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<document id="DE391E85E90EB1B7DFE57F1E8755077E" ID-DOI="10.11646/zootaxa.3937.2.4" ID-GBIF-Dataset="b66a4a8e-cd41-4362-ba56-cff77ff8adcc" ID-ISSN="1175-5326" ID-Zenodo-Dep="243111" ID-ZooBank="46253C57-B237-4A7C-B110-49F79290CAE9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461020628306" checkinUser="plazi" docAuthor="Gómez, Isrrael C., Sääksjärvi, Ilari E., Puhakka, Liisa, Castillo, Carol &amp; Bordera, Santiago" docDate="2015" docId="03FE87F5FFB45263FF06959FFE47F979" docLanguage="en" docName="zt03937p336.pdf" docOrigin="Zootaxa 3937 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epirhyssa wisei Porter 1978" docType="treatment" docVersion="8" lastPageNumber="332" masterDocId="FFC7FF8DFFA15276FF919428FFE5FFA7" masterDocTitle="The Peruvian Amazonian species of Epirhyssa Cresson (Hymenoptera: Ichneumonidae: Rhyssinae), with notes on tropical species richness" masterLastPageNumber="336" masterPageNumber="311" pageNumber="332" updateTime="1698570012193" updateUser="plazi">
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<mods:title id="870FA4BD169E7F68C575295583BC1D9F">The Peruvian Amazonian species of Epirhyssa Cresson (Hymenoptera: Ichneumonidae: Rhyssinae), with notes on tropical species richness</mods:title>
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<mods:namePart id="AFB1DBCE6F27093A755CD20E15A8D0CE">Gómez, Isrrael C.</mods:namePart>
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<mods:namePart id="F2AC6C008D0F291AEC43EDCE31232466">Sääksjärvi, Ilari E.</mods:namePart>
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<mods:namePart id="75799966A6CDE5B870AFC885DFDE45C3">Puhakka, Liisa</mods:namePart>
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<mods:namePart id="495A9AF8DFFD5E613F0E3742CB433599">Castillo, Carol</mods:namePart>
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<mods:namePart id="643ED2358550F066E74867F0FE0C9289">Bordera, Santiago</mods:namePart>
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<mods:date id="A2232DE667ABA7E999D1419D8A5C8EDA">2015</mods:date>
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<mods:number id="3EB859239F7ED3BC327920B227F03A31">3937</mods:number>
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<treatment id="03FE87F5FFB45263FF06959FFE47F979" ID-DOI="http://doi.org/10.5281/zenodo.6110505" ID-GBIF-Taxon="119602727" ID-Zenodo-Dep="6110505" LSID="urn:lsid:plazi:treatment:03FE87F5FFB45263FF06959FFE47F979" httpUri="http://treatment.plazi.org/id/03FE87F5FFB45263FF06959FFE47F979" lastPageNumber="332" pageId="21" pageNumber="332">
<subSubSection id="C34D6568FFB45263FF06959FFEC9FE54" pageId="21" pageNumber="332" type="nomenclature">
<paragraph id="8BE836E3FFB45263FF06959FFE0BFE75" blockId="21.[151,494,439,499]" box="[151,494,439,466]" pageId="21" pageNumber="332">
<heading id="D0A0818FFFB45263FF06959FFE0BFE75" bold="true" box="[151,494,439,466]" fontSize="11" level="1" pageId="21" pageNumber="332" reason="1">
<taxonomicName id="4C574D60FFB45263FF06959FFE0BFE75" ID-CoL="6G8BP" authority="Porter 1978" authorityName="Porter" authorityYear="1978" box="[151,494,439,466]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="332" phylum="Arthropoda" rank="species" species="wisei">
<emphasis id="B923EAF1FFB45263FF06959FFE0BFE75" bold="true" box="[151,494,439,466]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF06959FFEB6FE76" bold="true" box="[151,339,439,465]" italics="true" pageId="21" pageNumber="332">Epirhyssa wisei</emphasis>
<bibRefCitation id="EFC64B12FFB45263FECB9590FE0BFE75" author="Porter" box="[346,494,440,466]" pageId="21" pageNumber="332" refString="Porter, C. C. (1978) A revision of the genus Epirhyssa (Hymenoptera, Ichneumonidae). Studia Entomologica, 20, 297 - 318." type="journal article" year="1978">Porter 1978</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE836E3FFB45263FF0695F2FEC9FE54" blockId="21.[151,494,439,499]" box="[151,300,474,499]" pageId="21" pageNumber="332">
(
<figureCitation id="136C2A66FFB45263FF0E95F2FF00FE54" box="[159,229,474,499]" captionStart="FIGURE 2" captionStartId="4.[151,250,1833,1855]" captionTargetBox="[151,1434,629,1810]" captionTargetId="figure@4.[151,1436,629,1811]" captionTargetPageId="4" captionText="FIGURE 2. Head of Epirhyssa spp.: frontal view: A— E. cochabambae; B— E. braconoides (paratype); C— E. zaphyma; D— E. amazonica; E— E. diatropis; F— E. wisei, (female); G— E. isthmia; H— E. simpirae sp. n. (holotype)." httpUri="https://zenodo.org/record/243113/files/figure.png" pageId="21" pageNumber="332">Figs 2</figureCitation>
F, 4D)
</paragraph>
</subSubSection>
<subSubSection id="C34D6568FFB45263FF06960AFAD5FD6C" pageId="21" pageNumber="332" type="materials_examined">
<paragraph id="8BE836E3FFB45263FF06960AFF04FDF9" blockId="21.[151,1436,546,1759]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF06960AFCEDFD9C" bold="true" box="[151,776,546,571]" pageId="21" pageNumber="332">
Material examined.
<typeStatus id="54EC8841FFB45263FE01960AFE2EFD9C" box="[400,459,546,571]" pageId="21" pageNumber="332">Type</typeStatus>
material:
<typeStatus id="54EC8841FFB45263FDC1960AFD5AFD9C" box="[592,703,546,571]" pageId="21" pageNumber="332" type="holotype">Holotype</typeStatus>
, male
</emphasis>
(
<collectionCode id="ED46AE26FFB45263FC8D960BFC80FD9C" box="[796,869,547,571]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="332">FSCA</collectionCode>
)
<collectingCountry id="F3407673FFB45263FCE9960BFC4BFD9D" box="[888,942,547,570]" name="Peru" pageId="21" pageNumber="332">Peru</collectingCountry>
, Tingo María,
<specimenCount id="9D51FD6AFFB45263FBF0960AFB50FD9D" box="[1121,1205,546,571]" pageId="21" pageNumber="332" type="generic">
<quantity id="4CAF9B06FFB45263FBF0960AFB54FD9D" box="[1121,1201,546,571]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="21" pageNumber="332" unit="m" value="620.0">620 m</quantity>
.
</specimenCount>
,
<date id="FFE91023FFB45263FB54960AFAF3FD9C" box="[1221,1302,546,571]" pageId="21" pageNumber="332" value="2012-10-05">X-5/12</date>
-1964, C.C. Porter.
</paragraph>
<paragraph id="8BE836E3FFB45263FF569642FEBFFD00" blockId="21.[151,1436,546,1759]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF569642FDB2FD24" bold="true" box="[199,599,618,643]" pageId="21" pageNumber="332">New material examined: Female</emphasis>
(
<collectionCode id="ED46AE26FFB45263FDFB9643FD34FD24" box="[618,721,619,643]" pageId="21" pageNumber="332">MEKRB</collectionCode>
)
<collectingCountry id="F3407673FFB45263FD759643FCFFFD25" box="[740,794,619,642]" name="Peru" pageId="21" pageNumber="332">Peru</collectingCountry>
, La Merced, Fundo La Genova,
<collectionCode id="ED46AE26FFB45263FB319643FB13FD24" box="[1184,1270,619,643]" pageId="21" pageNumber="332">AECID</collectionCode>
A/013484/07,
<date id="FFE91023FFB45263FF0696A6FEB3FD00" box="[151,342,654,679]" pageId="21" pageNumber="332" value="2008-08-16" valueMax="2008-08-30" valueMin="2008-08-16">1630.VIII.2008</date>
.
</paragraph>
<paragraph id="8BE836E3FFB45263FF56969AFAD5FD6C" blockId="21.[151,1436,546,1759]" box="[199,1328,690,715]" pageId="21" pageNumber="332">
Female, (
<collectionCode id="ED46AE26FFB45263FEBE969BFE9BFD6C" box="[303,382,691,715]" pageId="21" pageNumber="332">CEUA</collectionCode>
)
<collectingCountry id="F3407673FFB45263FE1C969BFE26FD6D" box="[397,451,691,714]" name="Peru" pageId="21" pageNumber="332">Peru</collectingCountry>
, Malaise 2, La Merced, Fundo La Genova,
<collectionCode id="ED46AE26FFB45263FC3A969BFBE5FD6C" box="[939,1024,691,715]" pageId="21" pageNumber="332">AECID</collectionCode>
A/013484/07,
<date id="FFE91023FFB45263FB38969BFAC9FD6C" box="[1193,1324,691,715]" pageId="21" pageNumber="332" value="2008-06-21">21.VI.2008</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D6568FFB45263FF5696FEFCBBFA18" pageId="21" pageNumber="332" type="description">
<paragraph id="8BE836E3FFB45263FF5696FEFDC2FB40" blockId="21.[151,1436,546,1759]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF5696FEFEFBFD48" bold="true" box="[199,286,726,751]" pageId="21" pageNumber="332">Female</emphasis>
(
<figureCitation id="136C2A66FFB45263FEA596FEFE9BFD48" box="[308,382,726,751]" captionStart="FIGURE 4" captionStartId="7.[151,250,1471,1493]" captionTargetBox="[153,1433,193,1449]" captionTargetId="figure@7.[153,1433,193,1449]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of Epirhyssa spp.: (females): A— E. electronigra sp. n. (holotype); B— E. cuscoensis sp. n. (holotype); C— E. diatropis (paratype); D— E. wisei; E— E. isthmia; F— E. prolasia." httpUri="https://zenodo.org/record/243115/files/figure.png" pageId="21" pageNumber="332">Fig. 4</figureCitation>
D). Fore wing length about
<quantity id="4CAF9B06FFB45263FD4796FFFCC5FD49" box="[726,800,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="332" unit="mm" value="8.0">8 mm</quantity>
.
<emphasis id="B923EAF1FFB45263FCA096FEFC94FD48" bold="true" box="[817,881,726,751]" pageId="21" pageNumber="332">Head</emphasis>
with occipital carina absent; posterior ocellus separated from eye by about 1.5 times its diameter; mandibles stout, lower tooth clearly longer than upper one; clypeus (
<figureCitation id="136C2A66FFB45263FF6F9736FEA7FC90" box="[254,322,798,823]" captionStart="FIGURE 2" captionStartId="4.[151,250,1833,1855]" captionTargetBox="[151,1434,629,1810]" captionTargetId="figure@4.[151,1436,629,1811]" captionTargetPageId="4" captionText="FIGURE 2. Head of Epirhyssa spp.: frontal view: A— E. cochabambae; B— E. braconoides (paratype); C— E. zaphyma; D— E. amazonica; E— E. diatropis; F— E. wisei, (female); G— E. isthmia; H— E. simpirae sp. n. (holotype)." httpUri="https://zenodo.org/record/243113/files/figure.png" pageId="21" pageNumber="332">Fig. 2</figureCitation>
F) with central part concave, centrally with a small apical tubercle; lower face about 1.1 times as broad as medially long, polished, with sparse yellowish setae; antenna slender with about 32 flagellar segments; gena polished with sparse setae.
<emphasis id="B923EAF1FFB45263FD90974EFD9AFCD8" bold="true" box="[513,639,870,895]" pageId="21" pageNumber="332">Mesosoma</emphasis>
with epomia diverging from anterior margin of pronotum; mesoscutum with central lobe not overhanging pronotum, transverse rugae short, with fairly sparse, short pubescence; mesopleuron polished, with sparse, long setiferous punctures; subalar prominence strongly inflated; epicnemial carina ventrally reaching above level of lower corner of pronotum; hind coxa in profile about 2.2 times as wide as deep; fore wing with
<emphasis id="B923EAF1FFB45263FE1A97DFFE59FBA9" box="[395,444,1015,1038]" italics="true" pageId="21" pageNumber="332">cu-a</emphasis>
slightly distal to
<emphasis id="B923EAF1FFB45263FD1597DFFD29FBA9" box="[644,716,1015,1038]" italics="true" pageId="21" pageNumber="332">Rs&amp;M</emphasis>
; first subdiscal cell broadened distally, its width near outer end about 1.8 times its basal width; abscissa of
<emphasis id="B923EAF1FFB45263FD119034FD72FB94" box="[640,663,1052,1075]" italics="true" pageId="21" pageNumber="332">M</emphasis>
between
<emphasis id="B923EAF1FFB45263FC959033FCA0FB94" box="[772,837,1051,1075]" italics="true" pageId="21" pageNumber="332">2rs-m</emphasis>
and
<emphasis id="B923EAF1FFB45263FCEC9033FC26FB94" box="[893,963,1051,1075]" italics="true" pageId="21" pageNumber="332">2m-cu</emphasis>
about 0.4 times as long
<emphasis id="B923EAF1FFB45263FB469033FAFCFB94" box="[1239,1305,1051,1075]" italics="true" pageId="21" pageNumber="332">2rs-m</emphasis>
,
<emphasis id="B923EAF1FFB45263FAB69034FAA6FB94" box="[1319,1347,1052,1075]" italics="true" pageId="21" pageNumber="332">Rs</emphasis>
weakly and evenly bowed; propodeum short, highly polished, with few setae and with no trace of median longitudinal depression; submetapleural carina sharp throughout.
<emphasis id="B923EAF1FFB45263FD74904AFC84FBDC" bold="true" box="[741,865,1122,1147]" pageId="21" pageNumber="332">Metasoma</emphasis>
with tergite I about 2.0 times as long as posteriorly broad, dorsally smooth and highly polished; tergite II about 1.0 times as long as posteriorly broad, smooth and highly polished, tergites III+ with fine short dense pubescence; ovipositor projecting beyond apex of metasoma by about 4.5 times length of hind tibia.
</paragraph>
<paragraph id="8BE836E3FFB45263FF5690DAFCEAFA3C" blockId="21.[151,1436,546,1759]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF5690DAFEAAFAAC" bold="true" box="[199,335,1266,1291]" pageId="21" pageNumber="332">Coloration.</emphasis>
A primarily yellow species, with following areas black: interocellar area and dorsal transverse band from eye to eye, adjacent area to occipital carina, small line from pronotum going down to propleura, anterior and posterior margins of mesopleura and propodeum, scutoscutellar groove and lower part of metanotum; narrow apical bands on tergites IIIVII. Flagellum brownish, ovipositor sheath brownish. Wings yellowish, fore wing with a small weak dark patch apically; pterostigma brownish.
</paragraph>
<paragraph id="8BE836E3FFB45263FF56918EFCBBFA18" blockId="21.[151,1436,546,1759]" box="[199,862,1446,1471]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF56918EFEE1FA18" bold="true" box="[199,260,1446,1471]" pageId="21" pageNumber="332">Male</emphasis>
. Similar to female (male described by
<bibRefCitation id="EFC64B12FFB45263FD26918FFCBCFA18" author="Porter" box="[695,857,1447,1471]" pageId="21" pageNumber="332" refString="Porter, C. C. (1978) A revision of the genus Epirhyssa (Hymenoptera, Ichneumonidae). Studia Entomologica, 20, 297 - 318." type="journal article" year="1978">Porter (1978))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D6568FFB45263FF5691E2FB7BF91C" pageId="21" pageNumber="332" type="diagnosis">
<paragraph id="8BE836E3FFB45263FF5691E2FC30F9D5" blockId="21.[151,1436,546,1759]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF5691E2FEDFFA44" bold="true" box="[199,314,1482,1507]" pageId="21" pageNumber="332">Diagnosis</emphasis>
. This species can be distinguished from all other Neotropical
<taxonomicName id="4C574D60FFB45263FC6F91E2FB8BFA44" box="[1022,1134,1482,1507]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAF1FFB45263FC6F91E2FB8BFA44" box="[1022,1134,1482,1507]" italics="true" pageId="21" pageNumber="332">Epirhyssa</emphasis>
</taxonomicName>
by the combination of the following characters: 1) occipital carina absent; 2) tergite I highly polished with very few setae; 3) yellowish annulus over antennal segments 79; and 4) fore wing yellowish, weakly infumate.
<taxonomicName id="4C574D60FFB45263FBE1923AFAC1F98C" box="[1136,1316,1554,1579]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="332" phylum="Arthropoda" rank="species" species="wisei">
<emphasis id="B923EAF1FFB45263FBE1923AFAC1F98C" box="[1136,1316,1554,1579]" italics="true" pageId="21" pageNumber="332">Epirhyssa wisei</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C574D60FFB45263FE9D921FFDA0F9E9" authority="Porter. However" authorityName="Porter. However" box="[268,581,1590,1615]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="332" phylum="Arthropoda" rank="species" species="isthmia">
<emphasis id="B923EAF1FFB45263FE9D921FFEF8F9E9" box="[268,285,1591,1614]" italics="true" pageId="21" pageNumber="332">E</emphasis>
.
<emphasis id="B923EAF1FFB45263FEBC921EFE65F9E8" box="[301,384,1590,1615]" italics="true" pageId="21" pageNumber="332">isthmia</emphasis>
Porter. However
</taxonomicName>
, the less elongate body, the lack of a strong apical black band on the fore wing, and the shorter yellow annulus on the antenna differentiate
<taxonomicName id="4C574D60FFB45263FCE49274FC28F9D4" box="[885,973,1628,1651]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="332" phylum="Arthropoda" rank="species" species="wisei">
<emphasis id="B923EAF1FFB45263FCE49274FC28F9D4" box="[885,973,1628,1651]" italics="true" pageId="21" pageNumber="332">E. wisei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE836E3FFB45263FF569256FB7BF91C" blockId="21.[151,1436,546,1759]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF569256FEEFF930" bold="true" box="[199,266,1662,1687]" pageId="21" pageNumber="332">Notes</emphasis>
. The species was originally described based on a single male specimen, so we provide a full description of the female. It has only been found in the Peruvian Amazonia, close to the tropical Andes.
</paragraph>
</subSubSection>
<subSubSection id="C34D6568FFB45263FF5692EEFE47F979" box="[199,418,1734,1759]" pageId="21" pageNumber="332" type="distribution">
<paragraph id="8BE836E3FFB45263FF5692EEFE47F979" blockId="21.[151,1436,546,1759]" box="[199,418,1734,1759]" pageId="21" pageNumber="332">
<emphasis id="B923EAF1FFB45263FF5692EEFE85F978" bold="true" box="[199,352,1734,1759]" pageId="21" pageNumber="332">Distribution.</emphasis>
<collectingCountry id="F3407673FFB45263FEF692EFFE7BF979" box="[359,414,1735,1758]" name="Peru" pageId="21" pageNumber="332">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>