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<document id="42C5F0A2B175EB58D7BD1AC09D26D244" ID-DOI="10.11646/zootaxa.3963.2.1" ID-GBIF-Dataset="dedbd5f6-4d9e-4253-805d-a05d5c547bdf" ID-ISSN="1175-5326" ID-Zenodo-Dep="237609" ID-ZooBank="09EBB4DB-8A89-4E87-97E6-5372E8FD49B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461070129978" checkinUser="plazi" docAuthor="Stuke, Jens-Hermann" docDate="2015" docId="23492B464B6545743788A566FC0D43A1" docLanguage="en" docName="zt03963p159.pdf" docOrigin="Zootaxa 3963 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thecophora nigrifrons Stuke, 2015, spec. nov." docType="treatment" docVersion="7" lastPageNumber="117" masterDocId="DF70533E4B694564371FA035FFFF441A" masterDocTitle="New Conopid records from the Afrotropical Region (Diptera). Part 1: Paramyopa Kröber, Pseudoconops Camras, Stylogaster Macquart, Thecophora Rondani, and Zodion Latreille" masterLastPageNumber="159" masterPageNumber="101" pageNumber="113" updateTime="1698584218552" updateUser="plazi">
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<mods:title id="ADAF87C655B3EE2FD40FF8F0B0ED20E2">New Conopid records from the Afrotropical Region (Diptera). Part 1: Paramyopa Kröber, Pseudoconops Camras, Stylogaster Macquart, Thecophora Rondani, and Zodion Latreille</mods:title>
</mods:titleInfo>
<mods:name id="E88A3593ACD2326AC0D3C828EC5B2F1C" type="personal">
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<mods:namePart id="E5CD42F7C212753A20CC90F890D270D2">Stuke, Jens-Hermann</mods:namePart>
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<mods:typeOfResource id="BE4C048D444D9CA3600D6E3A87ACF203">text</mods:typeOfResource>
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<mods:title id="2A490A319D8A1A1C96042667E7F6596F">Zootaxa</mods:title>
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<mods:date id="F26E2F3A114E01A56C23DD457ADA8BB9">2015</mods:date>
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<mods:number id="8AFF6DB4E31756930333FCE17EF1F7B5">3963</mods:number>
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<mods:classification id="75A61B7F694FE0D1B53EAC8C391A73F7">journal article</mods:classification>
<mods:identifier id="30D7EA37AC54960E97B4CCE89EB04689" type="DOI">10.11646/zootaxa.3963.2.1</mods:identifier>
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<treatment id="23492B464B6545743788A566FC0D43A1" ID-DOI="http://doi.org/10.5281/zenodo.6097887" ID-GBIF-Taxon="119615777" ID-Zenodo-Dep="6097887" LSID="urn:lsid:plazi:treatment:23492B464B6545743788A566FC0D43A1" httpUri="http://treatment.plazi.org/id/23492B464B6545743788A566FC0D43A1" lastPageId="16" lastPageNumber="117" pageId="12" pageNumber="113">
<subSubSection id="E3FAC9DB4B6545683788A566FED84194" pageId="12" pageNumber="113" type="nomenclature">
<paragraph id="AB5F9A504B6545683788A566FDD54177" blockId="12.[151,554,1363,1422]" box="[151,554,1363,1389]" pageId="12" pageNumber="113">
<heading id="F0172D3C4B6545683788A566FDD54177" bold="true" box="[151,554,1363,1389]" fontSize="11" level="1" pageId="12" pageNumber="113" reason="1">
<emphasis id="999446424B6545683788A566FDD54177" bold="true" box="[151,554,1363,1389]" pageId="12" pageNumber="113">
<taxonomicName id="6CE0E1D34B6545683788A566FE544177" ID-CoL="8FDXB" box="[151,427,1363,1389]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="nigrifrons" status="sp. nov.">
<emphasis id="999446424B6545683788A566FE544177" bold="true" box="[151,427,1363,1389]" italics="true" pageId="12" pageNumber="113">Thecophora nigrifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82A7FB394B65456836ADA566FDD54177" box="[434,554,1363,1389]" pageId="12" pageNumber="113" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="AB5F9A504B6545683788A540FED84194" blockId="12.[151,554,1363,1422]" box="[151,295,1397,1422]" pageId="12" pageNumber="113">
(
<figureCitation id="33DB86D54B6545683780A540FEE04194" box="[159,287,1397,1422]" captionStart-0="FIGURES 21 24" captionStart-1="FIGURES 25 28" captionStartId-0="14.[151,264,1846,1869]" captionStartId-1="15.[151,264,1287,1310]" captionTargetBox-0="[151,1436,193,1824]" captionTargetBox-1="[151,1436,193,1265]" captionTargetId-0="figure@14.[151,1436,193,1826]" captionTargetId-1="figure@15.[151,1436,193,1266]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 21 24. Thecophora nigrifrons spec. nov. (♀, holotype). 21. Lateral view of habitus. 22. Dorsal view of frons. 23. Lateral view of antennae. 24. Posterior view of theca." captionText-1="FIGURES 25 28. ♂ postabdomen of Thecophora nigrifrons spec. nov. (♂, Kakamega forest). 25. Lateral view of phallus and hypandrium. 26. Lateral view of surstylus. 27. Dorsal view of phallus and hypandrium. 28. Frontal view of phallus and hypandrium. che—lateral connection hypandrium—epandrium; dp—distiphallus; epp—epiphallus; hya—hypandrial arms; hym—hypandrial membrane; hyr—hypandrial bars; hys—anteriorly directed phallus sheath of hypandrium; ipoe—inner plate of postgonite evagination; opoe—outer plate of postgonite evagination; pa—phallus apodeme; pdp—sclerotised plates on distiphallus; po—postgonite; poe—postgonite evagination; psu—posterior surstylus; rs—ring sclerite of phallus." httpUri-0="https://zenodo.org/record/237616/files/figure.png" httpUri-1="https://zenodo.org/record/237617/files/figure.png" pageId="12" pageNumber="113">Figs 2128</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E3FAC9DB4B6545683788A588FBDB4324" pageId="12" pageNumber="113" type="materials_examined">
<paragraph id="AB5F9A504B6545683788A588FF0E4303" blockId="12.[151,1437,1469,2034]" pageId="12" pageNumber="113">
<emphasis id="999446424B6545683788A588FE4341CC" bold="true" box="[151,444,1469,1494]" pageId="12" pageNumber="113">
<typeStatus id="745B24F24B6545683788A588FF2D41CC" box="[151,210,1469,1494]" pageId="12" pageNumber="113">Type</typeStatus>
material.
<typeStatus id="745B24F24B6545683650A588FE4341CC" box="[335,444,1469,1494]" pageId="12" pageNumber="113" type="holotype">Holotype</typeStatus>
</emphasis>
♀: (1)
<collectingCountry id="D3F7DAC04B654568350BA58BFD9F41CC" box="[532,608,1470,1494]" name="Kenya" pageId="12" pageNumber="113">Kenya</collectingCountry>
, Western Prov. /Kakamega forest / 014.13N, 3451.87E /
<date id="DF5EBC904B65456833E5A588FA6841CC" box="[1274,1431,1469,1494]" pageId="12" pageNumber="113" value="1999-11-06" valueMax="1999-11-13" valueMin="1999-11-06">6.13.xi.1999</date>
, Malaise / trap, R. Copeland; (2)
<typeStatus id="745B24F24B6545683515A5D4FD8041E0" box="[522,639,1505,1530]" pageId="12" pageNumber="113" type="holotype">Holotypus</typeStatus>
/
<taxonomicName id="6CE0E1D34B6545683588A5D4FC6941E0" box="[663,918,1505,1530]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="nigrifrons">
<emphasis id="999446424B6545683588A5D4FC6941E0" box="[663,918,1505,1530]" italics="true" pageId="12" pageNumber="113">Thecophora nigrifrons</emphasis>
</taxonomicName>
♀ / spec. nov. / det.
<bibRefCitation id="CF71E7A14B6545683398A5D4FAED41E0" author="Stuke" box="[1159,1298,1505,1530]" pageId="12" pageNumber="113" refString="Stuke, J. - H. (2014) A new genus of Conopidae from Europe (Diptera). Zootaxa, 3795, 472 - 482. http: // dx. doi. org / 10.11646 / zootaxa. 3795.4.6" type="journal article" year="2014">Stuke, 2014</bibRefCitation>
.
<emphasis id="999446424B6545683202A5D4FA6341E0" bold="true" box="[1309,1436,1505,1530]" pageId="12" pageNumber="113">
<typeStatus id="745B24F24B6545683202A5D4FA6841E0" box="[1309,1431,1505,1530]" pageId="12" pageNumber="113" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="D3F7DAC04B6545683788A633FEC34204" box="[151,316,1542,1566]" name="Cameroon" pageId="12" pageNumber="113">CAMEROON</collectingCountry>
:
<specimenCount id="BDE651D94B6545683658A633FE934204" box="[327,364,1542,1566]" pageId="12" pageNumber="113" type="generic">1♂</specimenCount>
,
<date id="DF5EBC904B6545683665A630FE0B4204" box="[378,500,1541,1566]" pageId="12" pageNumber="113" value="1987-11-18">18.xi.1987</date>
, Bambalang off Rt. N11, 35 km E. Bamenda,
<quantity id="6C1837B54B6545683319A633FBA04207" box="[1030,1119,1542,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="113" unit="m" value="1200.0">1200 m</quantity>
, leg. F. Kaplan, coll.
<collectionCode id="CDF102954B654568324AA633FA664204" box="[1365,1433,1542,1566]" pageId="12" pageNumber="113">TAUI</collectionCode>
, identified as
<taxonomicName id="6CE0E1D34B6545683637A61CFDD14258" box="[296,558,1577,1602]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="999446424B6545683637A61CFDD14258" box="[296,558,1577,1602]" italics="true" pageId="12" pageNumber="113">Thecophora flavicornis</emphasis>
</taxonomicName>
var. by Camras 2000; 1♀,
<date id="DF5EBC904B6545683441A61CFC274258" box="[862,984,1577,1602]" pageId="12" pageNumber="113" value="1987-11-20">20.xi.1987</date>
, Rt. N6, Bali-Batibo, West of Bamenda, leg. F. Kaplan, coll.
<collectionCode id="CDF102954B654568369AA67BFE36427C" box="[389,457,1614,1638]" pageId="12" pageNumber="113">TAUI</collectionCode>
, identified as
<taxonomicName id="6CE0E1D34B6545683573A678FC8A427C" box="[620,885,1613,1638]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="999446424B6545683573A678FC8A427C" box="[620,885,1613,1638]" italics="true" pageId="12" pageNumber="113">Thecophora flavicornis</emphasis>
</taxonomicName>
var. by Camras 2000;
<collectingCountry id="D3F7DAC04B654568339EA67BFB1A427C" box="[1153,1253,1614,1638]" name="Kenya" pageId="12" pageNumber="113">KENYA</collectingCountry>
: 1♀,
<date id="DF5EBC904B654568323AA67BFA67427C" box="[1317,1432,1614,1638]" pageId="12" pageNumber="113" value="1987-10-17">17.x.1987</date>
, Western Province, Kakamega Forest, leg. R. Copeland, coll.
<collectionCode id="CDF102954B654568345DA647FC674293" box="[834,920,1650,1673]" httpUri="http://grbio.org/cool/dhtn-7yaw" name="National Museum of Kenya" pageId="12" pageNumber="113">NMKE</collectionCode>
; 1♀,
<date id="DF5EBC904B65456834CAA647FBBB4290" box="[981,1092,1650,1674]" pageId="12" pageNumber="113" value="1991-05-11">11.v.1991</date>
, Western Province, Kakamega Forest, leg. A. Freidberg &amp; F. Kaplan, coll.
<collectionCode id="CDF102954B6545683598A6A3FD3442B4" box="[647,715,1686,1710]" pageId="12" pageNumber="113">TAUI</collectionCode>
, identified as
<taxonomicName id="6CE0E1D34B6545683476A6A0FB8F42B4" box="[873,1136,1685,1710]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="999446424B6545683476A6A0FB8F42B4" box="[873,1136,1685,1710]" italics="true" pageId="12" pageNumber="113">Thecophora flavicornis</emphasis>
</taxonomicName>
var. by Camras 2000; 1♀,
<date id="DF5EBC904B6545683788A68CFEEE42C8" box="[151,273,1721,1746]" pageId="12" pageNumber="113" value="2005-09-23">23.ix.2005</date>
, Western Province, Kakamega Forest [013N 3452E], leg. A. Freidberg, coll.
<collectionCode id="CDF102954B65456833BBA68FFB0542CB" box="[1188,1274,1722,1745]" httpUri="http://grbio.org/cool/dhtn-7yaw" name="National Museum of Kenya" pageId="12" pageNumber="113">NMKE</collectionCode>
;
<specimenCount id="BDE651D94B6545683216A68FFAD342C8" box="[1289,1324,1722,1746]" pageId="12" pageNumber="113" type="generic">1♂</specimenCount>
1♀,
<date id="DF5EBC904B6545683274A68FFEEE42EC" pageId="12" pageNumber="113" value="1999-11-20" valueMax="1999-11-27" valueMin="1999-11-20">20. 27.xi.1999</date>
, Western Province, Kakamega Forest, malaise trap [014.13N 3451.87E], leg. R. Copeland, coll.
<collectionCode id="CDF102954B6545683788A737FF114303" box="[151,238,1794,1817]" httpUri="http://grbio.org/cool/dhtn-7yaw" name="National Museum of Kenya" pageId="12" pageNumber="113">NMKE</collectionCode>
.
</paragraph>
<paragraph id="AB5F9A504B65456837D8A710FBDB4324" blockId="12.[151,1437,1469,2034]" box="[199,1060,1829,1854]" pageId="12" pageNumber="113">
<typeStatus id="745B24F24B65456837D8A710FED04324" box="[199,303,1829,1854]" pageId="12" pageNumber="113" type="holotype">Holotype</typeStatus>
is deposited in the
<collectionCode id="CDF102954B6545683517A713FDA14327" box="[520,606,1830,1853]" httpUri="http://grbio.org/cool/dhtn-7yaw" name="National Museum of Kenya" pageId="12" pageNumber="113">NMKE</collectionCode>
. The
<typeStatus id="745B24F24B6545683584A710FD024324" box="[667,765,1829,1854]" pageId="12" pageNumber="113" type="holotype">holotype</typeStatus>
is in very good condition.
</paragraph>
</subSubSection>
<subSubSection id="E3FAC9DB4B65457437D8A77CFD4F4281" lastPageId="16" lastPageNumber="117" pageId="12" pageNumber="113" type="description">
<paragraph id="AB5F9A504B65457437D8A77CFA724009" blockId="12.[151,1437,1469,2034]" lastBlockId="16.[151,1437,151,1979]" lastPageId="16" lastPageNumber="117" pageId="12" pageNumber="113">
<emphasis id="999446424B65456837D8A77CFDAB4378" bold="true" box="[199,596,1865,1890]" pageId="12" pageNumber="113">
Description of
<typeStatus id="745B24F24B6545683664A77CFE1D4378" box="[379,482,1865,1890]" pageId="12" pageNumber="113" type="holotype">holotype</typeStatus>
(female).
</emphasis>
Body length
<quantity id="6C1837B54B65456835ECA77CFCB1437B" box="[755,846,1865,1890]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="113" unit="mm" value="5.2">5.2 mm</quantity>
. Wing-length
<quantity id="6C1837B54B65456834EEA77FFBB3437B" box="[1009,1100,1866,1890]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="113" unit="mm" value="4.2">4.2 mm</quantity>
. Head-height
<quantity id="6C1837B54B65456833F2A77FFAB6437B" box="[1261,1353,1866,1890]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="113" unit="mm" value="1.1">1.1 mm</quantity>
.
<emphasis id="999446424B654568324AA77CFA634378" bold="true" box="[1365,1436,1865,1890]" pageId="12" pageNumber="113">Head.</emphasis>
Antenna orange-brown, basal flagellomere black distally and dorsally. Arista with 1 orange-brown aristomere situated dorsally on basal flagellomere. Aristomere about as long as basal flagellomere, broad at base and becoming abruptly narrower in the apical 2/3. Scape distinctly longer than broad, apically with a few bristles which are shorter than scape. Pedicel longer than maximum width, completely covered with short black setae. Basal flagellomere distinctly longer than high, and about as long as pedicel. Ocellar tubercle with three distinct brown ocelli. Ocellar triangle evident, about 1/2 of frons length. Frons black, with only a small yellow spot above antennae. Obvious silver dusting laterally at eye margin and an indistinct more-or-less v-shaped golden patch in front of the ocellar triangle. Black hairs all over frons which might represent frontal and orbital setae. Face orangebrown and slightly silver dusted, facial grooves shining. Gena and facial ridge with scattered black hairs. Postcranium black dorsally, becoming blackish brown ventrally. Adjacent to ventral posterior margin of eye there is a stripe of dense silver dusting; postgenae slightly silver dusted, remaining postcranium slightly dusted to shining. Postcranium with black hairs. No hairs on a small area adjacent to eye margin and bottom portion of postcranium. Median occipital sclerite with only a few scattered black hairs. Proboscis black, with an indistinct brown spot between frontoclypeal membrane and mouth edge. Frontoclypeal membrane narrow, widened basally, yellowish brown and distinctly separated from the brown clypeus. Palps long (longer than scape) blackish brown, hardly widened distally, and covered with long black hairs. Labium about as long as head-length, projecting out of mouth opening, hardly thickened basally; anterior section not fused into a tube. Labellum as long as labium, only divided apically, slightly narrower than adjacent haustellum, and covered with scattered hairs. Head with distinct ocellar setae, postocellar setae, 1 distinct medial and 2 lateral postocular setae.
<emphasis id="999446424B66456B3335A734FB754300" bold="true" box="[1066,1162,1793,1818]" pageId="15" pageNumber="116">Thorax.</emphasis>
Thorax blackish brown without any reflection, completely grey dusted, with only the anepisternum shining in ventral 2/3. Postpronotum and scutum along the transverse suture denser dusted. Basisternum narrow, narrowed to a point, with 34 black hairs apically. Proepisternum with 2 black hairs ventrally and a few hairs dorsally. Scutellum covered with long white hairs and 2 pairs of scutellar bristles (the lateral scutellar bristles distinctly smaller). Scutum with short black hairs, about as long as tibia diameter. 1 postpronotal, 2 notopleural, 1 supraalar, 2 postalar, and 1 prescutellar bristle. 1 black bristle posterodorsally and several bristles ventrally on katepisternum. Metakatepisternum without bristles. Anepisternum and anepimeron hairless and without bristles. Wing tinged inconspiciously brownish. Veins yellowish white basally and black apically. Basal-medial-cubital-crossvein bm-cu incomplete. Vein A1+CuA2 not quite reaching hind margin of wing. Upper and lower calypters whitish brown, upper calypter with long white hairs on margin. Alula broad (about 2 times wider than long), with long hairs on posterior margin. Haltere yellowish white with brown base. Knob of haltere with short, indistinct, light yellow hairs. Legs brown, fore coxa, basal 2/3 of femora and basal 3 tarsomeres yellow. Legs slightly dusted to shining. Posterior surfaces of fore and middle tibiae without obvious dusted fields distally. Legs all with long, semiadpressed or erect black hairs.
<collectingCountry id="D3F7DAC04B7945743207A147FAAF4591" box="[1304,1360,370,395]" name="India" pageId="16" pageNumber="117">Hind</collectingCountry>
femur anterodorsally and posterordorsally with single outstanding long hairs. 12 prepreapical bristles dorsally on tibiae. Fore and middle tibiae basally on ventral surface with one distinct line of small, thick, adpressed black bristles. All femora distally on ventral surface with two parallel lines of small, thick, black bristles. Fore coxa without outstanding black bristle which is stronger than the surrounding scattered black hairs. Middle and hind coxae with some distinct strong lateral bristles which are about as long as diameter of basal hind tibia.
<collectingCountry id="D3F7DAC04B79457433FAA213FAE24625" box="[1253,1309,550,575]" name="India" pageId="16" pageNumber="117">Hind</collectingCountry>
femur not obviously thickened in basal half. Pulvilli yellow-whitish. Claws brown with black tips. Empodium yellowish brown.
<emphasis id="999446424B79457437EFA25BFE93469D" bold="true" box="[240,364,622,647]" pageId="16" pageNumber="117">Abdomen.</emphasis>
Abdomen blackish brown, and covered with black hairs only. Tergites 15 slightly dusted. Tergites 23 with grey dusted spots laterally on hind margins. The
<typeStatus id="745B24F24B7945743491A2A7FC0F46B1" box="[910,1008,658,683]" pageId="16" pageNumber="117" type="holotype">holotype</typeStatus>
was not dissected and the description of internal structures of the female abdomen is based on a
<typeStatus id="745B24F24B794574342BA28DFC6746CA" box="[820,920,696,720]" pageId="16" pageNumber="117" type="paratype">paratype</typeStatus>
. Abdomen parallel-sided. Length: maximum width of tergite 2 = 1.4; length: maximum width of tergite 3 = 1.1. Sternites 34 protruding ventrally, posterior parts of sternites conspicious. Shape of theca as shown in
<figureCitation id="33DB86D54B7945743403A2CBFC91470D" box="[796,878,766,791]" captionStart="FIGURES 21 24" captionStartId="14.[151,264,1846,1869]" captionTargetBox="[151,1436,193,1824]" captionTargetId="figure@14.[151,1436,193,1826]" captionTargetPageId="14" captionText="FIGURES 21 24. Thecophora nigrifrons spec. nov. (♀, holotype). 21. Lateral view of habitus. 22. Dorsal view of frons. 23. Lateral view of antennae. 24. Posterior view of theca." httpUri="https://zenodo.org/record/237616/files/figure.png" pageId="16" pageNumber="117">Fig. 24</figureCitation>
. Anterior surface of theca without hairs. Apical 1/ 3 of posterior surface of theca with 5 horizontal lines of close-set, short, blunt spicules. Sternite 6 almost completely covered with short, blunt spicules which are almost entirely arranged in 9 horizontal lines. Sternite 7 longer than broad, slightly pointed anteriorly. Area between sides of syntergite 8+9 entirely lightly sclerotised, and without a distinct sternite 8. Sack-like ventral protrusion of vagina with annular sclerotisation and not covered with bristles. Base of accessory glands slightly sclerotised. Spermathecal ducts not sclerotised at base. A dorsal sclerotisation is evident on the sack-like ventral protrusion of the vagina. 2 pairs of round spermathecae, spermathecal ducts fused shortly after leaving spermathecae. Spermathecal ducts not sclerotised at spermathecae.
</paragraph>
<caption id="FF9FCAD84B6445693788A781FB4643F3" httpUri="https://zenodo.org/record/237615/files/figure.png" pageId="13" pageNumber="114" targetBox="[151,1436,409,1949]" targetPageId="13">
<paragraph id="AB5F9A504B6445693788A781FB4643F3" blockId="13.[151,1435,1972,2025]" pageId="13" pageNumber="114">
<emphasis id="999446424B6445693788A781FEA643D3" bold="true" box="[151,345,1972,1995]" pageId="13" pageNumber="114">FIGURES 1820.</emphasis>
<taxonomicName id="6CE0E1D34B644569367BA781FD5843D0" authority="Camras." authorityName="Camras." box="[356,679,1972,1994]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="999446424B644569367BA781FDB743D0" box="[356,584,1972,1994]" italics="true" pageId="13" pageNumber="114">Thecophora metallica</emphasis>
Camras.
</taxonomicName>
<emphasis id="999446424B64456935ACA781FD3343D3" bold="true" box="[691,716,1972,1993]" pageId="13" pageNumber="114">18</emphasis>
. Lateral view of thorax and head (♂, 12 km w Ranomafana).
<emphasis id="999446424B6445693263A781FA6A43D3" bold="true" box="[1404,1429,1972,1993]" pageId="13" pageNumber="114">19</emphasis>
. Frontolateral view of head (♂, 12 km w Ranomafana).
<emphasis id="999446424B64456935DAA7E6FD2143F2" bold="true" box="[709,734,2003,2024]" pageId="13" pageNumber="114">20</emphasis>
. Posterior view of theca (♀, Ranomafana NP).
</paragraph>
</caption>
<caption id="FF9FCAD84B67456A3788A703FD494376" httpUri="https://zenodo.org/record/237616/files/figure.png" pageId="14" pageNumber="115" targetBox="[151,1436,193,1824]" targetPageId="14">
<paragraph id="AB5F9A504B67456A3788A703FD494376" blockId="14.[151,1435,1846,1900]" pageId="14" pageNumber="115">
<emphasis id="999446424B67456A3788A703FEAA4356" bold="true" box="[151,341,1846,1869]" pageId="14" pageNumber="115">FIGURES 2124.</emphasis>
<taxonomicName id="6CE0E1D34B67456A3643A702FDBD4357" box="[348,578,1847,1869]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="nigrifrons" status="sp. nov.">
<emphasis id="999446424B67456A3643A702FDBD4357" box="[348,578,1847,1869]" italics="true" pageId="14" pageNumber="115">Thecophora nigrifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82A7FB394B67456A3556A702FD4E4356" box="[585,689,1847,1868]" pageId="14" pageNumber="115" rank="species">
<emphasis id="999446424B67456A3556A702FD4E4356" bold="true" box="[585,689,1847,1868]" pageId="14" pageNumber="115">spec. nov.</emphasis>
</taxonomicNameLabel>
(♀, holotype).
<emphasis id="999446424B67456A344EA702FC954356" bold="true" box="[849,874,1847,1868]" pageId="14" pageNumber="115">21</emphasis>
. Lateral view of habitus.
<emphasis id="999446424B67456A336CA702FB734356" bold="true" box="[1139,1164,1847,1868]" pageId="14" pageNumber="115">22</emphasis>
. Dorsal view of frons.
<emphasis id="999446424B67456A3263A702FA6A4356" bold="true" box="[1404,1429,1847,1868]" pageId="14" pageNumber="115">23</emphasis>
. Lateral view of antennae.
<emphasis id="999446424B67456A3681A763FE484371" bold="true" box="[414,439,1878,1899]" pageId="14" pageNumber="115">24</emphasis>
. Posterior view of theca.
</paragraph>
</caption>
<caption id="FF9FCAD84B66456B3788A532FAF241AD" httpUri="https://zenodo.org/record/237617/files/figure.png" pageId="15" pageNumber="116" targetBox="[151,1436,193,1265]" targetPageId="15">
<paragraph id="AB5F9A504B66456B3788A532FAF241AD" blockId="15.[151,1436,1287,1463]" pageId="15" pageNumber="116">
<emphasis id="999446424B66456B3788A532FEAB4107" bold="true" box="[151,340,1287,1310]" pageId="15" pageNumber="116">FIGURES 2528.</emphasis>
♂ postabdomen of
<taxonomicName id="6CE0E1D34B66456B3503A53DFCFE4104" box="[540,769,1288,1310]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="nigrifrons" status="sp. nov.">
<emphasis id="999446424B66456B3503A53DFCFE4104" box="[540,769,1288,1310]" italics="true" pageId="15" pageNumber="116">Thecophora nigrifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82A7FB394B66456B3417A53DFC904107" box="[776,879,1288,1309]" pageId="15" pageNumber="116" rank="species">
<emphasis id="999446424B66456B3417A53DFC904107" bold="true" box="[776,879,1288,1309]" pageId="15" pageNumber="116">spec. nov.</emphasis>
</taxonomicNameLabel>
(♂, Kakamega forest).
<emphasis id="999446424B66456B337FA53DFB864107" bold="true" box="[1120,1145,1288,1309]" pageId="15" pageNumber="116">25</emphasis>
. Lateral view of phallus and hypandrium.
<emphasis id="999446424B66456B363DA512FEC44126" bold="true" box="[290,315,1319,1340]" pageId="15" pageNumber="116">26</emphasis>
. Lateral view of surstylus.
<emphasis id="999446424B66456B3579A512FD804126" bold="true" box="[614,639,1319,1340]" pageId="15" pageNumber="116">27</emphasis>
. Dorsal view of phallus and hypandrium.
<emphasis id="999446424B66456B3356A512FB9D4126" bold="true" box="[1097,1122,1319,1340]" pageId="15" pageNumber="116">28</emphasis>
. Frontal view of phallus and hypandrium. che—lateral connection hypandrium—epandrium; dp—distiphallus; epp—epiphallus; hya—hypandrial arms; hym—hypandrial membrane; hyr—hypandrial bars; hys—anteriorlydirected phallus sheath of hypandrium; ipoe—inner plate of postgonite evagination; opoe—outer plate of postgonite evagination; pa—phallus apodeme; pdp—sclerotised plates on distiphallus; po—postgonite; poe—postgonite evagination; psu—posterior surstylus; rs—ring sclerite of phallus.
</paragraph>
</caption>
<paragraph id="AB5F9A504B79457437D8A42BFBA34249" blockId="16.[151,1437,151,1979]" pageId="16" pageNumber="117">
<emphasis id="999446424B79457437D8A42BFDCE402D" bold="true" box="[199,561,1054,1079]" pageId="16" pageNumber="117">Description of male abdomen.</emphasis>
Tergite 1 with black hairs laterally which are slightly shorter than the lateral hairs on tergite 2. Tergite 2 elongated, distinctly longer than wide. Tergite 2 without obvious lateral hair tufts, but with slightly longer hairs than laterally on tergite 3. Tergite 3 not widened posteriorly. Sternites 15 present, sternites 1 and 2 to some extent fused but still distinguishable. Sternite 4 slightly broader than long and obviously smaller than sternite 5, with several long black hairs. Sternite 5 apically with a small field of thick black bristles and long hairs. Ventrally the lateral edges of the protandrium are fused by a narrow strip. Epandrium fused behind cerci. Posterior surstylus slightly pointed distally. A dense field of strong black bristles at base of posterior surstylus reaching up to anterior surstylus. Two inner bristles elongated, about as long as posterior surstylus. Subepandrial plate distinctly sclerotised and u-shaped. Phallus sheath slightly fused dorsally; apically without obvious evagination and without any tooth or bristles, dorsomedially without a notch. Postgonite more-or-less distinct. Postgonite evagination small, hardly sclerotised, not projecting above distiphallus and covered with microtrichia. Two plates at inner side of postgonite evagination, the inner one obvious but the outer plate much smaller and therefore hardly evident. Both plates partly fused together. A u-shaped ring sclerite is obvious. Epiphallus distinct, covered with microtrichia. Distiphallus short, about 1/2 of length of epandrium, densely covered with blunt microtrichia, without any evaginations but with indistinct sclerotised dorsolateral plates. Ejaculatory apodeme elongate, distinctly sclerotised, attachment to sperm sac evident.
</paragraph>
<paragraph id="AB5F9A504B79457437D8A66BFD4F4281" blockId="16.[151,1437,151,1979]" pageId="16" pageNumber="117">
<emphasis id="999446424B79457437D8A66BFEB2426D" bold="true" box="[199,333,1630,1655]" pageId="16" pageNumber="117">Variability.</emphasis>
Wing length
<quantity id="6C1837B54B79457436F5A66BFD8A426D" box="[490,629,1630,1655]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.3" metricValueMin="4.2" pageId="16" pageNumber="117" unit="mm" value="4.75" valueMax="5.3" valueMin="4.2">4.25.3 mm</quantity>
. The dusting on tergites 24 varies and the abdomen might be shining with only grey dusted spots at the hind margins.
</paragraph>
</subSubSection>
<subSubSection id="E3FAC9DB4B79457437D8A693FC2142F9" pageId="16" pageNumber="117" type="etymology">
<paragraph id="AB5F9A504B79457437D8A693FC2142F9" blockId="16.[151,1437,151,1979]" pageId="16" pageNumber="117">
<emphasis id="999446424B79457437D8A693FEB142A5" bold="true" box="[199,334,1702,1727]" pageId="16" pageNumber="117">Etymology.</emphasis>
The name is combined from the Latin nigra (black) and frons. The black frons is an obvious feature of the species, and the character which first drew my attention to it.
</paragraph>
</subSubSection>
<subSubSection id="E3FAC9DB4B79457437D8A6DBFDCE438D" pageId="16" pageNumber="117" type="discussion">
<paragraph id="AB5F9A504B79457437D8A6DBFDCE438D" blockId="16.[151,1437,151,1979]" pageId="16" pageNumber="117">
<emphasis id="999446424B79457437D8A6DBFEB3431D" bold="true" box="[199,332,1774,1799]" pageId="16" pageNumber="117">Discussion.</emphasis>
<taxonomicName id="6CE0E1D34B7945743649A6DBFDA8431D" box="[342,599,1774,1799]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="nigrifrons">
<emphasis id="999446424B7945743649A6DBFDA8431D" box="[342,599,1774,1799]" italics="true" pageId="16" pageNumber="117">Thecophora nigrifrons</emphasis>
</taxonomicName>
is close to
<taxonomicName id="6CE0E1D34B79457435FBA6DBFC45431D" box="[740,954,1774,1799]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="999446424B79457435FBA6DBFC45431D" box="[740,954,1774,1799]" italics="true" pageId="16" pageNumber="117">Thecophora pilosa</emphasis>
</taxonomicName>
, which has light yellow legs and a more distinctly yellowish wing base (
<figureCitation id="33DB86D54B79457436E5A727FDB14331" box="[506,590,1810,1835]" captionStart="FIGURES 21 24" captionStartId="14.[151,264,1846,1869]" captionTargetBox="[151,1436,193,1824]" captionTargetId="figure@14.[151,1436,193,1826]" captionTargetPageId="14" captionText="FIGURES 21 24. Thecophora nigrifrons spec. nov. (♀, holotype). 21. Lateral view of habitus. 22. Dorsal view of frons. 23. Lateral view of antennae. 24. Posterior view of theca." httpUri="https://zenodo.org/record/237616/files/figure.png" pageId="16" pageNumber="117">Fig. 21</figureCitation>
). The mainly black frons (
<figureCitation id="33DB86D54B7945743469A727FC374331" box="[886,968,1810,1835]" captionStart="FIGURES 21 24" captionStartId="14.[151,264,1846,1869]" captionTargetBox="[151,1436,193,1824]" captionTargetId="figure@14.[151,1436,193,1826]" captionTargetPageId="14" captionText="FIGURES 21 24. Thecophora nigrifrons spec. nov. (♀, holotype). 21. Lateral view of habitus. 22. Dorsal view of frons. 23. Lateral view of antennae. 24. Posterior view of theca." httpUri="https://zenodo.org/record/237616/files/figure.png" pageId="16" pageNumber="117">Fig. 22</figureCitation>
) is a first clue in recognising
<taxonomicName id="6CE0E1D34B794574320BA727FEF94355" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="nigrifrons">
<emphasis id="999446424B794574320BA727FEF94355" italics="true" pageId="16" pageNumber="117">Thecophora nigrifrons</emphasis>
</taxonomicName>
. More important, however, is the different shape of the theca in female, which is broader and has 45 distinct lines of black spicules in
<taxonomicName id="6CE0E1D34B794574350FA769FD644369" box="[528,667,1883,1907]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="nigrifrons">
<emphasis id="999446424B794574350FA769FD644369" box="[528,667,1883,1907]" italics="true" pageId="16" pageNumber="117">T. nigrifrons</emphasis>
</taxonomicName>
(
<figureCitation id="33DB86D54B79457435B4A76FFCFF4369" box="[683,768,1882,1907]" captionStart="FIGURES 21 24" captionStartId="14.[151,264,1846,1869]" captionTargetBox="[151,1436,193,1824]" captionTargetId="figure@14.[151,1436,193,1826]" captionTargetPageId="14" captionText="FIGURES 21 24. Thecophora nigrifrons spec. nov. (♀, holotype). 21. Lateral view of habitus. 22. Dorsal view of frons. 23. Lateral view of antennae. 24. Posterior view of theca." httpUri="https://zenodo.org/record/237616/files/figure.png" pageId="16" pageNumber="117">Fig. 24</figureCitation>
). The epandrium of male
<taxonomicName id="6CE0E1D34B794574333BA769FB504369" box="[1060,1199,1883,1907]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="nigrifrons">
<emphasis id="999446424B794574333BA769FB504369" box="[1060,1199,1883,1907]" italics="true" pageId="16" pageNumber="117">T. nigrifrons</emphasis>
</taxonomicName>
has no long, pointed setae close to the surstylus (
<figureCitation id="33DB86D54B79457436D0A74BFDDB438D" box="[463,548,1918,1943]" captionStart="FIGURES 25 28" captionStartId="15.[151,264,1287,1310]" captionTargetBox="[151,1436,193,1265]" captionTargetId="figure@15.[151,1436,193,1266]" captionTargetPageId="15" captionText="FIGURES 25 28. ♂ postabdomen of Thecophora nigrifrons spec. nov. (♂, Kakamega forest). 25. Lateral view of phallus and hypandrium. 26. Lateral view of surstylus. 27. Dorsal view of phallus and hypandrium. 28. Frontal view of phallus and hypandrium. che—lateral connection hypandrium—epandrium; dp—distiphallus; epp—epiphallus; hya—hypandrial arms; hym—hypandrial membrane; hyr—hypandrial bars; hys—anteriorly directed phallus sheath of hypandrium; ipoe—inner plate of postgonite evagination; opoe—outer plate of postgonite evagination; pa—phallus apodeme; pdp—sclerotised plates on distiphallus; po—postgonite; poe—postgonite evagination; psu—posterior surstylus; rs—ring sclerite of phallus." httpUri="https://zenodo.org/record/237617/files/figure.png" pageId="16" pageNumber="117">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E3FAC9DB4B79457437D8A797FC0D43A1" box="[199,1010,1954,1979]" pageId="16" pageNumber="117" type="distribution">
<paragraph id="AB5F9A504B79457437D8A797FC0D43A1" blockId="16.[151,1437,151,1979]" box="[199,1010,1954,1979]" pageId="16" pageNumber="117">
<emphasis id="999446424B79457437D8A797FE9F43A1" bold="true" box="[199,352,1954,1979]" pageId="16" pageNumber="117">Distribution.</emphasis>
<taxonomicName id="6CE0E1D34B7945743678A797FD9C43A1" box="[359,611,1954,1979]" class="Insecta" family="Conopidae" genus="Thecophora" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="nigrifrons">
<emphasis id="999446424B7945743678A797FD9C43A1" box="[359,611,1954,1979]" italics="true" pageId="16" pageNumber="117">Thecophora nigrifrons</emphasis>
</taxonomicName>
is only known from
<collectingCountry id="D3F7DAC04B7945743451A796FC6843A1" box="[846,919,1955,1979]" name="Kenya" pageId="16" pageNumber="117">Kenya</collectingCountry>
to date.
</paragraph>
</subSubSection>
</treatment>
</document>