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<document id="F22BEB6AB3FF78E2E9CAC0AB2E157947" ID-DOI="10.5281/zenodo.5352700" ID-ISSN="2345-7600" ID-Zenodo-Dep="5352700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208580763" checkinUser="tatiana" docAuthor="Polhemus, Dan A. &amp; Polhemus, John T." docDate="2013" docId="03FE7253F509FFF4FC99FCBAFD12FDCA" docLanguage="en" docName="61rbz665-686.pdf" docOrigin="Raffles Bulletin of Zoology 61 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Aphelocheirus Westwood 1833" docType="key" docVersion="3" lastPageNumber="682" masterDocId="FFC70A2BF519FFE5FFFEFFA4FFFDFFE7" masterDocTitle="Guide To The Aquatic Heteroptera Of Singapore And Peninsular Malaysia. Xi. Infraorder Nepomorpha- Families Naucoridae And Aphelocheiridae" masterLastPageNumber="686" masterPageNumber="665" pageNumber="681" updateTime="1699186152900" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E1B311AA66B82DD6565E2B26E504E8FC">Guide To The Aquatic Heteroptera Of Singapore And Peninsular Malaysia. Xi. Infraorder Nepomorpha- Families Naucoridae And Aphelocheiridae</mods:title>
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<mods:name id="C64E13E1EE864EA4B5AD240B1C79547C" type="personal">
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<mods:namePart id="E41356EDF4E1CCA07146B4EEAB10BB26">Polhemus, Dan A.</mods:namePart>
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<mods:namePart id="3F936EAB5D00105CF927AECC880E5FAF">Polhemus, John T.</mods:namePart>
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<mods:typeOfResource id="F4177575EB2EE5F9A204828425721879">text</mods:typeOfResource>
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<mods:title id="1EB07D2FBDFD6EB14C0F9132CF99184E">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="06D17124235AE9C720EDA6993609AD5B">
<mods:date id="335D69028334D9A172C675B9C0CB67D9">2013</mods:date>
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<mods:number id="73B2EF5D9005CFA491A8E2C719E5970C">2013-08-30</mods:number>
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<mods:detail id="8B95BAA2AAE1DC936EDB5896148460D9" type="volume">
<mods:number id="2BF6E178D52BE59ADD334F3EB3E589F9">61</mods:number>
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<mods:number id="706690ED03D49C18B4913FBA0DA632E5">2</mods:number>
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<mods:start id="F081706FDA90C9989346C9BD5587F393">665</mods:start>
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<mods:classification id="62D7A396CEFC355210B8721666FE1566">journal article</mods:classification>
<mods:identifier id="681438BC61D1544573DA557712B27AC9" type="DOI">10.5281/zenodo.5352700</mods:identifier>
<mods:identifier id="D67D46A1A90FD3B20A86BF9AA12ED96D" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03FE7253F509FFF4FC99FCBAFD12FDCA" LSID="urn:lsid:plazi:treatment:03FE7253F509FFF4FC99FCBAFD12FDCA" httpUri="http://treatment.plazi.org/id/03FE7253F509FFF4FC99FCBAFD12FDCA" lastPageId="17" lastPageNumber="682" pageId="16" pageNumber="681">
<subSubSection id="C34D90CEF509FFF5FC99FCBAFA64FC91" pageId="16" pageNumber="681" type="nomenclature">
<paragraph id="8BE8C345F509FFF5FC99FCBAFA62FCD1" blockId="16.[871,1439,798,886]" box="[871,1439,798,822]" pageId="16" pageNumber="681">
<heading id="D0A07429F509FFF5FC99FCBAFA62FCD1" allCaps="true" box="[871,1439,798,822]" centered="true" fontSize="10" level="1" pageId="16" pageNumber="681" reason="1">
<emphasis id="B9231F57F509FFF5FC99FCBAFA62FCD1" bold="true" box="[871,1439,798,822]" pageId="16" pageNumber="681">
KEY TO THE SPECIES OF
<taxonomicName id="4C57B8C6F509FFF5FB42FCBAFA62FCD1" ID-CoL="62CXH" authorityName="Westwood" authorityYear="1833" box="[1212,1439,798,822]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="681" phylum="Arthropoda" rank="genus">
<emphasis id="B9231F57F509FFF5FB42FCBAFA62FCD1" bold="true" box="[1212,1439,798,822]" italics="true" pageId="16" pageNumber="681">APHELOCHEIRUS</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE8C345F509FFF5FC81FC9AFA64FC91" blockId="16.[871,1439,798,886]" pageId="16" pageNumber="681">
occurring in
<collectingCountry id="F34083D5F509FFF5FBF4FC9AFB8BFCB1" box="[1034,1142,830,854]" name="Singapore" pageId="16" pageNumber="681">Singapore</collectingCountry>
and Peninsular
<collectingCountry id="F34083D5F509FFF5FADBFC9AFA7AFCB1" box="[1317,1415,830,854]" name="Malaysia" pageId="16" pageNumber="681">Malaysia</collectingCountry>
(modified from D. Polhemus &amp; J. Polhemus, 1989)
</paragraph>
</subSubSection>
<subSubSection id="C34D90CEF509FFF4FCBAFC39FD12FDCA" lastPageId="17" lastPageNumber="682" pageId="16" pageNumber="681" type="key">
<keyStep id="30A3DBE0F509FFF5FCBAFC39FA3FFBC2" pageId="16" pageNumber="681">
<paragraph id="8BE8C345F509FFF5FCBAFC39FA3FFC0B" blockId="16.[836,1475,925,1120]" pageId="16" pageNumber="681">
<keyLead id="30A66070F509FFF5FCBAFC39FA3FFC0B" pageId="16" pageNumber="681">
1. Small species, body length less than
<quantity id="4CAF6EA0F509FFF5FAF2FC39FAB6FC55" box="[1292,1355,925,947]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="681" unit="mm" value="5.0">5 mm</quantity>
(subgenus
<taxonomicName id="4C57B8C6F509FFF5FC94FC1FFBDCFC28" ID-CoL="8P3Q9" authorityName="Hoberlandt &amp; Stys" authorityYear="1979" box="[874,1057,955,975]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="681" phylum="Arthropoda" rank="subGenus" subGenus="Micraphelocheirus">
<emphasis id="B9231F57F509FFF5FC94FC1FFBDCFC28" box="[874,1057,955,975]" italics="true" pageId="16" pageNumber="681">Micraphelocheirus</emphasis>
</taxonomicName>
) ................................................................... ...........................
<taxonomicName id="4C57B8C6F509FFF5FBEBFC7CFA3FFC0B" ID-CoL="7QYQ7" authority="Zettel &amp; Papacek" authorityName="Zettel &amp; Papacek" authorityYear="2006" box="[1045,1474,982,1004]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="681" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F509FFF5FBEBFC7CFAECFC0B" box="[1045,1297,984,1004]" italics="true" pageId="16" pageNumber="681">Aphelocheirus malayensis</emphasis>
Zettel &amp; Papáček
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE8C345F509FFF5FCBAFC50FA3FFBC2" blockId="16.[836,1475,925,1120]" pageId="16" pageNumber="681">
<keyLead id="30A66070F509FFF5FCBAFC50FA3FFBC2" pageId="16" pageNumber="681">
– Larger species, body length exceeding
<quantity id="4CAF6EA0F509FFF5FAEFFC50FAB3FBEE" box="[1297,1358,1012,1033]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="681" unit="mm" value="7.0">7 mm</quantity>
(subgenus
<taxonomicName id="4C57B8C6F509FFF5FC94FBB5FC08FBC2" ID-CoL="62CXH" authorityName="Westwood" authorityYear="1833" box="[874,1013,1041,1061]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="681" phylum="Arthropoda" rank="genus">
<emphasis id="B9231F57F509FFF5FC94FBB5FC08FBC2" box="[874,1013,1041,1061]" italics="true" pageId="16" pageNumber="681">Aphelocheirus</emphasis>
</taxonomicName>
).........................................................................2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A3DBE0F509FFF4FCBAFB89FD12FEA1" lastPageId="17" lastPageNumber="682" pageId="16" pageNumber="681">
<paragraph id="8BE8C345F509FFF4FCBAFB89FD12FF17" blockId="16.[836,1475,925,1120]" lastBlockId="17.[113,753,189,557]" lastPageId="17" lastPageNumber="682" pageId="16" pageNumber="681">
<keyLead id="30A66070F509FFF4FCBAFB89FD12FF17" lastPageId="17" lastPageNumber="682" pageId="16" pageNumber="681">
2. Male with conspicuous projecting tab on abdominal ventrite IV (
<figureCitation id="136CDFC0F509FFF5FC6CFBEEFC21FB87" box="[914,988,1098,1120]" captionStart="Figs" captionStartId="18.[137,181,1918,1940]" captionTargetBox="[136,776,862,1891]" captionTargetId="figure-371@18.[118,795,850,1903]" captionTargetPageId="18" captionText="Figs. 47–52.Aphelocheirus (Aphelocheirus) grik D. Polhemus &amp; J. Polhemus, structural details. 47. Brachypterous male, dorsal habitus, Malaysia, Perak, Grik. 48. Macropterous female, dorsal habitus, specimen from Thailand, Chiang Mai Prov.. Mae Ping River. 49. Male ventral abdomen, showing tab on abdominal ventrite IV. 50. Female subgenital plate. 51. Male parameres and phallotheca. 52. Proacetabula." figureDoi="http://doi.org/10.5281/zenodo.5352734" httpUri="https://zenodo.org/record/5352734/files/figure.png" pageId="16" pageNumber="681">Fig. 49</figureCitation>
); female subgenital plate very short and truncate, broader than long, with two long hair tufts on posterior margin (
<figureCitation id="136CDFC0F508FFF4FF60FF7EFF18FF17" box="[158,229,218,240]" captionStart="Figs" captionStartId="18.[137,181,1918,1940]" captionTargetBox="[136,776,862,1891]" captionTargetId="figure-371@18.[118,795,850,1903]" captionTargetPageId="18" captionText="Figs. 47–52.Aphelocheirus (Aphelocheirus) grik D. Polhemus &amp; J. Polhemus, structural details. 47. Brachypterous male, dorsal habitus, Malaysia, Perak, Grik. 48. Macropterous female, dorsal habitus, specimen from Thailand, Chiang Mai Prov.. Mae Ping River. 49. Male ventral abdomen, showing tab on abdominal ventrite IV. 50. Female subgenital plate. 51. Male parameres and phallotheca. 52. Proacetabula." figureDoi="http://doi.org/10.5281/zenodo.5352734" httpUri="https://zenodo.org/record/5352734/files/figure.png" pageId="17" pageNumber="682">Fig. 50</figureCitation>
) ...........................
<taxonomicName id="4C57B8C6F508FFF4FE6FFF7FFD12FF17" ID-CoL="7QYLB" authority="D. Polhemus &amp; J. Polhemus" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="1989" box="[401,751,218,240]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="682" phylum="Arthropoda" rank="species" species="grik">
<emphasis id="B9231F57F508FFF4FE6FFF7FFE5DFF08" box="[401,416,219,239]" italics="true" pageId="17" pageNumber="682">A</emphasis>
.
<emphasis id="B9231F57F508FFF4FE53FF7FFE29FF08" box="[429,468,219,239]" italics="true" pageId="17" pageNumber="682">grik</emphasis>
D. Polhemus &amp; J. Polhemus
</taxonomicName>
</keyLead>
</paragraph>
<caption id="DF2893CDF509FFF5FF74F81CFE14F7A5" ID-DOI="http://doi.org/10.5281/zenodo.5352730" ID-Zenodo-Dep="5352730" httpUri="https://zenodo.org/record/5352730/files/figure.png" pageId="16" pageNumber="681" startId="16.[138,182,1976,1998]" targetBox="[159,1460,1145,1971]" targetPageId="16">
<paragraph id="8BE8C345F509FFF5FF74F81CFE14F7A5" blockId="16.[136,776,1976,2114]" pageId="16" pageNumber="681">
Figs. 37–39.
<taxonomicName id="4C57B8C6F509FFF5FEF2F81EFF44F80C" ID-CoL="3FY5F" authority="D. Polhemus, J. Polhemus &amp; Sites" authorityName=", D. Polhemus, J. Polhemus &amp; Sites" authorityYear="2008" class="Insecta" family="Naucoridae" genus="Gestroiella" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="681" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B9231F57F509FFF5FEF2F81EFE20F829" box="[268,477,1978,1998]" italics="true" pageId="16" pageNumber="681">Gestroiella siamensis</emphasis>
D. Polhemus, J. Polhemus &amp; Sites
</taxonomicName>
, structural details, specimen from Thailand, Thailand, Songkla Prov., Ton Nga Chang. 37. Female terminal abdomen, ventral view, showing shape of subgenital plate. 38. Male left paramere. 39. Medial process of male phallotheca.
</paragraph>
</caption>
<caption id="DF2893CDF509FFF5FCBAF872FC09F7A5" pageId="16" pageNumber="681" startId="16.[836,875,2006,2028]">
<paragraph id="8BE8C345F509FFF5FCBAF872FC09F7A5" blockId="16.[836,1474,2006,2114]" pageId="16" pageNumber="681">
Fig. 40.
<taxonomicName id="4C57B8C6F509FFF5FC59F873FA41F80B" ID-CoL="7QYNG" authority="Horvath" authorityName="Horvath" authorityYear="1899" box="[935,1468,2006,2028]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="681" phylum="Arthropoda" rank="species" species="pallens" subGenus="Aphelocheirus">
<emphasis id="B9231F57F509FFF5FC59F873FBBEF80C" box="[935,1091,2007,2027]" italics="true" pageId="16" pageNumber="681">Aphelocheirus</emphasis>
(
<emphasis id="B9231F57F509FFF5FBA6F873FB09F80C" box="[1112,1268,2007,2027]" italics="true" pageId="16" pageNumber="681">Aphelocheirus</emphasis>
)
<emphasis id="B9231F57F509FFF5FAF7F873FAAAF80C" box="[1289,1367,2007,2027]" italics="true" pageId="16" pageNumber="681">pallens</emphasis>
Horváth
</taxonomicName>
, macropterous male, dorsal habitus, specimen from Papua New Guinea, Gulf Prov., Sapoi River nr. Lakekamu, CL 7146 (Young Sohn illustration).
</paragraph>
</caption>
<paragraph id="8BE8C345F508FFF4FF8FFF53FD12FEA1" blockId="17.[113,753,189,557]" pageId="17" pageNumber="682">
<keyLead id="30A66070F508FFF4FF8FFF53FD12FEA1" pageId="17" pageNumber="682">
– Male lacking a projecting tab on abdominal ventrite IV; female subgenital plate longer, shape roughly triangular or elongate trapezoidal with elongate lateral hair tufts (
<figureCitation id="136CDFC0F508FFF4FDC2FE95FD70FEA1" box="[572,653,305,327]" captionStart="Figs" captionStartId="17.[113,157,1948,1970]" captionTargetBox="[109,757,1020,1917]" captionTargetId="figure-458@17.[97,769,1010,1932]" captionTargetPageId="17" captionText="Figs.41–46.Aphelocheirus (Aphelocheirus) femoratus D. Polhemus &amp; J. Polhemus, structural details, specimens from Malaysia, Pahang, Cameron Highlands. 41. Brachypterous male, dorsal habitus. 42. Macropterous male, dorsal habitus. 43. Male parameres and phallotheca. 44. Female subgenital plate. 45. Male hind femur, ventral view. 46. Proacetabula." figureDoi="http://doi.org/10.5281/zenodo.5352732" httpUri="https://zenodo.org/record/5352732/files/figure.png" pageId="17" pageNumber="682">Figs. 44</figureCitation>
,
<figureCitation id="136CDFC0F508FFF4FD67FE94FD4FFEA1" box="[665,690,304,326]" captionStart="Figs" captionStartId="18.[836,879,1948,1970]" captionTargetBox="[834,1477,1158,1911]" captionTargetId="figure-430@18.[811,1503,1147,1916]" captionTargetPageId="18" captionText="Figs.53–57.Aphelocheirus (Aphelocheirus) malayanus D.Polhemus &amp; J. Polhemus, structural details. 53. Brachypterous male, dorsal habitus, specimen from Malaysia, Perak, near Jeli. 54. Macropterous female, dorsal habitus, specimen from Malaysia, Pahang, Cameron Highlands. 55. Male parameres and phallotheca. 56. Proacetabula. 57. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5352736" httpUri="https://zenodo.org/record/5352736/files/figure.png" pageId="17" pageNumber="682">57</figureCitation>
)......3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A3DBE0F508FFF4FF8FFEEAFD12FDCA" pageId="17" pageNumber="682">
<paragraph id="8BE8C345F508FFF4FF8FFEEAFD12FE5D" blockId="17.[113,753,189,557]" pageId="17" pageNumber="682">
<keyLead id="30A66070F508FFF4FF8FFEEAFD12FE5D" pageId="17" pageNumber="682">
3. Male with dark, raised, ovate swelling on ventral face of hind femur (
<figureCitation id="136CDFC0F508FFF4FF21FECEFED4FE67" box="[223,297,362,384]" captionStart="Figs" captionStartId="17.[113,157,1948,1970]" captionTargetBox="[109,757,1020,1917]" captionTargetId="figure-458@17.[97,769,1010,1932]" captionTargetPageId="17" captionText="Figs.41–46.Aphelocheirus (Aphelocheirus) femoratus D. Polhemus &amp; J. Polhemus, structural details, specimens from Malaysia, Pahang, Cameron Highlands. 41. Brachypterous male, dorsal habitus. 42. Macropterous male, dorsal habitus. 43. Male parameres and phallotheca. 44. Female subgenital plate. 45. Male hind femur, ventral view. 46. Proacetabula." figureDoi="http://doi.org/10.5281/zenodo.5352732" httpUri="https://zenodo.org/record/5352732/files/figure.png" pageId="17" pageNumber="682">Fig. 45</figureCitation>
); body length in both sexes less than 8.0 mm; head produced ahead of eyes for 0.70 the dorsal length of an eye (
<figureCitation id="136CDFC0F508FFF4FF39FE00FEC0FE5D" box="[199,317,420,442]" captionStart="Figs" captionStartId="17.[113,157,1948,1970]" captionTargetBox="[109,757,1020,1917]" captionTargetId="figure-458@17.[97,769,1010,1932]" captionTargetPageId="17" captionText="Figs.41–46.Aphelocheirus (Aphelocheirus) femoratus D. Polhemus &amp; J. Polhemus, structural details, specimens from Malaysia, Pahang, Cameron Highlands. 41. Brachypterous male, dorsal habitus. 42. Macropterous male, dorsal habitus. 43. Male parameres and phallotheca. 44. Female subgenital plate. 45. Male hind femur, ventral view. 46. Proacetabula." figureDoi="http://doi.org/10.5281/zenodo.5352732" httpUri="https://zenodo.org/record/5352732/files/figure.png" pageId="17" pageNumber="682">Figs. 41, 42</figureCitation>
)...
<taxonomicName id="4C57B8C6F508FFF4FEA6FE01FD12FE5D" ID-CoL="7QYKV" authority="D. Polhemus &amp; J. Polhemus" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="1989" box="[344,751,420,442]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="682" phylum="Arthropoda" rank="species" species="femoratus">
<emphasis id="B9231F57F508FFF4FEA6FE01FE9AFE5E" box="[344,359,421,441]" italics="true" pageId="17" pageNumber="682">A</emphasis>
.
<emphasis id="B9231F57F508FFF4FE8AFE01FE29FE5E" box="[372,468,421,441]" italics="true" pageId="17" pageNumber="682">femoratus</emphasis>
D. Polhemus &amp; J. Polhemus
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE8C345F508FFF4FF8FFE65FD12FDCA" blockId="17.[113,753,189,557]" pageId="17" pageNumber="682">
<keyLead id="30A66070F508FFF4FF8FFE65FD12FDCA" pageId="17" pageNumber="682">
– Male lacking dark, raised, ovate swelling on ventral face of hind femur; body length in both sexes exceeding
<quantity id="4CAF6EA0F508FFF4FD90FE79FD44FE14" box="[622,697,477,499]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="17" pageNumber="682" unit="mm" value="8.5">8.5 mm</quantity>
; head produced ahead of eyes for 0.60 the dorsal length of an eye (
<figureCitation id="136CDFC0F508FFF4FF60FDB3FEE9FDCA" box="[158,276,535,557]" captionStart="Figs" captionStartId="18.[836,879,1948,1970]" captionTargetBox="[834,1477,1158,1911]" captionTargetId="figure-430@18.[811,1503,1147,1916]" captionTargetPageId="18" captionText="Figs.53–57.Aphelocheirus (Aphelocheirus) malayanus D.Polhemus &amp; J. Polhemus, structural details. 53. Brachypterous male, dorsal habitus, specimen from Malaysia, Perak, near Jeli. 54. Macropterous female, dorsal habitus, specimen from Malaysia, Pahang, Cameron Highlands. 55. Male parameres and phallotheca. 56. Proacetabula. 57. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5352736" httpUri="https://zenodo.org/record/5352736/files/figure.png" pageId="17" pageNumber="682">Figs. 53, 54</figureCitation>
) ........
<taxonomicName id="4C57B8C6F508FFF4FEAEFDBCFD12FDCA" ID-CoL="7QYMS" authority="D. Polhemus &amp; J. Polhemus" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="1989" box="[336,751,535,557]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="682" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="B9231F57F508FFF4FEAEFDBCFEA2FDCB" box="[336,351,536,556]" italics="true" pageId="17" pageNumber="682">A</emphasis>
.
<emphasis id="B9231F57F508FFF4FE92FDBCFE29FDCB" box="[364,468,536,556]" italics="true" pageId="17" pageNumber="682">malayanus</emphasis>
D. Polhemus &amp; J. Polhemus
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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