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<document id="C696916FC673C76DDC469796EF26E7D9" ID-CLB-Dataset="5710" ID-DOI="10.11646/zootaxa.4985.3.11" ID-GBIF-Dataset="7dc8160d-98f9-4c8a-8382-994e97b7d71f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4955877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623756031967" checkinUser="plazi" docAuthor="Makarchenko Usgs, Eugenyi A." docDate="2021" docId="0B149642A5742D4041DCFA89FDF5FE12" docLanguage="en" docName="zootaxa.4985.3.11.pdf" docOrigin="Zootaxa 4985 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diamesa japonica Tokunaga 1936" docType="treatment" docVersion="8" lastPageNumber="442" masterDocId="F72DEE3AA5752D43414BFF9FFFF1FF9A" masterDocTitle="Confirmation of a species status of Diamesa leoniella Hansen, 1976, stat. resurr (Diptera, Chironomidae) from North America" masterLastPageNumber="442" masterPageNumber="439" pageNumber="440" updateTime="1699016569100" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="3F37BFF30CD7AF47040A5BAD0F5045FB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D96725034C93C76A660A02EB8372C3AE">Confirmation of a species status of Diamesa leoniella Hansen, 1976, stat. resurr (Diptera, Chironomidae) from North America</mods:title>
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<mods:namePart id="6EAA5CA253BDD2AA7DCE2F3C6F35FA5D">Makarchenko Usgs, Eugenyi A.</mods:namePart>
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<mods:typeOfResource id="69DD8C023326379F5240E356E91F53ED">text</mods:typeOfResource>
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<mods:date id="9FC6680A556087135AF3FDDAB8494973">2021</mods:date>
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<mods:number id="96A5446C1DB8DF203FB1C6C27322FAC2">2021-06-15</mods:number>
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<mods:number id="E5F0C0AD9F9A62B3CB6DADCB472CA22B">4985</mods:number>
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<mods:identifier id="A42EECB8531B0CC6CEFAD4CD68C95965" type="DOI">10.11646/zootaxa.4985.3.11</mods:identifier>
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<treatment id="0B149642A5742D4041DCFA89FDF5FE12" ID-DOI="http://doi.org/10.5281/zenodo.4955883" ID-GBIF-Taxon="183325885" ID-Zenodo-Dep="4955883" LSID="urn:lsid:plazi:treatment:0B149642A5742D4041DCFA89FDF5FE12" httpUri="http://treatment.plazi.org/id/0B149642A5742D4041DCFA89FDF5FE12" lastPageId="3" lastPageNumber="442" pageId="1" pageNumber="440">
<subSubSection id="CBA774DFA5742D4241DCFA89FDCFFAAA" box="[151,574,1302,1328]" pageId="1" pageNumber="440" type="nomenclature">
<paragraph id="83022754A5742D4241DCFA89FDCFFAAA" blockId="1.[151,1437,1302,2038]" box="[151,574,1302,1328]" pageId="1" pageNumber="440">
<heading id="D84A9038A5742D4241DCFA89FDCFFAAA" bold="true" box="[151,574,1302,1328]" fontSize="11" level="1" pageId="1" pageNumber="440" reason="1">
<taxonomicName id="44BD5CD7A5742D4241DCFA89FDCFFAAA" ID-CoL="6CQN4" authority="Tokunaga, 1936" authorityName="Tokunaga" authorityYear="1936" box="[151,574,1302,1328]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B1C9FB46A5742D4241DCFA89FDCFFAAA" bold="true" box="[151,574,1302,1328]" pageId="1" pageNumber="440">
<emphasis id="B1C9FB46A5742D4241DCFA89FE81FAAA" bold="true" box="[151,368,1302,1328]" italics="true" pageId="1" pageNumber="440">Diamesa japonica</emphasis>
<bibRefCitation id="E72C5AA5A5742D424033FA89FDCFFAAA" author="Tokunaga, M." box="[376,574,1302,1328]" pageId="1" pageNumber="440" pagination="525 - 552" refId="ref2701" refString="Tokunaga, M. (1936) Chironomidae from Japan (Diptera). VI. Diamesinae. Philippine Journal of Science, 59, 525 - 552." type="journal article" year="1936">Tokunaga, 1936</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CBA774DFA5742D4241DCFAA3FEFDFACE" box="[151,268,1340,1364]" pageId="1" pageNumber="440" type="description">
<paragraph id="83022754A5742D4241DCFAA3FEFDFACE" blockId="1.[151,1437,1302,2038]" box="[151,268,1340,1364]" pageId="1" pageNumber="440">
(
<figureCitation id="1B863BD1A5742D4241D4FAA3FEF5FACE" box="[159,260,1340,1364]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1950,1974]" captionTargetBox="[151,1435,596,1927]" captionTargetId="figure-144@2.[151,1435,596,1927]" captionTargetPageId="2" captionText="FIGURES 1–4. Males of Diamesa leoniella Hansen (1–2) and D. japonica Tokunaga (3–4). 1, 4, central part of hypopygium without tergite IX; 2–3, hypopygium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4955879" httpUri="https://zenodo.org/record/4955879/files/figure.png" pageId="1" pageNumber="440">Figs. 3–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CBA774DFA5742D4241DCFAC4FC2CFA08" pageId="1" pageNumber="440" type="reference_group">
<paragraph id="83022754A5742D4241DCFAC4FAB5FAE9" blockId="1.[151,1437,1302,2038]" box="[151,1348,1371,1395]" pageId="1" pageNumber="440">
<treatmentCitationGroup id="A3AD007AA5742D4241DCFAC4FAB5FAE9" box="[151,1348,1371,1395]" pageId="1" pageNumber="440">
<treatmentCitation id="021C0145A5742D4241DCFAC4FDDEFAE9" author="Tokunaga, M." box="[151,559,1371,1395]" page="542" pageId="1" pageNumber="440" year="1936">
<taxonomicName id="44BD5CD7A5742D4241DCFAC4FDDEFAE9" ID-CoL="6CQN4" authority="Tokunaga, 1936: 542" authorityName="Tokunaga" authorityPageNumber="542" authorityYear="1936" box="[151,559,1371,1395]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B1C9FB46A5742D4241DCFAC4FEA0FAE8" box="[151,337,1371,1394]" italics="true" pageId="1" pageNumber="440">Diamesa japonica</emphasis>
<bibRefCitation id="E72C5AA5A5742D42401DFAC4FDDEFAE9" author="Tokunaga, M." box="[342,559,1371,1395]" pageId="1" pageNumber="440" pagination="525 - 552" refId="ref2701" refString="Tokunaga, M. (1936) Chironomidae from Japan (Diptera). VI. Diamesinae. Philippine Journal of Science, 59, 525 - 552." type="journal article" year="1936">Tokunaga, 1936: 542</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="021C0145A5742D424371FAC4FCC3FAE9" author="Makarchenko, E. A." box="[570,818,1371,1395]" page="110" pageId="1" pageNumber="440" year="1981">
<bibRefCitation id="E72C5AA5A5742D424371FAC4FCC3FAE9" author="Makarchenko, E. A." box="[570,818,1371,1395]" pageId="1" pageNumber="440" pagination="89 - 113" refId="ref2375" refString="Makarchenko, E. A. (1981) Taxonomy and distribution of some chironomids of Diamesinae (Diptera, Chironomidae) from the Far East of the USSR. In: Invertebrates in the ecosystems of Far Eastern salmon rivers. DVNC AN SSSR Press, Vladivostok, pp. 89 - 113. [in Russian]" type="book chapter" year="1981">Makarchenko 1981: 110</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="021C0145A5742D424276FAC4FC68FAE9" author="Makarchenko, E. A." box="[829,921,1371,1395]" page="76" pageId="1" pageNumber="440" year="1985">
<bibRefCitation id="E72C5AA5A5742D424276FAC4FC68FAE9" author="Makarchenko, E. A." box="[829,921,1371,1395]" pageId="1" pageNumber="440" refId="ref2434" refString="Makarchenko, E. A. (1985) Chironomids of the Soviet Far East. Subfamilies Podonominae, Diamesinae and Prodiamesinae (Diptera, Chironomidae). DVNC AN SSSR Press, Vladivostok, 208 pp. [in Russian]" type="book" year="1985">1985: 76</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="021C0145A5742D4242E8FAC4FBFDFAE9" author="Makarchenko, E. A." box="[931,1036,1371,1395]" page="260" pageId="1" pageNumber="440" year="2006">
<bibRefCitation id="E72C5AA5A5742D4242E8FAC4FBFDFAE9" author="Makarchenko, E. A." box="[931,1036,1371,1395]" pageId="1" pageNumber="440" pagination="607 - 621" refId="ref2476" refString="Makarchenko, E. A. (2006) 3. Subfamily Diamesinae. In: Lelej, A. (Ed.), Key to the insects of Russian Far East. Vol. 6. Diptera and Siphonaptera. Pt 4. Dal'nauka, Vladivostok, pp. 253 - 276 + 468 - 480 + 607 - 621. [in Russian]" type="book chapter" year="2006">2006: 260</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="021C0145A5742D42455EFAC4FAB1FAE9" author="Ashe, P. &amp; O'Connor, J. P." box="[1045,1344,1371,1395]" page="278" pageId="1" pageNumber="440" year="2009">
<bibRefCitation id="E72C5AA5A5742D42455EFAC4FAB1FAE9" author="Ashe, P. &amp; O'Connor, J. P." box="[1045,1344,1371,1395]" pageId="1" pageNumber="440" refId="ref2269" refString="Ashe, P. &amp; O'Connor, J. P. (2009) A World Catalogue of Chironomidae (Diptera). Part 1. Buchonomyiinae, Chilenomyiinae, Podonominae, Aphroteniinae, Tanypodinae, Usambaromyiinae, Diamesinae, Prodiamesinae and Telmatogetoninae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, 445 pp." type="book" year="2009">Ashe &amp; O’Connor 2009: 278</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="83022754A5742D4241DCFAE5FC2CFA08" blockId="1.[151,1437,1302,2038]" box="[151,989,1401,1426]" pageId="1" pageNumber="440">
<treatmentCitationGroup id="A3AD007AA5742D4241DCFAE5FC2CFA08" box="[151,989,1401,1426]" pageId="1" pageNumber="440">
<taxonomicName id="44BD5CD7A5742D4241DCFAE5FD54FA0B" ID-CoL="9HWDB" authority="Sasa et Okazava, 1992: 60" authorityName="Sasa et Okazava" authorityPageNumber="60" authorityYear="1992" box="[151,677,1401,1426]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="kurobemijikaia">
<emphasis id="B1C9FB46A5742D4241DCFAE5FE61FA0B" box="[151,400,1401,1425]" italics="true" pageId="1" pageNumber="440">Diamesa kurobemijikaia</emphasis>
Sasa et Okazava, 1992: 60
</taxonomicName>
;
<treatmentCitation id="021C0145A5742D4243E5FAE6FC28FA08" author="Ashe, P. &amp; O'Connor, J. P." box="[686,985,1401,1426]" page="278" pageId="1" pageNumber="440" year="2009">
<bibRefCitation id="E72C5AA5A5742D4243E5FAE6FC28FA08" author="Ashe, P. &amp; O'Connor, J. P." box="[686,985,1401,1426]" pageId="1" pageNumber="440" refId="ref2269" refString="Ashe, P. &amp; O'Connor, J. P. (2009) A World Catalogue of Chironomidae (Diptera). Part 1. Buchonomyiinae, Chilenomyiinae, Podonominae, Aphroteniinae, Tanypodinae, Usambaromyiinae, Diamesinae, Prodiamesinae and Telmatogetoninae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, 445 pp." type="book" year="2009">Ashe &amp; O’Connor 2009: 278</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CBA774DFA5742D4041DCFA02FBB8FEDA" lastPageId="3" lastPageNumber="442" pageId="1" pageNumber="440" type="materials_examined">
<paragraph id="83022754A5742D4241DCFA02FB88FA43" blockId="1.[151,1437,1302,2038]" pageId="1" pageNumber="440">
<emphasis id="B1C9FB46A5742D4241DCFA02FE82FA2F" bold="true" box="[151,371,1437,1461]" inLineHeading="true" pageId="1" pageNumber="440" reason="1">Material examined.</emphasis>
<materialsCitation id="33D52D09A5742D42403CFA01FE88FA43" ID-GBIF-Occurrence="3314008304" collectingDate="1978-04-24" collectorName="E. Makarchenko" country="Russia" county="Tiurino River" location="Kurile Islands" municipality="Sernovodsk Village" pageId="1" pageNumber="440" specimenCount="2" specimenCount-adult="2">
<specimenCount id="95BBECDDA5742D42403CFA01FE4FFA2F" box="[375,446,1437,1462]" pageId="1" pageNumber="440" type="adult">2 adult</specimenCount>
males,
<collectingCountry id="FBAA67C4A5742D424340FA02FDA7FA2F" box="[523,598,1437,1461]" name="Russia" pageId="1" pageNumber="440">Russia</collectingCountry>
:
<location id="8662718FA5742D424315FA02FD05FA2F" LSID="urn:lsid:plazi:treatment:0B149642A5742D4041DCFA89FDF5FE12:8662718FA5742D424315FA02FD05FA2F" box="[606,756,1437,1461]" country="Russia" county="Tiurino River" municipality="Sernovodsk Village" name="Kurile Islands" pageId="1" pageNumber="440">Kurile Islands</location>
,
<location id="8662718FA5742D4243B7FA02FC57FA2F" LSID="urn:lsid:plazi:treatment:0B149642A5742D4041DCFA89FDF5FE12:8662718FA5742D4243B7FA02FC57FA2F" box="[764,934,1437,1461]" country="Russia" county="Tiurino River" municipality="Sernovodsk Village" name="Kunashir Island" pageId="1" pageNumber="440">Kunashir Island</location>
,
<collectingMunicipality id="6366BD2EA5742D4242E5FA02FB71FA2F" box="[942,1152,1437,1461]" pageId="1" pageNumber="440">Sernovodsk Village</collectingMunicipality>
,
<collectingCounty id="6A635FD8A5742D4245CCFA02FAE6FA2F" box="[1159,1303,1437,1461]" pageId="1" pageNumber="440">Tiurino River</collectingCounty>
,
<date id="F7030194A5742D42446BFA01FA69FA2C" box="[1312,1432,1438,1462]" pageId="1" pageNumber="440" value="1978-04-24">
<collectingDate id="E747F87CA5742D42446BFA01FA69FA2C" box="[1312,1432,1438,1462]" pageId="1" pageNumber="440" value="1978-04-24">24.IV.1978</collectingDate>
</date>
, leg.
<collectorName id="2E484282A5742D42418EFA5DFE88FA43" box="[197,377,1473,1497]" pageId="1" pageNumber="440">E. Makarchenko</collectorName>
</materialsCitation>
;
<materialsCitation id="33D52D09A5742D4240C8FA5DFB88FA43" ID-GBIF-Occurrence="3314008301" box="[387,1145,1473,1498]" collectingDate="1978-04-26" collectorName="E. Makarchenko." country="Russia" county="Tiurino River" location="Kurile Islands" municipality="Sernovodsk Village" pageId="1" pageNumber="440" specimenCount="23" specimenCount-adult="23">
<specimenCount id="95BBECDDA5742D4240C8FA5DFE2BFA43" box="[387,474,1473,1498]" pageId="1" pageNumber="440" type="adult">23 adult</specimenCount>
males, the same data except
<date id="F7030194A5742D42425BFA5EFC78FA43" box="[784,905,1473,1497]" pageId="1" pageNumber="440" value="1978-04-26">
<collectingDate id="E747F87CA5742D42425BFA5EFC78FA43" box="[784,905,1473,1497]" pageId="1" pageNumber="440" value="1978-04-26">26.IV.1978</collectingDate>
</date>
, leg.
<collectorName id="2E484282A5742D424289FA5DFB88FA43" box="[962,1145,1473,1497]" pageId="1" pageNumber="440">E. Makarchenko.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="83022754A5742D42418CFA7AFA9CFA64" blockId="1.[151,1437,1302,2038]" box="[199,1389,1509,1534]" pageId="1" pageNumber="440">
<emphasis id="B1C9FB46A5742D42418CFA7AFE55FA67" box="[199,420,1509,1534]" inLineHeading="true" pageId="1" pageNumber="440" reason="1">
<emphasis id="B1C9FB46A5742D42418CFA7AFEB1FA67" bold="true" box="[199,320,1509,1533]" italics="true" pageId="1" pageNumber="440">Adult male</emphasis>
(n = 12).
</emphasis>
Total length 3.1–4.0 mm. Wing length
<quantity id="44458AB1A5742D42420CFA7AFC17FA67" box="[839,998,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="2.965" metricValueMax="3.17" metricValueMin="2.76" pageId="1" pageNumber="440" unit="mm" value="2.965" valueMax="3.17" valueMin="2.76">2.76–3.17 mm</quantity>
. Total length/wing length 0.90–1.38.
</paragraph>
<paragraph id="83022754A5742D42418CF996FD8DF9DF" blockId="1.[151,1437,1302,2038]" pageId="1" pageNumber="440">Colouration. Head, thorax, legs and hypopygium dark brown; antenna and palpomeres brown; abdomen light brown to dark brown; wing greyish, veins brownish.</paragraph>
<paragraph id="83022754A5742D42418CF9CEFD73F8D8" blockId="1.[151,1437,1302,2038]" pageId="1" pageNumber="440">
Head. Eyes hairy. Temporal setae including 12–15 verticals, 6–12 preoculars, 4–5 postorbitals. Clypeus with 6–8 setae. Antenna with 8 flagellomeres and reduced plume of setae; number and length of these setae on 1–7 flagellomeres respectively: 2–3 (28–40 μm), 1–2 (28–36 μm), 1–2 (26–40 μm), 0, 0, 0, 0; terminal flagellomere in basal part with 5–6 setae, 60–80 μm long and with 2 subapical setae, 40–52 μm long. Flagellomeres 1–8 length (μm): 71–88, 34–38, 25–34, 25–30, 25–34, 25–34, 34–38, 193–218; AR 0.68–0.81. Antennal length/palp length 1.05–1.11. Palpomeres lengths (in μm): 36–40; 55–63; 88–105; 76–105; 105–147. Palpomere
<quantity id="44458AB1A5742D424245F899FCC6F887" box="[782,823,1797,1822]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="440" unit="in" value="3.0">3 in</quantity>
distal part with sensilla capitata with diameter 20–24 μm. Palpomeres 1–5 length/head width 0.80–0.84.
</paragraph>
<paragraph id="83022754A5742D42418CF8D2FAFDF8FC" blockId="1.[151,1437,1302,2038]" box="[199,1292,1869,1894]" pageId="1" pageNumber="440">Thorax. Antepronotum with 7–12 ventrolateral setae. Dorsocentrals 6–10, prealars 5, scutellars 11–24.</paragraph>
<paragraph id="83022754A5742D42418CF8EEFB3FF837" blockId="1.[151,1437,1302,2038]" pageId="1" pageNumber="440">
Wing.
<collectionCode id="E5ACBF91A5742D42405AF8EDFED2F813" box="[273,291,1906,1929]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="440">R</collectionCode>
and
<collectionCode id="E5ACBF91A5742D424010F8EDFE9CF813" box="[347,365,1906,1929]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="440">R</collectionCode>
<subScript id="1F392511A5742D424026F81DFE84F80A" attach="left" box="[365,373,1922,1936]" fontSize="6" pageId="1" pageNumber="440">1</subScript>
with 15–25 setae;
<collectionCode id="E5ACBF91A5742D424303F8EDFDABF813" box="[584,602,1906,1929]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="440">R</collectionCode>
<subScript id="1F392511A5742D424311F81EFD83F815" attach="left" box="[602,626,1921,1935]" fontSize="6" pageId="1" pageNumber="440">4+5</subScript>
with 8–12 setae. Costa extension
<emphasis id="B1C9FB46A5742D4242A7F8EDFBF4F813" box="[1004,1029,1906,1929]" italics="true" pageId="1" pageNumber="440">ca</emphasis>
48 μm long.
<collectionCode id="E5ACBF91A5742D4245D6F8EDFB37F813" box="[1181,1222,1906,1929]" country="Canada" name="McGill University, Redpath Museum" pageId="1" pageNumber="440" type="Museum">RM</collectionCode>
length/MCu length 2.0–3.6. Anal lobe well developed, rounded. Squama with 18–25 setae 28–52 μm long. VR 0.96–1.0.
</paragraph>
<paragraph id="83022754A5742D42418CF825FB69F86C" blockId="1.[151,1437,1302,2038]" pageId="1" pageNumber="440">
Legs. Spur of fore tibia 34–46 μm long; spurs of mid tibia 46–55 μm long; of hind tibia 50–71 μm and 34–50 μm long. Hind tibial comb with 16
<emphasis id="B1C9FB46A5742D4240ABF841FE1CF86F" box="[480,493,2014,2037]" italics="true" pageId="1" pageNumber="440">–</emphasis>
20 setae. Lengths and proportions of leg segments as in
<tableCitation id="CE3F12EFA5742D424508F842FB65F86C" box="[1091,1172,2013,2038]" captionStart="TABLE 2" captionStartId="2.[151,239,151,175]" captionTargetBox="[166,1394,192,342]" captionText="TABLE 2. Lengths (in µm) and proportions of leg segments of Diamesa japonica Tokunaga, male (n=12)" httpUri="http://table.plazi.org/id/D7C277DCA5772D4141DCFF08FAF3FF2A" pageId="1" pageNumber="440" tableUuid="D7C277DCA5772D4141DCFF08FAF3FF2A">Table 2</tableCitation>
.
</paragraph>
<caption id="D7C277DCA5772D4141DCFF08FAF3FF2A" ID-Table-UUID="D7C277DCA5772D4141DCFF08FAF3FF2A" box="[151,1282,151,176]" httpUri="http://table.plazi.org/id/D7C277DCA5772D4141DCFF08FAF3FF2A" pageId="2" pageNumber="441" startId="2.[151,239,151,175]" targetBox="[166,1394,192,342]" targetIsTable="true" targetPageId="2">
<paragraph id="83022754A5772D4141DCFF08FAF3FF2A" blockId="2.[151,1282,151,176]" box="[151,1282,151,176]" pageId="2" pageNumber="441">
<emphasis id="B1C9FB46A5772D4141DCFF08FEFBFF35" bold="true" box="[151,266,151,175]" pageId="2" pageNumber="441">TABLE 2.</emphasis>
Lengths (in µm) and proportions of leg segments of
<taxonomicName id="44BD5CD7A5772D414274FF07FB84FF35" authority="Tokunaga" authorityName="Tokunaga" authorityYear="1936" box="[831,1141,151,175]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B1C9FB46A5772D414274FF07FBF3FF35" box="[831,1026,152,175]" italics="true" pageId="2" pageNumber="441">Diamesa japonica</emphasis>
Tokunaga
</taxonomicName>
, male (n=12)
</paragraph>
</caption>
<paragraph id="83022754A5772D414066FF5FFA83FECA" pageId="2" pageNumber="441">
<table id="F1BDD5F4A577D2BC41EDFF5FFA83FECC" box="[166,1394,192,342]" gridcols="8" gridrows="4" pageId="2" pageNumber="441" rowsContinueIn="2.[166,1326,426,577]">
<tr id="3D8D2516A577D2BC41EDFF5FFA83FF45" box="[166,1394,192,223]" gridrow="0" pageId="2" pageNumber="441" rowspan-0="1">
<th id="7E5C4C6AA577D2BC41B5FF5FFE81FF45" box="[254,368,192,223]" gridcol="1" gridrow="0" pageId="2" pageNumber="441">fe</th>
<th id="7E5C4C6AA577D2BC40E0FF5FFDECFF45" box="[427,541,192,223]" gridcol="2" gridrow="0" pageId="2" pageNumber="441">ti</th>
<th id="7E5C4C6AA577D2BC4315FF5FFD35FF45" box="[606,708,192,223]" gridcol="3" gridrow="0" pageId="2" pageNumber="441">ta 1</th>
<th id="7E5C4C6AA577D2BC4259FF5FFC9AFF45" box="[786,875,192,223]" gridcol="4" gridrow="0" pageId="2" pageNumber="441">ta 2</th>
<th id="7E5C4C6AA577D2BC42F4FF5FFBE9FF45" box="[959,1048,192,223]" gridcol="5" gridrow="0" pageId="2" pageNumber="441">ta 3</th>
<th id="7E5C4C6AA577D2BC4527FF5FFB34FF45" box="[1132,1221,192,223]" gridcol="6" gridrow="0" pageId="2" pageNumber="441">ta 4</th>
<th id="7E5C4C6AA577D2BC4452FF5FFA83FF45" box="[1305,1394,192,223]" gridcol="7" gridrow="0" pageId="2" pageNumber="441">ta 5</th>
</tr>
<tr id="3D8D2516A577D2BC41EDFF77FA83FE9C" box="[166,1394,232,262]" gridrow="1" pageId="2" pageNumber="441">
<th id="7E5C4C6AA577D2BC41EDFF77FF4AFE9C" box="[166,187,232,262]" gridcol="0" gridrow="1" pageId="2" pageNumber="441">P1</th>
<td id="7E5C4C6AA577D2BC41B5FF77FE81FE9C" box="[254,368,232,262]" gridcol="1" gridrow="1" pageId="2" pageNumber="441">1496–1768</td>
<td id="7E5C4C6AA577D2BC40E0FF77FDECFE9C" box="[427,541,232,262]" gridcol="2" gridrow="1" pageId="2" pageNumber="441">1513–1716</td>
<td id="7E5C4C6AA577D2BC4315FF77FD35FE9C" box="[606,708,232,262]" gridcol="3" gridrow="1" pageId="2" pageNumber="441">918–1105</td>
<td id="7E5C4C6AA577D2BC4259FF77FC9AFE9C" box="[786,875,232,262]" gridcol="4" gridrow="1" pageId="2" pageNumber="441">425–527</td>
<td id="7E5C4C6AA577D2BC42F4FF77FBE9FE9C" box="[959,1048,232,262]" gridcol="5" gridrow="1" pageId="2" pageNumber="441">255–306</td>
<td id="7E5C4C6AA577D2BC4527FF77FB34FE9C" box="[1132,1221,232,262]" gridcol="6" gridrow="1" pageId="2" pageNumber="441">102–136</td>
<td id="7E5C4C6AA577D2BC4452FF77FA83FE9C" box="[1305,1394,232,262]" gridcol="7" gridrow="1" pageId="2" pageNumber="441">136–153</td>
</tr>
<tr id="3D8D2516A577D2BC41EDFE8FFA83FEB4" box="[166,1394,272,302]" gridrow="2" pageId="2" pageNumber="441">
<th id="7E5C4C6AA577D2BC41EDFE8FFF4AFEB4" box="[166,187,272,302]" gridcol="0" gridrow="2" pageId="2" pageNumber="441">P2</th>
<td id="7E5C4C6AA577D2BC41B5FE8FFE81FEB4" box="[254,368,272,302]" gridcol="1" gridrow="2" pageId="2" pageNumber="441">1496–1768</td>
<td id="7E5C4C6AA577D2BC40E0FE8FFDECFEB4" box="[427,541,272,302]" gridcol="2" gridrow="2" pageId="2" pageNumber="441">1292–1479</td>
<td id="7E5C4C6AA577D2BC4315FE8FFD35FEB4" box="[606,708,272,302]" gridcol="3" gridrow="2" pageId="2" pageNumber="441">476–544</td>
<td id="7E5C4C6AA577D2BC4259FE8FFC9AFEB4" box="[786,875,272,302]" gridcol="4" gridrow="2" pageId="2" pageNumber="441">238–289</td>
<td id="7E5C4C6AA577D2BC42F4FE8FFBE9FEB4" box="[959,1048,272,302]" gridcol="5" gridrow="2" pageId="2" pageNumber="441">170–204</td>
<td id="7E5C4C6AA577D2BC4527FE8FFB34FEB4" box="[1132,1221,272,302]" gridcol="6" gridrow="2" pageId="2" pageNumber="441">85–119</td>
<td id="7E5C4C6AA577D2BC4452FE8FFA83FEB4" box="[1305,1394,272,302]" gridcol="7" gridrow="2" pageId="2" pageNumber="441">119–153</td>
</tr>
<tr id="3D8D2516A577D2BC41EDFEA7FA83FECC" box="[166,1394,312,342]" gridrow="3" pageId="2" pageNumber="441">
<th id="7E5C4C6AA577D2BC41EDFEA7FF4AFECC" box="[166,187,312,342]" gridcol="0" gridrow="3" pageId="2" pageNumber="441">P3</th>
<td id="7E5C4C6AA577D2BC41B5FEA7FE81FECC" box="[254,368,312,342]" gridcol="1" gridrow="3" pageId="2" pageNumber="441">1615–1872</td>
<td id="7E5C4C6AA577D2BC40E0FEA7FDECFECC" box="[427,541,312,342]" gridcol="2" gridrow="3" pageId="2" pageNumber="441">1530–1768</td>
<td id="7E5C4C6AA577D2BC4315FEA7FD35FECC" box="[606,708,312,342]" gridcol="3" gridrow="3" pageId="2" pageNumber="441">850–1054</td>
<td id="7E5C4C6AA577D2BC4259FEA7FC9AFECC" box="[786,875,312,342]" gridcol="4" gridrow="3" pageId="2" pageNumber="441">442–527</td>
<td id="7E5C4C6AA577D2BC42F4FEA7FBE9FECC" box="[959,1048,312,342]" gridcol="5" gridrow="3" pageId="2" pageNumber="441">221–255</td>
<td id="7E5C4C6AA577D2BC4527FEA7FB34FECC" box="[1132,1221,312,342]" gridcol="6" gridrow="3" pageId="2" pageNumber="441">102–119</td>
<td id="7E5C4C6AA577D2BC4452FEA7FA83FECC" box="[1305,1394,312,342]" gridcol="7" gridrow="3" pageId="2" pageNumber="441">136–153</td>
</tr>
</table>
</paragraph>
<paragraph id="83022754A5772D4141DCFE1DFEE7FE00" blockId="2.[151,278,386,410]" box="[151,278,386,410]" pageId="2" pageNumber="441">
<emphasis id="B1C9FB46A5772D4141DCFE1DFEE7FE00" bold="true" box="[151,278,386,410]" pageId="2" pageNumber="441">
<tableNote id="7E5B26DAA5772D4141DCFE1DFEE7FE00" box="[151,278,386,410]" pageId="2" pageNumber="441" targetBox="[166,1394,192,342]" targetPageId="2">Continued.</tableNote>
</emphasis>
</paragraph>
<paragraph id="83022754A5772D4140DBFE34FADFFDA1" pageId="2" pageNumber="441">
<table id="F1BDD5F4A577D2BC41EDFE35FADFFDDB" box="[166,1326,426,577]" gridcols="5" gridrows="4" pageId="2" pageNumber="441" rowsContinueFrom="2.[166,1394,192,342]">
<tr id="3D8D2516A577D2BC41EDFE35FADFFE58" box="[166,1326,426,450]" gridrow="0" pageId="2" pageNumber="441" rowspan-0="1">
<th id="7E5C4C6AA577D2BC4025FE35FE22FE58" box="[366,467,426,450]" gridcol="1" gridrow="0" pageId="2" pageNumber="441">LR</th>
<th id="7E5C4C6AA577D2BC43DBFE35FD07FE58" box="[656,758,426,450]" gridcol="2" gridrow="0" pageId="2" pageNumber="441">BV</th>
<th id="7E5C4C6AA577D2BC42F8FE35FBE9FE58" box="[947,1048,426,450]" gridcol="3" gridrow="0" pageId="2" pageNumber="441">SV</th>
<th id="7E5C4C6AA577D2BC45A9FE35FADFFE58" box="[1250,1326,426,450]" gridcol="4" gridrow="0" pageId="2" pageNumber="441">BR</th>
</tr>
<tr id="3D8D2516A577D2BC41EDFE4DFADFFE6A" box="[166,1326,466,496]" gridrow="1" pageId="2" pageNumber="441">
<th id="7E5C4C6AA577D2BC41EDFE4DFF4AFE6A" box="[166,187,466,496]" gridcol="0" gridrow="1" pageId="2" pageNumber="441">P1</th>
<td id="7E5C4C6AA577D2BC4025FE4DFE22FE6A" box="[366,467,466,496]" gridcol="1" gridrow="1" pageId="2" pageNumber="441">0.57–0.64</td>
<td id="7E5C4C6AA577D2BC43DBFE4DFD07FE6A" box="[656,758,466,496]" gridcol="2" gridrow="1" pageId="2" pageNumber="441">4.05–4.42</td>
<td id="7E5C4C6AA577D2BC42F8FE4DFBE9FE6A" box="[947,1048,466,496]" gridcol="3" gridrow="1" pageId="2" pageNumber="441">3.15–3.41</td>
<td id="7E5C4C6AA577D2BC45A9FE4DFADFFE6A" box="[1250,1326,466,496]" gridcol="4" gridrow="1" pageId="2" pageNumber="441">0.9–1.3</td>
</tr>
<tr id="3D8D2516A577D2BC41EDFE65FADFFD82" box="[166,1326,506,536]" gridrow="2" pageId="2" pageNumber="441">
<th id="7E5C4C6AA577D2BC41EDFE65FF4AFD82" box="[166,187,506,536]" gridcol="0" gridrow="2" pageId="2" pageNumber="441">P2</th>
<td id="7E5C4C6AA577D2BC4025FE65FE22FD82" box="[366,467,506,536]" gridcol="1" gridrow="2" pageId="2" pageNumber="441">0.34–0,39</td>
<td id="7E5C4C6AA577D2BC43DBFE65FD07FD82" box="[656,758,506,536]" gridcol="2" gridrow="2" pageId="2" pageNumber="441">4.91–5.53</td>
<td id="7E5C4C6AA577D2BC42F8FE65FBE9FD82" box="[947,1048,506,536]" gridcol="3" gridrow="2" pageId="2" pageNumber="441">5.41–6.21</td>
<td id="7E5C4C6AA577D2BC45A9FE65FADFFD82" box="[1250,1326,506,536]" gridcol="4" gridrow="2" pageId="2" pageNumber="441">0.8–1.3</td>
</tr>
<tr id="3D8D2516A577D2BC41EDFDBCFADFFDDB" box="[166,1326,547,577]" gridrow="3" pageId="2" pageNumber="441">
<th id="7E5C4C6AA577D2BC41EDFDBCFF4AFDDB" box="[166,187,547,577]" gridcol="0" gridrow="3" pageId="2" pageNumber="441">P 3</th>
<td id="7E5C4C6AA577D2BC4025FDBCFE22FDDB" box="[366,467,547,577]" gridcol="1" gridrow="3" pageId="2" pageNumber="441">053–0.60</td>
<td id="7E5C4C6AA577D2BC43DBFDBCFD07FDDB" box="[656,758,547,577]" gridcol="2" gridrow="3" pageId="2" pageNumber="441">4.31–4.84</td>
<td id="7E5C4C6AA577D2BC42F8FDBCFBE9FDDB" box="[947,1048,547,577]" gridcol="3" gridrow="3" pageId="2" pageNumber="441">4.31–4.84</td>
<td id="7E5C4C6AA577D2BC45A9FDBCFADFFDDB" box="[1250,1326,547,577]" gridcol="4" gridrow="3" pageId="2" pageNumber="441">0.8–1.2</td>
</tr>
</table>
</paragraph>
<caption id="D7C277DCA5772D4141DCF801FD59F841" ID-DOI="http://doi.org/10.5281/zenodo.4955879" ID-Zenodo-Dep="4955879" httpUri="https://zenodo.org/record/4955879/files/figure.png" pageId="2" pageNumber="441" startId="2.[151,264,1950,1974]" targetBox="[151,1435,596,1927]" targetPageId="2">
<paragraph id="83022754A5772D4141DCF801FD59F841" blockId="2.[151,1436,1950,2011]" pageId="2" pageNumber="441">
<emphasis id="B1C9FB46A5772D4141DCF801FECAF82D" bold="true" box="[151,315,1950,1975]" pageId="2" pageNumber="441">FIGURES 1–4.</emphasis>
Males of
<taxonomicName id="44BD5CD7A5772D4140E2F800FD47F82C" authority="Hansen" authorityName="Hansen" authorityYear="1976" box="[425,694,1951,1975]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="leoniella">
<emphasis id="B1C9FB46A5772D4140E2F800FD92F82D" box="[425,611,1951,1975]" italics="true" pageId="2" pageNumber="441">Diamesa leoniella</emphasis>
Hansen
</taxonomicName>
(1–2) and
<taxonomicName id="44BD5CD7A5772D41426DF800FBF9F82D" authority="Tokunaga" authorityName="Tokunaga" authorityYear="1936" box="[806,1032,1951,1975]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B1C9FB46A5772D41426DF800FC6FF82C" box="[806,926,1951,1974]" italics="true" pageId="2" pageNumber="441">D. japonica</emphasis>
Tokunaga
</taxonomicName>
(3–4).
<emphasis id="B1C9FB46A5772D414518F800FB8EF82D" bold="true" box="[1107,1151,1951,1975]" pageId="2" pageNumber="441">1, 4,</emphasis>
central part of hypopygium without tergite IX;
<emphasis id="B1C9FB46A5772D414011F85CFE77F841" bold="true" box="[346,390,1987,2011]" pageId="2" pageNumber="441">2–3,</emphasis>
hypopygium in dorsal view.
</paragraph>
</caption>
<paragraph id="83022754A5762D40418CFF08FBB8FEDA" blockId="3.[151,1436,151,392]" pageId="3" pageNumber="442">
Hypopygium (
<figureCitation id="1B863BD1A5762D404028FF08FE3AFF2A" box="[355,459,151,176]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1950,1974]" captionTargetBox="[151,1435,596,1927]" captionTargetId="figure-144@2.[151,1435,596,1927]" captionTargetPageId="2" captionText="FIGURES 1–4. Males of Diamesa leoniella Hansen (1–2) and D. japonica Tokunaga (3–4). 1, 4, central part of hypopygium without tergite IX; 2–3, hypopygium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4955879" httpUri="https://zenodo.org/record/4955879/files/figure.png" pageId="3" pageNumber="442">Figs. 3–4</figureCitation>
). Tergite IX with 10–11 setae (from one side), 20–28 μm long and wedge-shaped anal point 63–136 μm long, covered in basal 2/3 with numerous microtrichia forward and laterally directed. Laterosternite IX with 9–10 short setae. Transverse sternapodeme triangular
<emphasis id="B1C9FB46A5762D404394FF7FFD09FF6D" box="[735,760,224,247]" italics="true" pageId="3" pageNumber="442">ca</emphasis>
128 μm long. Gonocoxite 350–476 μm long. Inferior volsella absent. Superior volsella in form of angular tubercle. Gonostylus slightly curved, 220–236 μm long, with numerous, proximally directed strong setae, 44–52 μm long; megaseta 6–8 μm long. HR 1.92–2.23.
</paragraph>
</subSubSection>
<subSubSection id="CBA774DFA5762D40418CFED4FDF5FE12" pageId="3" pageNumber="442" type="distribution">
<paragraph id="83022754A5762D40418CFED4FDF5FE12" blockId="3.[151,1436,151,392]" pageId="3" pageNumber="442">
<emphasis id="B1C9FB46A5762D40418CFED4FEABFEF9" bold="true" box="[199,346,331,355]" inLineHeading="true" pageId="3" pageNumber="442" reason="1">Distribution.</emphasis>
East Palaearctic island species. Known from
<collectingCountry id="FBAA67C4A5762D40421CFED3FC65FEF9" box="[855,916,332,355]" name="Japan" pageId="3" pageNumber="442">Japan</collectingCountry>
and Russian Far East (Kunashir Island, Kurile Islands (
<bibRefCitation id="E72C5AA5A5762D4041BAFEF0FE09FE12" author="Ashe, P. &amp; O'Connor, J. P." box="[241,504,367,392]" pageId="3" pageNumber="442" refId="ref2269" refString="Ashe, P. &amp; O'Connor, J. P. (2009) A World Catalogue of Chironomidae (Diptera). Part 1. Buchonomyiinae, Chilenomyiinae, Podonominae, Aphroteniinae, Tanypodinae, Usambaromyiinae, Diamesinae, Prodiamesinae and Telmatogetoninae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, 445 pp." type="book" year="2009">Ashe &amp; O’Connor 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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