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<document id="C65846D96962413830A327FE24CE3E10" ID-CLB-Dataset="5710" ID-DOI="10.11646/zootaxa.4985.3.11" ID-GBIF-Dataset="7dc8160d-98f9-4c8a-8382-994e97b7d71f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4955877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623756031967" checkinUser="plazi" docAuthor="Makarchenko Usgs, Eugenyi A." docDate="2021" docId="0B149642A5752D4241DCFAEDFD35FB5E" docLanguage="en" docName="zootaxa.4985.3.11.pdf" docOrigin="Zootaxa 4985 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diamesa leoniella Hansen 1976" docType="treatment" docVersion="9" lastPageNumber="440" masterDocId="F72DEE3AA5752D43414BFF9FFFF1FF9A" masterDocTitle="Confirmation of a species status of Diamesa leoniella Hansen, 1976, stat. resurr (Diptera, Chironomidae) from North America" masterLastPageNumber="442" masterPageNumber="439" pageNumber="439" updateTime="1699016569100" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7E2B810E700D3E3BF09E5F2684BE0DE8">Confirmation of a species status of Diamesa leoniella Hansen, 1976, stat. resurr (Diptera, Chironomidae) from North America</mods:title>
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<mods:namePart id="F3D1E3D7AA3DBFA6861ABCA3EB6D8C03">Makarchenko Usgs, Eugenyi A.</mods:namePart>
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<mods:typeOfResource id="3D128D690C70DBEF8CDE949F6446AF0F">text</mods:typeOfResource>
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<mods:title id="5C852304630ACEC35D14E11F9D9E9BC3">Zootaxa</mods:title>
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<mods:date id="04B38EF0D664D312BAE42B733BE620FF">2021</mods:date>
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<mods:number id="360D298F8EDB479003C76140D46D9BF7">2021-06-15</mods:number>
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<mods:number id="19D0E3B82896BC42CD6B73B404D3F30D">4985</mods:number>
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<mods:number id="C104A720D7C5F712BF97694EF543D853">3</mods:number>
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<mods:classification id="6823868DDEE34AA3DEF8D9CF4ED69BDC">journal article</mods:classification>
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<mods:identifier id="54922513E10337E2F1DA5157EC9DB481" type="DOI">10.11646/zootaxa.4985.3.11</mods:identifier>
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<treatment id="0B149642A5752D4241DCFAEDFD35FB5E" ID-DOI="http://doi.org/10.5281/zenodo.5073283" ID-GBIF-Taxon="183325884" ID-Zenodo-Dep="5073283" LSID="urn:lsid:plazi:treatment:0B149642A5752D4241DCFAEDFD35FB5E" httpUri="http://treatment.plazi.org/id/0B149642A5752D4241DCFAEDFD35FB5E" lastPageId="1" lastPageNumber="440" pageId="0" pageNumber="439">
<subSubSection id="CBA774DFA5752D4341DCFAEDFD4EFA16" box="[151,703,1393,1420]" pageId="0" pageNumber="439" type="nomenclature">
<paragraph id="83022754A5752D4341DCFAEDFD4EFA16" blockId="0.[151,1437,1358,2027]" box="[151,703,1393,1420]" pageId="0" pageNumber="439">
<heading id="D84A9038A5752D4341DCFAEDFD4EFA16" bold="true" box="[151,703,1393,1420]" fontSize="11" level="1" pageId="0" pageNumber="439" reason="1">
<emphasis id="B1C9FB46A5752D4341DCFAEDFD4EFA16" bold="true" box="[151,703,1393,1420]" pageId="0" pageNumber="439">
<taxonomicName id="44BD5CD7A5752D4341DCFAEDFDD0FA16" ID-CoL="359FB" authority="Hansen, 1976" authorityName="Hansen" authorityYear="1976" box="[151,545,1393,1420]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="leoniella">
<emphasis id="B1C9FB46A5752D4341DCFAEDFE81FA16" bold="true" box="[151,368,1393,1420]" italics="true" pageId="0" pageNumber="439">Diamesa leoniella</emphasis>
<bibRefCitation id="E72C5AA5A5752D434033FAEDFDD0FA16" author="Hansen, D. C. &amp; Cook, E. F." box="[376,545,1394,1420]" pageId="0" pageNumber="439" pagination="1 - 203" refId="ref2327" refString="Hansen, D. C. &amp; Cook, E. F. (1976) The systematics and morphology of the Nearctic species of Diamesa Meigen, 1835 (Diptera, Chironomidae). Memoirs of the American Entomological Society, 30, 1 - 203." type="journal article" year="1976">Hansen, 1976</bibRefCitation>
</taxonomicName>
, stat. resurr.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CBA774DFA5752D4341DCFA07FEFDFA2A" box="[151,268,1432,1456]" pageId="0" pageNumber="439" type="description">
<paragraph id="83022754A5752D4341DCFA07FEFDFA2A" blockId="0.[151,1437,1358,2027]" box="[151,268,1432,1456]" pageId="0" pageNumber="439">
(
<figureCitation id="1B863BD1A5752D4341D4FA07FEF5FA2A" box="[159,260,1432,1456]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1950,1974]" captionTargetBox="[151,1435,596,1927]" captionTargetId="figure-144@2.[151,1435,596,1927]" captionTargetPageId="2" captionText="FIGURES 1–4. Males of Diamesa leoniella Hansen (1–2) and D. japonica Tokunaga (3–4). 1, 4, central part of hypopygium without tergite IX; 2–3, hypopygium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4955879" httpUri="https://zenodo.org/record/4955879/files/figure.png" pageId="0" pageNumber="439">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CBA774DFA5752D4341DCFA28FD29FA55" box="[151,728,1462,1487]" pageId="0" pageNumber="439" type="reference_group">
<paragraph id="83022754A5752D4341DCFA28FD29FA55" blockId="0.[151,1437,1358,2027]" box="[151,728,1462,1487]" pageId="0" pageNumber="439">
<treatmentCitationGroup id="A3AD007AA5752D4341DCFA28FD29FA55" box="[151,728,1462,1487]" pageId="0" pageNumber="439">
<taxonomicName id="44BD5CD7A5752D4341DCFA28FE52FA54" ID-CoL="359FB" authority="Hansen" authorityName="Hansen" authorityYear="1976" box="[151,419,1462,1486]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="leoniella">
<emphasis id="B1C9FB46A5752D4341DCFA28FEA0FA54" box="[151,337,1462,1486]" italics="true" pageId="0" pageNumber="439">Diamesa leoniella</emphasis>
Hansen
</taxonomicName>
in
<treatmentCitation id="021C0145A5752D434088FA28FD25FA55" author="Hansen, D. C. &amp; Cook, E. F." box="[451,724,1462,1487]" page="111" pageId="0" pageNumber="439" year="1976">
<bibRefCitation id="E72C5AA5A5752D434088FA28FD25FA55" author="Hansen, D. C. &amp; Cook, E. F." box="[451,724,1462,1487]" pageId="0" pageNumber="439" pagination="1 - 203" refId="ref2327" refString="Hansen, D. C. &amp; Cook, E. F. (1976) The systematics and morphology of the Nearctic species of Diamesa Meigen, 1835 (Diptera, Chironomidae). Memoirs of the American Entomological Society, 30, 1 - 203." type="journal article" year="1976">Hansen &amp; Cook 1976: 111</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CBA774DFA5752D4241DCFA45FA97FBE6" lastPageId="1" lastPageNumber="440" pageId="0" pageNumber="439" type="materials_examined">
<paragraph id="83022754A5752D4341DCFA45FEC8F9A0" blockId="0.[151,1437,1358,2027]" pageId="0" pageNumber="439">
<emphasis id="B1C9FB46A5752D4341DCFA45FE86FA68" bold="true" box="[151,375,1498,1522]" inLineHeading="true" pageId="0" pageNumber="439" reason="1">Material examined.</emphasis>
<materialsCitation id="33D52D09A5752D434035FA44FE16F98C" ID-GBIF-Occurrence="3314008303" collectingDate="2013-03-26" collectorName="K. Frangos &amp; P. Hudson" country="United States of America" latitude="58.4217" location="Mendenhall Lake" longLatPrecision="6" longitude="134.5388" municipality="Juneau City" pageId="0" pageNumber="439" specimenCount="1" specimenCount-adult="1" stateProvince="Alaska">
<specimenCount id="95BBECDDA5752D434035FA44FE39FA68" box="[382,456,1498,1523]" pageId="0" pageNumber="439" type="adult">1 adult</specimenCount>
male,
<collectingCountry id="FBAA67C4A5752D434358FA44FD91FA69" box="[531,608,1499,1523]" name="United States of America" pageId="0" pageNumber="439">U.S.A.</collectingCountry>
:
<collectingRegion id="4179E9B6A5752D434321FA45FD46FA68" box="[618,695,1498,1522]" country="United States of America" name="Alaska" pageId="0" pageNumber="439">Alaska</collectingRegion>
,
<collectingMunicipality id="6366BD2EA5752D434389FA44FCB2FA68" box="[706,835,1498,1522]" pageId="0" pageNumber="439">Juneau City</collectingMunicipality>
,
<location id="8662718FA5752D434204FA45FBFFFA68" LSID="urn:lsid:plazi:treatment:0B149642A5752D4241DCFAEDFD35FB5E:8662718FA5752D434204FA45FBFFFA68" box="[847,1038,1498,1522]" country="United States of America" latitude="58.4217" longLatPrecision="6" longitude="134.5388" municipality="Juneau City" name="Mendenhall Lake" pageId="0" pageNumber="439" stateProvince="Alaska">Mendenhall Lake</location>
,
<geoCoordinate id="E6894193A5752D434551FA44FB7DFA68" box="[1050,1164,1498,1522]" degrees="58.4217" direction="north" orientation="latitude" pageId="0" pageNumber="439" precision="5" value="58.4217">N 58.4217</geoCoordinate>
<geoCoordinate id="E6894193A5752D4345D8FA44FAE3FA68" box="[1171,1298,1498,1522]" degrees="134.5388" direction="east" orientation="longitude" pageId="0" pageNumber="439" precision="5" value="134.5388">E 134.5388</geoCoordinate>
,
<date id="F7030194A5752D434455FA45FA69FA68" box="[1310,1432,1498,1522]" pageId="0" pageNumber="439" value="2013-03-26">
<collectingDate id="E747F87CA5752D434455FA45FA69FA68" box="[1310,1432,1498,1522]" pageId="0" pageNumber="439" value="2013-03-26">26.III.2013</collectingDate>
</date>
, leg.
<collectorName id="2E484282A5752D43418CFA60FECEF98D" box="[199,319,1535,1559]" pageId="0" pageNumber="439">K. Frangos</collectorName>
and
<collectorName id="2E484282A5752D43403DFA60FE16F98C" box="[374,487,1534,1558]" pageId="0" pageNumber="439">P. Hudson</collectorName>
</materialsCitation>
;
<materialsCitation id="33D52D09A5752D4340B8FA60FEC8F9A0" ID-GBIF-Occurrence="3314008302" collectingDate="2013-07-17" collectorName="P. Hudson." country="United States of America" latitude="58.427" location="Nugget Falls" longLatPrecision="6" longitude="134.5373" municipality="Juneau City" pageId="0" pageNumber="439" specimenCount="1" specimenCount-adult="1" stateProvince="Alaska">
<specimenCount id="95BBECDDA5752D4340B8FA60FDCFF98C" box="[499,574,1534,1559]" pageId="0" pageNumber="439" type="adult">1 adult</specimenCount>
male, the same data except
<location id="8662718FA5752D43423FFA60FBF1F98C" LSID="urn:lsid:plazi:treatment:0B149642A5752D4241DCFAEDFD35FB5E:8662718FA5752D43423FFA60FBF1F98C" box="[884,1024,1534,1559]" latitude="58.427" longLatPrecision="6" longitude="134.5373" name="Nugget Falls" pageId="0" pageNumber="439">Nugget Falls</location>
,
<geoCoordinate id="E6894193A5752D434546FA60FB71F98C" box="[1037,1152,1534,1558]" degrees="58.4270" direction="north" orientation="latitude" pageId="0" pageNumber="439" precision="5" value="58.427">N 58.4270</geoCoordinate>
<geoCoordinate id="E6894193A5752D4345C3FA60FAF6F98C" box="[1160,1287,1534,1558]" degrees="134.5373" direction="east" orientation="longitude" pageId="0" pageNumber="439" precision="5" value="134.5373">E 134.5373</geoCoordinate>
,
<date id="F7030194A5752D434458FA60FA69F98D" box="[1299,1432,1535,1559]" pageId="0" pageNumber="439" value="2013-07-17">
<collectingDate id="E747F87CA5752D434458FA60FA69F98D" box="[1299,1432,1535,1559]" pageId="0" pageNumber="439" value="2013-07-17">17.VII.2013</collectingDate>
</date>
, leg.
<collectorName id="2E484282A5752D43418EF9BCFEC8F9A0" box="[197,313,1570,1594]" pageId="0" pageNumber="439">P. Hudson.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="83022754A5752D43418CF9D9FB04F9C5" blockId="0.[151,1437,1358,2027]" box="[199,1269,1606,1631]" pageId="0" pageNumber="439">
<emphasis id="B1C9FB46A5752D43418CF9D9FE67F9C4" box="[199,406,1606,1631]" inLineHeading="true" pageId="0" pageNumber="439" reason="1">
<emphasis id="B1C9FB46A5752D43418CF9D9FEB1F9C4" bold="true" box="[199,320,1606,1630]" italics="true" pageId="0" pageNumber="439">Adult male</emphasis>
(n = 2).
</emphasis>
Total length
<quantity id="44458AB1A5752D434368F9D8FD89F9C4" box="[547,632,1607,1631]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="0" pageNumber="439" unit="mm" value="4.7">4.7 mm</quantity>
. Wing length
<quantity id="44458AB1A5752D434240F9D9FC5BF9C4" box="[779,938,1606,1630]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.44" metricValueMin="2.56" pageId="0" pageNumber="439" unit="mm" value="3.0" valueMax="3.44" valueMin="2.56">2.56–3.44 mm</quantity>
. Total length/wing length 1.37.
</paragraph>
<paragraph id="83022754A5752D43418CF9F5FD38F93C" blockId="0.[151,1437,1358,2027]" pageId="0" pageNumber="439">Colouration. Head, thorax, legs and hypopygium dark brown; antenna brown; palpomeres light brown; abdomen light brown to brown; wing greyish, veins brownish.</paragraph>
<paragraph id="83022754A5752D43418CF92DFC7AF85D" blockId="0.[151,1437,1358,2027]" pageId="0" pageNumber="439">
Head. Eyes hairy, i.e., length of eye microtrichia about 1.5 or more times the height of ommatidial lenses and visible along lateral eye margin when head is viewed from front (
<bibRefCitation id="E72C5AA5A5752D43427EF948FBD4F974" author="Hansen, D. C. &amp; Cook, E. F." box="[821,1061,1750,1775]" pageId="0" pageNumber="439" pagination="1 - 203" refId="ref2327" refString="Hansen, D. C. &amp; Cook, E. F. (1976) The systematics and morphology of the Nearctic species of Diamesa Meigen, 1835 (Diptera, Chironomidae). Memoirs of the American Entomological Society, 30, 1 - 203." type="journal article" year="1976">Hansen &amp; Cook 1976</bibRefCitation>
). Frontal tubercles 16–44 μm long, covered with microtrichia. Temporal setae including about 13–15 verticals, 8–12 preoculars, 5 postorbitals. Clypeus with 5–6 setae. Antenna with 8 flagellomeres and reduced plume of setae; flagellomeres 1–7 with 2–3 setae, 24–40 μm long; terminal flagellomere in basal part with 4–5 setae, 60–72 μm long and with 2 subapical setae, 16–20 μm long. Flagellomeres 1–8 length (μm): 88–104, 36–40, 40, 32, 32, 26–32, 32–44, 164–248; AR 0.41–0.78. Antennal length/palp length 1.09–1.21. Palpomeres lengths (in μm): 32–40; 56–64; 108; 84–100; 136–156. Palpomere
<quantity id="44458AB1A5752D434525F814FB68F838" box="[1134,1177,1930,1955]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="0" pageNumber="439" unit="in" value="3.0">3 in</quantity>
distal part with sensilla capitata with diameter 20 μm. Palpomeres 1–5 length/head width 0.92.
</paragraph>
<paragraph id="83022754A5752D43418CF84DFB01F871" blockId="0.[151,1437,1358,2027]" box="[199,1264,2002,2027]" pageId="0" pageNumber="439">Thorax. Antepronotum with 9–11 ventrolateral setae. Dorsocentrals 5–7, prealars 7, scutellars 9–11.</paragraph>
<paragraph id="83022754A5742D42418CFF08FB18FF49" blockId="1.[151,1437,151,284]" pageId="1" pageNumber="440">
Wing.
<collectionCode id="E5ACBF91A5742D42405BFF07FED3FF35" box="[272,290,152,175]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="440">R</collectionCode>
and
<collectionCode id="E5ACBF91A5742D424013FF07FE9BFF35" box="[344,362,152,175]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="440">R</collectionCode>
<subScript id="1F392511A5742D424021FF37FE83FF2C" attach="left" box="[362,370,168,182]" fontSize="6" pageId="1" pageNumber="440">1</subScript>
with 25–34 setae;
<collectionCode id="E5ACBF91A5742D42430AFF07FDA2FF35" box="[577,595,152,175]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="440">R</collectionCode>
<subScript id="1F392511A5742D424319FF37FD9BFF2C" attach="left" box="[594,618,168,182]" fontSize="6" pageId="1" pageNumber="440">4+5</subScript>
with 18–19 setae. Costa extension 52–60 μm long.
<collectionCode id="E5ACBF91A5742D4245D4FF07FB39FF35" box="[1183,1224,152,175]" country="Canada" name="McGill University, Redpath Museum" pageId="1" pageNumber="440" type="Museum">RM</collectionCode>
length/MCu length 2.8–3.5. Anal lobe slightly reduced, angularly. Squama with 14–17 setae, 52–68 μm long. VR 0.96–1.0.
</paragraph>
<paragraph id="83022754A5742D42418CFF7FFBADFE86" blockId="1.[151,1437,151,284]" pageId="1" pageNumber="440">
Legs. Spur of fore tibia 40–44 μm long; spurs of mid tibia 44 μm long; of hind tibia 74–80 μm and 44–48 μm long. Hind tibial comb with 15–16 setae. Lengths and proportions of leg segments as in
<tableCitation id="CE3F12EFA5742D42454DFE9CFBA4FE86" box="[1030,1109,259,284]" captionStart="TABLE 1" captionStartId="1.[151,239,331,355]" captionTargetBox="[166,1402,372,522]" captionText="TABLE 1. Lengths (in µm) and proportions of leg segments of Diamesa leoniella Hansen, male (n=2)" httpUri="http://table.plazi.org/id/D7C277DCA5742D4241DCFED4FB2DFEFE" pageId="1" pageNumber="440" tableUuid="D7C277DCA5742D4241DCFED4FB2DFEFE">Table 1</tableCitation>
.
</paragraph>
<caption id="D7C277DCA5742D4241DCFED4FB2DFEFE" ID-Table-UUID="D7C277DCA5742D4241DCFED4FB2DFEFE" box="[151,1244,331,356]" httpUri="http://table.plazi.org/id/D7C277DCA5742D4241DCFED4FB2DFEFE" pageId="1" pageNumber="440" startId="1.[151,239,331,355]" targetBox="[166,1402,372,522]" targetIsTable="true" targetPageId="1">
<paragraph id="83022754A5742D4241DCFED4FB2DFEFE" blockId="1.[151,1244,331,356]" box="[151,1244,331,356]" pageId="1" pageNumber="440">
<emphasis id="B1C9FB46A5742D4241DCFED4FEFBFEF9" bold="true" box="[151,266,331,355]" pageId="1" pageNumber="440">TABLE 1.</emphasis>
Lengths (in µm) and proportions of leg segments of
<taxonomicName id="44BD5CD7A5742D424274FED3FBADFEF9" authority="Hansen" authorityName="Hansen" authorityYear="1976" box="[831,1116,331,355]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="leoniella">
<emphasis id="B1C9FB46A5742D424274FED3FBF3FEF9" box="[831,1026,331,355]" italics="true" pageId="1" pageNumber="440">Diamesa leoniella</emphasis>
Hansen
</taxonomicName>
, male (n=2)
</paragraph>
</caption>
<paragraph id="83022754A5742D424071FEEBFA8BFD9F" pageId="1" pageNumber="440">
<table id="F1BDD5F4A574D2BC41EDFEEBFA8BFD90" box="[166,1402,372,522]" gridcols="8" gridrows="4" pageId="1" pageNumber="440" rowsContinueIn="1.[166,1329,606,757]">
<tr id="3D8D2516A574D2BC41EDFEEBFA8BFE09" box="[166,1402,372,403]" gridrow="0" pageId="1" pageNumber="440" rowspan-0="1">
<th id="7E5C4C6AA574D2BC4040FEEBFE8CFE09" box="[267,381,372,403]" gridcol="1" gridrow="0" pageId="1" pageNumber="440">fe</th>
<th id="7E5C4C6AA574D2BC4080FEEBFDCFFE09" box="[459,574,372,403]" gridcol="2" gridrow="0" pageId="1" pageNumber="440">ti</th>
<th id="7E5C4C6AA574D2BC43C7FEEBFD0FFE09" box="[652,766,372,403]" gridcol="3" gridrow="0" pageId="1" pageNumber="440">ta 1</th>
<th id="7E5C4C6AA574D2BC4203FEEBFC50FE09" box="[840,929,372,403]" gridcol="4" gridrow="0" pageId="1" pageNumber="440">ta 2</th>
<th id="7E5C4C6AA574D2BC42ADFEEBFBCEFE09" box="[998,1087,372,403]" gridcol="5" gridrow="0" pageId="1" pageNumber="440">ta 3</th>
<th id="7E5C4C6AA574D2BC45C8FEEBFB2DFE09" box="[1155,1244,372,403]" gridcol="6" gridrow="0" pageId="1" pageNumber="440">ta 4</th>
<th id="7E5C4C6AA574D2BC446AFEEBFA8BFE09" box="[1313,1402,372,403]" gridcol="7" gridrow="0" pageId="1" pageNumber="440">ta 5</th>
</tr>
<tr id="3D8D2516A574D2BC41EDFE03FA8BFE20" box="[166,1402,412,442]" gridrow="1" pageId="1" pageNumber="440">
<th id="7E5C4C6AA574D2BC41EDFE03FF4AFE20" box="[166,187,412,442]" gridcol="0" gridrow="1" pageId="1" pageNumber="440">P 1</th>
<td id="7E5C4C6AA574D2BC4040FE03FE8CFE20" box="[267,381,412,442]" gridcol="1" gridrow="1" pageId="1" pageNumber="440">1607–2280</td>
<td id="7E5C4C6AA574D2BC4080FE03FDCFFE20" box="[459,574,412,442]" gridcol="2" gridrow="1" pageId="1" pageNumber="440">1591–2120</td>
<td id="7E5C4C6AA574D2BC43C7FE03FD0FFE20" box="[652,766,412,442]" gridcol="3" gridrow="1" pageId="1" pageNumber="440">1082–1400</td>
<td id="7E5C4C6AA574D2BC4203FE03FC50FE20" box="[840,929,412,442]" gridcol="4" gridrow="1" pageId="1" pageNumber="440">541–720</td>
<td id="7E5C4C6AA574D2BC42ADFE03FBCEFE20" box="[998,1087,412,442]" gridcol="5" gridrow="1" pageId="1" pageNumber="440">295–440</td>
<td id="7E5C4C6AA574D2BC45C8FE03FB2DFE20" box="[1155,1244,412,442]" gridcol="6" gridrow="1" pageId="1" pageNumber="440">107–140</td>
<td id="7E5C4C6AA574D2BC446AFE03FA8BFE20" box="[1313,1402,412,442]" gridcol="7" gridrow="1" pageId="1" pageNumber="440">148–180</td>
</tr>
<tr id="3D8D2516A574D2BC41EDFE5BFA8BFE78" box="[166,1402,452,482]" gridrow="2" pageId="1" pageNumber="440">
<th id="7E5C4C6AA574D2BC41EDFE5BFF4AFE78" box="[166,187,452,482]" gridcol="0" gridrow="2" pageId="1" pageNumber="440">P2</th>
<td id="7E5C4C6AA574D2BC4040FE5BFE8CFE78" box="[267,381,452,482]" gridcol="1" gridrow="2" pageId="1" pageNumber="440">1706–2160</td>
<td id="7E5C4C6AA574D2BC4080FE5BFDCFFE78" box="[459,574,452,482]" gridcol="2" gridrow="2" pageId="1" pageNumber="440">1394–1760</td>
<td id="7E5C4C6AA574D2BC43C7FE5BFD0FFE78" box="[652,766,452,482]" gridcol="3" gridrow="2" pageId="1" pageNumber="440">763–960</td>
<td id="7E5C4C6AA574D2BC4203FE5BFC50FE78" box="[840,929,452,482]" gridcol="4" gridrow="2" pageId="1" pageNumber="440">377–440</td>
<td id="7E5C4C6AA574D2BC42ADFE5BFBCEFE78" box="[998,1087,452,482]" gridcol="5" gridrow="2" pageId="1" pageNumber="440">230–360</td>
<td id="7E5C4C6AA574D2BC45C8FE5BFB2DFE78" box="[1155,1244,452,482]" gridcol="6" gridrow="2" pageId="1" pageNumber="440">98–140</td>
<td id="7E5C4C6AA574D2BC446AFE5BFA8BFE78" box="[1313,1402,452,482]" gridcol="7" gridrow="2" pageId="1" pageNumber="440">148–180</td>
</tr>
<tr id="3D8D2516A574D2BC41EDFE73FA8BFD90" box="[166,1402,492,522]" gridrow="3" pageId="1" pageNumber="440">
<th id="7E5C4C6AA574D2BC41EDFE73FF4AFD90" box="[166,187,492,522]" gridcol="0" gridrow="3" pageId="1" pageNumber="440">P3</th>
<td id="7E5C4C6AA574D2BC4040FE73FE8CFD90" box="[267,381,492,522]" gridcol="1" gridrow="3" pageId="1" pageNumber="440">1738–2400</td>
<td id="7E5C4C6AA574D2BC4080FE73FDCFFD90" box="[459,574,492,522]" gridcol="2" gridrow="3" pageId="1" pageNumber="440">1624–2160</td>
<td id="7E5C4C6AA574D2BC43C7FE73FD0FFD90" box="[652,766,492,522]" gridcol="3" gridrow="3" pageId="1" pageNumber="440">1033–1440</td>
<td id="7E5C4C6AA574D2BC4203FE73FC50FD90" box="[840,929,492,522]" gridcol="4" gridrow="3" pageId="1" pageNumber="440">607–800</td>
<td id="7E5C4C6AA574D2BC42ADFE73FBCEFD90" box="[998,1087,492,522]" gridcol="5" gridrow="3" pageId="1" pageNumber="440">312–400</td>
<td id="7E5C4C6AA574D2BC45C8FE73FB2DFD90" box="[1155,1244,492,522]" gridcol="6" gridrow="3" pageId="1" pageNumber="440">98–160</td>
<td id="7E5C4C6AA574D2BC446AFE73FA8BFD90" box="[1313,1402,492,522]" gridcol="7" gridrow="3" pageId="1" pageNumber="440">148–200</td>
</tr>
</table>
</paragraph>
<paragraph id="83022754A5742D4241DCFDA9FEE7FDD4" blockId="1.[151,278,566,590]" box="[151,278,566,590]" pageId="1" pageNumber="440">
<emphasis id="B1C9FB46A5742D4241DCFDA9FEE7FDD4" bold="true" box="[151,278,566,590]" pageId="1" pageNumber="440">
<tableNote id="7E5B26DAA5742D4241DCFDA9FEE7FDD4" box="[151,278,566,590]" pageId="1" pageNumber="440" targetBox="[166,1402,372,522]" targetPageId="1">Continued.</tableNote>
</emphasis>
</paragraph>
<paragraph id="83022754A5742D4240DCFDC0FAC0FD75" pageId="1" pageNumber="440">
<table id="F1BDD5F4A574D2BC41EDFDC1FAC0FD6F" box="[166,1329,606,757]" gridcols="5" gridrows="4" pageId="1" pageNumber="440" rowsContinueFrom="1.[166,1402,372,522]">
<tr id="3D8D2516A574D2BC41EDFDC1FAC0FDEC" box="[166,1329,606,630]" gridrow="0" pageId="1" pageNumber="440" rowspan-0="1">
<th id="7E5C4C6AA574D2BC403FFDC1FE2BFDEC" box="[372,474,606,630]" gridcol="1" gridrow="0" pageId="1" pageNumber="440">LR</th>
<th id="7E5C4C6AA574D2BC43DDFDC1FD0AFDEC" box="[662,763,606,630]" gridcol="2" gridrow="0" pageId="1" pageNumber="440">BV</th>
<th id="7E5C4C6AA574D2BC42FCFDC1FBECFDEC" box="[951,1053,606,630]" gridcol="3" gridrow="0" pageId="1" pageNumber="440">SV</th>
<th id="7E5C4C6AA574D2BC45AEFDC1FAC0FDEC" box="[1253,1329,606,630]" gridcol="4" gridrow="0" pageId="1" pageNumber="440">BR</th>
</tr>
<tr id="3D8D2516A574D2BC41EDFD19FAC0FD3E" box="[166,1329,646,676]" gridrow="1" pageId="1" pageNumber="440">
<th id="7E5C4C6AA574D2BC41EDFD19FF4AFD3E" box="[166,187,646,676]" gridcol="0" gridrow="1" pageId="1" pageNumber="440">P1</th>
<td id="7E5C4C6AA574D2BC403FFD19FE2BFD3E" box="[372,474,646,676]" gridcol="1" gridrow="1" pageId="1" pageNumber="440">0.66–0.68</td>
<td id="7E5C4C6AA574D2BC43DDFD19FD0AFD3E" box="[662,763,646,676]" gridcol="2" gridrow="1" pageId="1" pageNumber="440">3.92</td>
<td id="7E5C4C6AA574D2BC42FCFD19FBECFD3E" box="[951,1053,646,676]" gridcol="3" gridrow="1" pageId="1" pageNumber="440">2.96–3.14</td>
<td id="7E5C4C6AA574D2BC45AEFD19FAC0FD3E" box="[1253,1329,646,676]" gridcol="4" gridrow="1" pageId="1" pageNumber="440">0.8–1.2</td>
</tr>
<tr id="3D8D2516A574D2BC41EDFD31FAC0FD56" box="[166,1329,686,716]" gridrow="2" pageId="1" pageNumber="440">
<th id="7E5C4C6AA574D2BC41EDFD31FF4AFD56" box="[166,187,686,716]" gridcol="0" gridrow="2" pageId="1" pageNumber="440">P2</th>
<td id="7E5C4C6AA574D2BC403FFD31FE2BFD56" box="[372,474,686,716]" gridcol="1" gridrow="2" pageId="1" pageNumber="440">0.55</td>
<td id="7E5C4C6AA574D2BC43DDFD31FD0AFD56" box="[662,763,686,716]" gridcol="2" gridrow="2" pageId="1" pageNumber="440">4.36–4.53</td>
<td id="7E5C4C6AA574D2BC42FCFD31FBECFD56" box="[951,1053,686,716]" gridcol="3" gridrow="2" pageId="1" pageNumber="440">4.06–4.08</td>
<td id="7E5C4C6AA574D2BC45AEFD31FAC0FD56" box="[1253,1329,686,716]" gridcol="4" gridrow="2" pageId="1" pageNumber="440">0.8–1.1</td>
</tr>
<tr id="3D8D2516A574D2BC41EDFD48FAC0FD6F" box="[166,1329,727,757]" gridrow="3" pageId="1" pageNumber="440">
<th id="7E5C4C6AA574D2BC41EDFD48FF4AFD6F" box="[166,187,727,757]" gridcol="0" gridrow="3" pageId="1" pageNumber="440">P 3</th>
<td id="7E5C4C6AA574D2BC403FFD48FE2BFD6F" box="[372,474,727,757]" gridcol="1" gridrow="3" pageId="1" pageNumber="440">0.64–0.67</td>
<td id="7E5C4C6AA574D2BC43DDFD48FD0AFD6F" box="[662,763,727,757]" gridcol="2" gridrow="3" pageId="1" pageNumber="440">3.77–3.85</td>
<td id="7E5C4C6AA574D2BC42FCFD48FBECFD6F" box="[951,1053,727,757]" gridcol="3" gridrow="3" pageId="1" pageNumber="440">3.17–3.25</td>
<td id="7E5C4C6AA574D2BC45AEFD48FAC0FD6F" box="[1253,1329,727,757]" gridcol="4" gridrow="3" pageId="1" pageNumber="440">0.8–1.2</td>
</tr>
</table>
</paragraph>
<paragraph id="83022754A5742D42418CFCBFFC9BFC76" blockId="1.[151,1437,800,1220]" pageId="1" pageNumber="440">
Hypopygium (
<figureCitation id="1B863BD1A5742D424028FCBFFE3BFCA2" box="[355,458,800,824]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1950,1974]" captionTargetBox="[151,1435,596,1927]" captionTargetId="figure-144@2.[151,1435,596,1927]" captionTargetPageId="2" captionText="FIGURES 1–4. Males of Diamesa leoniella Hansen (1–2) and D. japonica Tokunaga (3–4). 1, 4, central part of hypopygium without tergite IX; 2–3, hypopygium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4955879" httpUri="https://zenodo.org/record/4955879/files/figure.png" pageId="1" pageNumber="440">Figs. 1–2</figureCitation>
). Tergite IX with 11–13 setae (from one side), 16–24 μm long and wedge-shaped anal point 100–140 μm long, covered in basal half with numerous microtrichia forward and laterally directed. Laterosternite IX with 10–11 short setae. Transverse sternapodeme triangular 128 μm long. Gonocoxite 400–496 μm long. Inferior volsella finger-like, 40–48 μm long and 24 μm width, covered with short setae and microtrichia. Superior volsella collar-like, covered with microtrichia and short setae. Gonostylus slightly curved, 212–246 μm long, with numerous, proximally directed strong setae 48–64 μm long; megaseta 6–8 μm long. HR 1.65–1.93.
</paragraph>
<paragraph id="83022754A5742D42418CFC67FA97FBE6" blockId="1.[151,1437,800,1220]" pageId="1" pageNumber="440">
<emphasis id="B1C9FB46A5742D42418CFC67FE31FB8A" bold="true" box="[199,448,1016,1040]" inLineHeading="true" pageId="1" pageNumber="440" reason="1">Diagnostic characters.</emphasis>
Male of
<taxonomicName id="44BD5CD7A5742D42436BFC66FD6DFB8A" authorityName="Hansen" authorityYear="1976" box="[544,668,1016,1040]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="leoniella">
<emphasis id="B1C9FB46A5742D42436BFC66FD6DFB8A" box="[544,668,1016,1040]" italics="true" pageId="1" pageNumber="440">D. leoniella</emphasis>
</taxonomicName>
is very closely related to
<taxonomicName id="44BD5CD7A5742D4242E1FC66FBD7FB8A" authorityName="Tokunaga" authorityYear="1936" box="[938,1062,1017,1040]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B1C9FB46A5742D4242E1FC66FBD7FB8A" box="[938,1062,1017,1040]" italics="true" pageId="1" pageNumber="440">D. japonica</emphasis>
</taxonomicName>
and distinguished from later by the following main features: 1—antennal flagellomeres 1–7 with 2–3 short setae, 2—inferior volsella finger-like, covered with short setae and microtrichia, 3—superior volsella collar-like, covered with microtrichia and short setae. Male of
<emphasis id="B1C9FB46A5742D424452FBDEFA6CFBC2" box="[1305,1437,1089,1112]" italics="true" pageId="1" pageNumber="440">
<taxonomicName id="44BD5CD7A5742D424452FBDEFA67FBC2" authorityName="Tokunaga" authorityYear="1936" box="[1305,1430,1089,1112]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="japonica">D. japonica</taxonomicName>
:
</emphasis>
1–without setae on flagellomeres 4–7, 2—inferior volsella absent, 3—superior volsella in form of an angular tubercle.
</paragraph>
</subSubSection>
<subSubSection id="CBA774DFA5742D42418CFB17FD35FB5E" pageId="1" pageNumber="440" type="distribution">
<paragraph id="83022754A5742D42418CFB17FD35FB5E" blockId="1.[151,1437,800,1220]" pageId="1" pageNumber="440">
<emphasis id="B1C9FB46A5742D42418CFB17FEABFB3A" bold="true" box="[199,346,1160,1184]" inLineHeading="true" pageId="1" pageNumber="440" reason="1">Distribution.</emphasis>
Nearctic species. Known from
<collectingCountry id="FBAA67C4A5742D4243ECFB17FD16FB3A" box="[679,743,1160,1184]" name="United States of America" pageId="1" pageNumber="440">U.S.A.</collectingCountry>
—
<collectingRegion id="4179E9B6A5742D42424FFB17FC8BFB3A" box="[772,890,1160,1184]" country="United States of America" name="California" pageId="1" pageNumber="440">California</collectingRegion>
,
<collectingRegion id="4179E9B6A5742D4242CFFB16FC17FB3A" box="[900,998,1161,1184]" country="United States of America" name="Montana" pageId="1" pageNumber="440">Montana</collectingRegion>
,
<collectingRegion id="4179E9B6A5742D4242A4FB17FBD7FB3A" box="[1007,1062,1160,1184]" country="United States of America" name="Utah" pageId="1" pageNumber="440">Utah</collectingRegion>
,
<collectingRegion id="4179E9B6A5742D424564FB17FB40FB3A" box="[1071,1201,1160,1184]" country="United States of America" name="Washington" pageId="1" pageNumber="440">Washington</collectingRegion>
,
<collectingRegion id="4179E9B6A5742D4245F1FB17FAD2FB3A" box="[1210,1315,1160,1184]" country="United States of America" name="Wyoming" pageId="1" pageNumber="440">Wyoming</collectingRegion>
(
<bibRefCitation id="E72C5AA5A5742D42447BFB17FEFEFB5E" author="Hansen, D. C. &amp; Cook, E. F." pageId="1" pageNumber="440" pagination="1 - 203" refId="ref2327" refString="Hansen, D. C. &amp; Cook, E. F. (1976) The systematics and morphology of the Nearctic species of Diamesa Meigen, 1835 (Diptera, Chironomidae). Memoirs of the American Entomological Society, 30, 1 - 203." type="journal article" year="1976">Hansen &amp; Cook 1976</bibRefCitation>
) and
<collectingRegion id="4179E9B6A5742D424003FB33FE63FB5E" box="[328,402,1196,1220]" country="United States of America" name="Alaska" pageId="1" pageNumber="440">Alaska</collectingRegion>
(recorded for the first time).
</paragraph>
</subSubSection>
</treatment>
</document>
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