32">Diphipyga</emphasis>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="88B7654F081BFF91FF3F71BF1003D5DF" pageId="5" pageNumber="32" type="description">
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<paragraph id="C01236C4081BFF91FF3F71BF1615D6C7" blockId="5.[151,1438,701,1727]" pageId="5" pageNumber="32">
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Parsimony analysis of the characters in
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<tableCitation id="8D2F037F081BFF91FDC671BF1099D1B7" box="[622,710,701,727]" captionStart="TABLE 1. A" captionStartId="2.[151,239,471,495]" captionTargetBox="[151,1434,597,1239]" captionTargetPageId="2" captionText="TABLE 1. A list of characters potentially distinguishing Icaia from Athysanella, scored for multiple representatives of each genus. Characters state observations for which different states were difficult to distinguish have a “? ” beside them. Numbers (or?) before the character states indicate the codes used for parsimony analysis." httpUri="http://table.plazi.org/id/94D2664C081CFF96FF3F72D5160DD154" pageId="5" pageNumber="32" tableUuid="94D2664C081CFF96FF3F72D5160DD154">Table 1</tableCitation>
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yielded 21 most parsimonious (MP) trees, and the strict consensus tree (
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<figureCitation id="58962A41081BFF91FE8771E713D5D19F" box="[303,394,741,767]" captionStart="FIGURE 46" captionStartId="11.[151,255,1940,1964]" captionTargetBox="[489,1114,1378,1904]" captionTargetId="figure@11.[470,1116,1369,1916]" captionTargetPageId="11" captionText="FIGURE 46. The strict consensus of 21 most parsimonious trees. Clade labeled “ A ” is referred to in the text. Numbers below branches indicate Bremer support values / bootstrap values over 50 %." httpUri="https://zenodo.org/record/175220/files/figure.png" pageId="5" pageNumber="32">Fig. 46</figureCitation>
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) generally reflects the observations outlined above.
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<taxonomicName id="07AD4D47081BFF91FBA571E716C8D19E" box="[1037,1175,741,766]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FBA571E716C8D19E" box="[1037,1175,741,766]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="07AD4D47081BFF91FB7171E71749D19E" box="[1241,1302,741,766]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FB7171E71749D19E" box="[1241,1302,741,766]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
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</taxonomicName>
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were each recovered as monophyletic groups. Characters that supported the monophyly of
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<taxonomicName id="07AD4D47081BFF91FBC6700F16A7D046" box="[1134,1272,781,806]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FBC6700F16A7D046" box="[1134,1272,781,806]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
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</taxonomicName>
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are the wide basal separation of the anterior arms of the connective (Figs. 38,39)— a character that was difficult to score for
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<taxonomicName id="07AD4D47081BFF91FF69705F13B5D016" box="[193,490,861,887]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="mexicana" subGenus="Diphipyga">
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<emphasis id="F2D9EAD6081BFF91FF69705F128CD016" box="[193,211,861,886]" italics="true" pageId="5" pageNumber="32">A</emphasis>
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. (
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<emphasis id="F2D9EAD6081BFF91FF44705F1337D016" box="[236,360,861,886]" italics="true" pageId="5" pageNumber="32">Diphipyga</emphasis>
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)
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<emphasis id="F2D9EAD6081BFF91FED1705F13B5D016" box="[377,490,861,886]" italics="true" pageId="5" pageNumber="32">mexicana</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="07AD4D47081BFF91FD8B705F10EED016" box="[547,689,861,886]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="straminea">
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<emphasis id="F2D9EAD6081BFF91FD8B705F10EED016" box="[547,689,861,886]" italics="true" pageId="5" pageNumber="32">I. straminea</emphasis>
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</taxonomicName>
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, and the presence of an apodeme above the aedeagus, although this character is not consistent in either genus. This phylogeny suggests that
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<emphasis id="F2D9EAD6081BFF91FB897087166CD0FE" box="[1057,1075,901,926]" italics="true" pageId="5" pageNumber="32">A</emphasis>
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. (
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<emphasis id="F2D9EAD6081BFF91FBE470871698D0FE" box="[1100,1223,901,926]" italics="true" pageId="5" pageNumber="32">Diphipyga</emphasis>
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)
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<emphasis id="F2D9EAD6081BFF91FB72708716BAD0FE" box="[1242,1253,901,926]" italics="true" pageId="5" pageNumber="32">s</emphasis>
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.
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<emphasis id="F2D9EAD6081BFF91FB45708716A7D0FE" box="[1261,1272,901,926]" italics="true" pageId="5" pageNumber="32">s</emphasis>
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. is basal with respect to the rest of
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<taxonomicName id="07AD4D47081BFF91FE2770AF1045D0A6" box="[399,538,941,966]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FE2770AF1045D0A6" box="[399,538,941,966]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
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</taxonomicName>
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, apparently in agreement with our observations of the similarity in the male genitalia of
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<emphasis id="F2D9EAD6081BFF91FE8B70D7136AD08E" box="[291,309,981,1006]" italics="true" pageId="5" pageNumber="32">A</emphasis>
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. (
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<emphasis id="F2D9EAD6081BFF91FEE570D71396D08E" box="[333,457,981,1006]" italics="true" pageId="5" pageNumber="32">Diphipyga</emphasis>
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)
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<emphasis id="F2D9EAD6081BFF91FE7270D713BAD08E" box="[474,485,981,1006]" italics="true" pageId="5" pageNumber="32">s</emphasis>
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.
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<emphasis id="F2D9EAD6081BFF91FE4470D713A8D08E" box="[492,503,981,1006]" italics="true" pageId="5" pageNumber="32">s</emphasis>
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. and
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<taxonomicName id="07AD4D47081BFF91FD9070D7102BD08E" box="[568,628,981,1006]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FD9070D7102BD08E" box="[568,628,981,1006]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
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</taxonomicName>
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.
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<emphasis id="F2D9EAD6081BFF91FD2B70D710CAD08E" box="[643,661,981,1006]" italics="true" pageId="5" pageNumber="32">A</emphasis>
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. (
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<emphasis id="F2D9EAD6081BFF91FD0570D71176D08E" box="[685,809,981,1006]" italics="true" pageId="5" pageNumber="32">Diphipyga</emphasis>
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)
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<emphasis id="F2D9EAD6081BFF91FC9270D7113FD08E" box="[826,864,981,1006]" italics="true" pageId="5" pageNumber="32">s.s.</emphasis>
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was never resolved as a monophyletic group. The only character supporting the monophyly of
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<taxonomicName id="07AD4D47081BFF91FD4270FF1179D776" box="[746,806,1021,1046]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FD4270FF1179D776" box="[746,806,1021,1046]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
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</taxonomicName>
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is the presence of a pair of apical or subapical processes of the aedeagus— a character that is not universally shared among species of
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<taxonomicName id="07AD4D47081BFF91FBDE772716EDD75E" box="[1142,1202,1061,1086]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FBDE772716EDD75E" box="[1142,1202,1061,1086]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
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</taxonomicName>
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, but appears in 6 or 7 of the 9 described species. This character does not occur in
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<taxonomicName id="07AD4D47081BFF91FCC1774F11ABD706" box="[873,1012,1101,1126]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FCC1774F11ABD706" box="[873,1012,1101,1126]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
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</taxonomicName>
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and rarely occurs in other genera of Chiasmini (one exception is
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<taxonomicName id="07AD4D47081BFF91FE4C7777109DD7EF" authority="Lethierry" authorityName="Lethierry" box="[484,706,1141,1167]" class="Insecta" family="Cicadellidae" genus="Aconura" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FE4C77771016D7EE" box="[484,585,1141,1166]" italics="true" pageId="5" pageNumber="32">Aconura</emphasis>
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Lethierry
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</taxonomicName>
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).
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<taxonomicName id="07AD4D47081BFF91FD7C7777114FD7EE" box="[724,784,1141,1166]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FD7C7777114FD7EE" box="[724,784,1141,1166]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
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</taxonomicName>
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spp. lack the wide separation of the anterior arms of the connective which supported monophyly of
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<taxonomicName id="07AD4D47081BFF91FD33779F1179D7D6" box="[667,806,1181,1206]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FD33779F1179D7D6" box="[667,806,1181,1206]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
|
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</taxonomicName>
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(including
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<emphasis id="F2D9EAD6081BFF91FC07779F1675D7D6" box="[943,1066,1181,1206]" italics="true" pageId="5" pageNumber="32">Diphipyga</emphasis>
|
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) and lack the characters defining the
|
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<taxonomicName id="07AD4D47081BFF91FF5A77C71322D7BE" box="[242,381,1221,1246]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="F2D9EAD6081BFF91FF5A77C71322D7BE" box="[242,381,1221,1246]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
|
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</taxonomicName>
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clade A. Also, preliminary molecular data (Zahniser and Dietrich, unpublished) suggest that each genus is monophyletic. Thus, retention of
|
||
<taxonomicName id="07AD4D47081BFF91FD5677EF1165D666" box="[766,826,1261,1286]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FD5677EF1165D666" box="[766,826,1261,1286]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
||
</taxonomicName>
|
||
as a genus separate from
|
||
<taxonomicName id="07AD4D47081BFF91FBD877EF16A5D666" box="[1136,1274,1261,1286]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FBD877EF16A5D666" box="[1136,1274,1261,1286]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
|
||
</taxonomicName>
|
||
is justified at this time. We did not identify any unambiguous characters that define
|
||
<taxonomicName id="07AD4D47081BFF91FC4876171635D64E" box="[992,1130,1301,1326]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FC4876171635D64E" box="[992,1130,1301,1326]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
|
||
</taxonomicName>
|
||
(including
|
||
<emphasis id="F2D9EAD6081BFF91FB5D7617172ED64E" box="[1269,1393,1301,1326]" italics="true" pageId="5" pageNumber="32">Diphipyga</emphasis>
|
||
) or that would unite
|
||
<taxonomicName id="07AD4D47081BFF91FEF4763F13C6D636" box="[348,409,1341,1366]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FEF4763F13C6D636" box="[348,409,1341,1366]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="07AD4D47081BFF91FE7A763F1003D636" box="[466,604,1341,1366]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FE7A763F1003D636" box="[466,604,1341,1366]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
|
||
</taxonomicName>
|
||
as a monophyletic group distinguishable from other chiasmine genera. Further collecting and morphological characterization of more species of
|
||
<taxonomicName id="07AD4D47081BFF91FC5176671669D61E" box="[1017,1078,1381,1406]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FC5176671669D61E" box="[1017,1078,1381,1406]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
||
</taxonomicName>
|
||
will undoubtedly help to clarify the relationship between these genera and other Old World chiasmine genera.
|
||
</paragraph>
|
||
<paragraph id="C01236C4081BFF91FF6E76B71003D5DF" blockId="5.[151,1438,701,1727]" pageId="5" pageNumber="32">
|
||
The apparent lack of unique and shared characters in
|
||
<taxonomicName id="07AD4D47081BFF91FC9376B71128D6AE" box="[827,887,1461,1486]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FC9376B71128D6AE" box="[827,887,1461,1486]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
||
</taxonomicName>
|
||
and overlap in character states with some species of
|
||
<taxonomicName id="07AD4D47081BFF91FF4476DF1329D696" box="[236,374,1501,1526]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FF4476DF1329D696" box="[236,374,1501,1526]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
|
||
</taxonomicName>
|
||
makes placing newly discovered species into
|
||
<taxonomicName id="07AD4D47081BFF91FC3F76DF118BD696" box="[919,980,1501,1526]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FC3F76DF118BD696" box="[919,980,1501,1526]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
||
</taxonomicName>
|
||
somewhat difficult, but a combination of some external characters cited by
|
||
<bibRefCitation id="A43C4B35081BFF91FDEF7507117DD57F" author="Linnavuori" box="[583,802,1541,1567]" pageId="5" pageNumber="39" refString="Linnavuori, R. (1973) Additional notes on the Cicadellidae fauna of Peru. Revista Peruana de Entomologia, 16, 14 - 16." type="journal article" year="1973">Linnavuori (1973)</bibRefCitation>
|
||
, especially of the head and face, and characters of the male genitalia identified here should separate
|
||
<taxonomicName id="07AD4D47081BFF91FD18752F10B3D526" box="[688,748,1581,1606]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FD18752F10B3D526" box="[688,748,1581,1606]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
||
</taxonomicName>
|
||
spp. from most
|
||
<taxonomicName id="07AD4D47081BFF91FC03752F166AD526" box="[939,1077,1581,1606]" class="Insecta" family="Cicadellidae" genus="Athysanella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FC03752F166AD526" box="[939,1077,1581,1606]" italics="true" pageId="5" pageNumber="32">Athysanella</emphasis>
|
||
</taxonomicName>
|
||
. Placement of the new species described here in
|
||
<taxonomicName id="07AD4D47081BFF91FECF755713FCD50E" box="[359,419,1621,1646]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FECF755713FCD50E" box="[359,419,1621,1646]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
||
</taxonomicName>
|
||
is based on the absence of characters defining other chiasmine genera, their similarity to other
|
||
<taxonomicName id="07AD4D47081BFF91FF55757F1366D5F6" box="[253,313,1661,1686]" class="Insecta" family="Cicadellidae" genus="Icaia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F2D9EAD6081BFF91FF55757F1366D5F6" box="[253,313,1661,1686]" italics="true" pageId="5" pageNumber="32">Icaia</emphasis>
|
||
</taxonomicName>
|
||
species externally, the short and falcate style apophysis, the mostly membranous segment X, and their South American distribution.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |