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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1186.105736" ID-Pensoft-Pub="1313-2970-1186-175" ID-Pensoft-UUID="F4B1AC951B1A5CDB9BAF35DA11360CDA" ID-ZooBank="AB71200E4EB640F9B981C78C0B3BD662" ModsDocID="1313-2970-1186-175" checkinTime="1702549276600" checkinUser="pensoft" docAuthor="Yang, Lan, Zhao, Fangyu, He, Qiaoqiao &amp; Yao, Zhiyuan" docDate="2023" docId="54A4991B1E4F5AC3A776572025532A9C" docLanguage="en" docName="ZooKeys 1186: 175-184" docOrigin="ZooKeys 1186" docPubDate="2023-12-13" docSource="http://dx.doi.org/10.3897/zookeys.1186.105736" docTitle="Belisana yuhaoi Yang &amp; Yao 2023, sp. nov." docType="treatment" docUuid="35D4BC52-53E7-47D4-9E54-2FC5291E33DC" docUuidSource="ZooBank" docVersion="1" id="F4B1AC951B1A5CDB9BAF35DA11360CDA" lastPageNumber="175" masterDocId="F4B1AC951B1A5CDB9BAF35DA11360CDA" masterDocTitle="A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China" masterLastPageNumber="184" masterPageNumber="175" pageNumber="175" updateTime="1702549276600" updateUser="pensoft">
<mods:mods id="ECAD6F6DA1730ECD281C154BE12FE191" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B047391BDF0A391B730A6E177E6A6296">
<mods:title id="D4DBA1B5DF00C64F7833076CAA4B110F">A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China</mods:title>
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<mods:name id="1A17D4EB940F1EF1DD71D2899F1EAB53" type="personal">
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<mods:namePart id="E6E29BA738705D8EC435C29F63A0EE24">Yang, Lan</mods:namePart>
<mods:nameIdentifier id="8D47298C7447199A92AA15684381DE92" type="ORCID">https://orcid.org/0000-0002-7754-9275</mods:nameIdentifier>
<mods:affiliation id="3BC698EA5977C3ABF1E5DE05157C7BD3">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:namePart id="7CA3C9D4EF3AC8BC1189F6B085DDA0E7">Zhao, Fangyu</mods:namePart>
<mods:nameIdentifier id="089033B57CB5F7DB24A6B46B0657D9D7" type="ORCID">https://orcid.org/0000-0002-1005-8471</mods:nameIdentifier>
<mods:affiliation id="963E251FEE1EEC8903EE30DF12A17827">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
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<mods:namePart id="4DE8E3149AC64BE83A66E60F8B3C40A4">He, Qiaoqiao</mods:namePart>
<mods:nameIdentifier id="EAAE968D1EFBD14832EF3CA120F611E1" type="ORCID">https://orcid.org/0000-0002-9381-7444</mods:nameIdentifier>
<mods:affiliation id="B11F066BCF747111CD683DC7F45D518F">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; Liaoning Key Laboratory of Evolution and Biodiversity, Shenyang 110034, Liaoning, China &amp; Liaoning Key Laboratory for Biological Evolution and Agricultural Ecology, Shenyang 110034, Liaoning, China</mods:affiliation>
<mods:nameIdentifier id="252FB77D2DD3FB939A3891B8EF2A2C7B" type="email">heqq@synu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="A08B95CB8BE363F5120F678925580A02">Yao, Zhiyuan</mods:namePart>
<mods:nameIdentifier id="07D6D91998C600A678F9D7D7A0FE378E" type="ORCID">https://orcid.org/0000-0002-1631-0949</mods:nameIdentifier>
<mods:affiliation id="848ACB87F6415530883BAF455798F2B4">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; Liaoning Key Laboratory of Evolution and Biodiversity, Shenyang 110034, Liaoning, China &amp; Liaoning Key Laboratory for Biological Evolution and Agricultural Ecology, Shenyang 110034, Liaoning, China</mods:affiliation>
<mods:nameIdentifier id="05D7AC32966B92560D7FD9D0975A7E8B" type="email">yaozy@synu.edu.cn</mods:nameIdentifier>
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<mods:typeOfResource id="FFA8C21599DB9BB0DD10A8C755F1A2AC">text</mods:typeOfResource>
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<mods:date id="BCFDF1031C9F5093491B6185DC388010">2023</mods:date>
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<mods:number id="3DC61CABEAA5499FF1A134B5286D4331">2023-12-13</mods:number>
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<mods:number id="B96F8696F6B28FB928233A8444C3834F">1186</mods:number>
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<treatment id="54A4991B1E4F5AC3A776572025532A9C" LSID="urn:lsid:zoobank.org:act:35D4BC52-53E7-47D4-9E54-2FC5291E33DC" httpUri="http://treatment.plazi.org/id/54A4991B1E4F5AC3A776572025532A9C" lastPageNumber="175" pageId="0" pageNumber="175">
<subSubSection id="A46C9216D8853640B6F6FC1284546A34" pageId="0" pageNumber="175" type="nomenclature">
<paragraph id="5F497A9DCECDE4247DEC5932B46F49A5" pageId="0" pageNumber="175">
<taxonomicName id="D51AD3C8E79799F3E9AE762F9DB23D77" LSID="https://zoobank.org/35D4BC52-53E7-47D4-9E54-2FC5291E33DC" authority="Yang &amp; Yao" authorityName="Yang &amp; Yao" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana yuhaoi" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="yuhaoi" status="sp. nov.">Belisana yuhaoi Yang &amp; Yao</taxonomicName>
<taxonomicNameLabel id="971704AE19CC47A378D41CD6D0CBCE38" pageId="0" pageNumber="175">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F2FA2478E090AA835F2414FBF1FA3C47" pageId="0" pageNumber="175" type="description">
<paragraph id="9E49029B05FD952EF0BD8104467387ED" pageId="0" pageNumber="175">
<figureCitation id="9CD375EB51B8D3DFAB318FF6CD7F6416" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">Figs 3</figureCitation>
<figureCitation id="CAAE1A4766DCC987F09913D542939A3D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FE1183AEB4ED4BC87650E7FB0A602476" pageId="0" pageNumber="175" type="type material">
<paragraph id="BF8512B013B39EF4826941D4E4574D7A" pageId="0" pageNumber="175">Type material.</paragraph>
<paragraph id="C8F3176224AEAC68A8D226D8D710E284" pageId="0" pageNumber="175">
Holotype: ♂ (SYNU-Ar00301), cave without a name (
<geoCoordinate id="586EDDAEEFD57A80412CF0C0641F1A62" degrees="27" direction="north" minutes="5.40" orientation="latitude" precision="9" value="27.09">27°5.40'N</geoCoordinate>
,
<geoCoordinate id="D3FE1B53CCAB7B1E80D70E2CEE7930B5" degrees="106" direction="east" minutes="30.00" orientation="longitude" precision="9" value="106.5">106°30.00'E</geoCoordinate>
, 1109 m), Liutong Town, Xiuwen County, Guiyang,
<emphasis id="1E82A418B5FB0892DE24EB7970783505" bold="true" pageId="0" pageNumber="175">Guizhou</emphasis>
,
<emphasis id="AB46A2AD0A3EEA8096171A44339DCDFB" bold="true" pageId="0" pageNumber="175">China</emphasis>
, 5 June 2022, H Yu &amp; Q Lu leg. Paratypes: 2♂ (SYNU-Ar00302, Ar00303) and 3♀ (SYNU-Ar00304-00306), same data as for the holotype. 1♀ (SYNU-Ar00307), Duobing Cave (
<geoCoordinate id="0949511386DFFD3E5564633AFC1C72FB" degrees="27" direction="north" minutes="6.00" orientation="latitude" precision="9" value="27.1">27°6.00'N</geoCoordinate>
,
<geoCoordinate id="D97C7A7AF32C75D56760BFA0B2AF3ABB" degrees="106" direction="east" minutes="30.00" orientation="longitude" precision="9" value="106.5">106°30.00'E</geoCoordinate>
, 1026 m), other data as for the holotype.
</paragraph>
</subSubSection>
<subSubSection id="B9C543CC967DCE10ACAF8DF502A6ABE0" pageId="0" pageNumber="175" type="etymology">
<paragraph id="E7D6714CE17401D9DA5F2B2FB1A405BF" pageId="0" pageNumber="175">Etymology.</paragraph>
<paragraph id="AB346CDF108BD486E47280C198625D6B" pageId="0" pageNumber="175">The specific name is a patronym in honor of the collector Hao Yu; noun (name) in genitive case.</paragraph>
</subSubSection>
<subSubSection id="6D242037BBB1130E9D5CAC69CB55CDC3" pageId="0" pageNumber="175" type="diagnosis">
<paragraph id="BA3F24D9E4EC2278F36F32FD10A88173" pageId="0" pageNumber="175">Diagnosis.</paragraph>
<paragraph id="6B92735172E6D281972A0DF5B6E97696" pageId="0" pageNumber="175">
The new species resembles
<taxonomicName id="EFC502E72E0583093602F3D6F7607F99" lsidName="B. galeiformis" pageId="0" pageNumber="175" rank="species" species="galeiformis">
<emphasis id="90ECFC06FF74BC32FEC98EB0EFC6F1CF" italics="true" pageId="0" pageNumber="175">B. galeiformis</emphasis>
</taxonomicName>
Zhang &amp; Peng, 2011 (
<bibRefCitation id="7A0D12876C41BB1B88729CD54001C24D" DOI="https://doi.org/10.11646/zootaxa.2989.1.2" author="Zhang, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="175" pagination="51 - 68" refId="B21" refString="Zhang, F, Peng, Y, 2011. Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from south China. Zootaxa 2989 (1): 51 - 68, DOI: https://doi.org/10.11646/zootaxa.2989.1.2" title="Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from south China." url="https://doi.org/10.11646/zootaxa.2989.1.2" volume="2989" year="2011">Zhang and Peng 2011</bibRefCitation>
: 52, fig. 1A-F) with similar male chelicerae and bulbal apophysis (Fig.
<figureCitation id="B69D40A7E88B071B5B7F311F887C6DFB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4C, D</figureCitation>
), but it can be distinguished by prolatero-ventral lamella of procursus nearly round (arrow 2 in Fig.
<figureCitation id="11DB94955AC8F8232603DAC3A01857BF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3C</figureCitation>
; nearly angular in
<taxonomicName id="F49300BE159440EB06C56CC8C9E6BAF3" lsidName="B. galeiformis" pageId="0" pageNumber="175" rank="species" species="galeiformis">
<emphasis id="CC705492C24003FFC4D40121A2B40EF6" italics="true" pageId="0" pageNumber="175">B. galeiformis</emphasis>
</taxonomicName>
), by distal membranous lamella of procursus laterally weak sclerotized (sclerotized part nearly half-round, arrow 3 in Fig.
<figureCitation id="170405BA059646460CFB68DBB00E4674" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3C</figureCitation>
; with triangular sclerite in
<taxonomicName id="D5D9E22604B0B8AF9678C29F1A74BC11" lsidName="B. galeiformis" pageId="0" pageNumber="175" rank="species" species="galeiformis">
<emphasis id="582D8C4E0D4F27FB3614F365A773F976" italics="true" pageId="0" pageNumber="175">B. galeiformis</emphasis>
</taxonomicName>
), by procursus with triangular retrolateral membranous flap (f in Fig.
<figureCitation id="D49BE8741F80C2F963A6443E7759D2F5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3D</figureCitation>
; flap large and half-round in
<taxonomicName id="CC4F067C5E98B08531E6D63727C46177" lsidName="B. galeiformis" pageId="0" pageNumber="175" rank="species" species="galeiformis">
<emphasis id="3C9F50CDCB69751179E9704ACAF7C8BE" italics="true" pageId="0" pageNumber="175">B. galeiformis</emphasis>
</taxonomicName>
), by epigynal plate nearly round, posteriorly strongly curved (Fig.
<figureCitation id="5892C27335D8BC0C6995D71D12650CEE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4A</figureCitation>
; hat-shaped, posteriorly straight in
<taxonomicName id="A599DF4D3BF21CE5DC432877C75EC5B0" lsidName="B. galeiformis" pageId="0" pageNumber="175" rank="species" species="galeiformis">
<emphasis id="825152D57A99629EF3AFDB690D39B9AB" italics="true" pageId="0" pageNumber="175">B. galeiformis</emphasis>
</taxonomicName>
), and by vulval pore plates nearly triangular (Fig.
<figureCitation id="76B8D7DF5900E391EA327E8394D7CD98" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4B</figureCitation>
; long elliptic in
<taxonomicName id="5EAD878CF3266D7E528C70EA0CD708B3" lsidName="B. galeiformis" pageId="0" pageNumber="175" rank="species" species="galeiformis">
<emphasis id="0A505EE08BDE2E090F228758353E64E3" italics="true" pageId="0" pageNumber="175">B. galeiformis</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="8B7261AC5D23CEA9363D7D3882E1B726" doi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175" start="Figure 3" startId="F3">
<paragraph id="97506D33D6519B26F81DF49DCBD0CE4C" pageId="0" pageNumber="175">
<emphasis id="BAF6B449975EF0C3F72CC9AD4D341794" bold="true" pageId="0" pageNumber="175">Figure 3.</emphasis>
<taxonomicName id="FB928047F057EFBC17D436B2680D479E" authorityName="Yang &amp; Yao" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana yuhaoi" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="yuhaoi">
<emphasis id="BFE68CA8684F1D8CF2A1538E5ED786B4" italics="true" pageId="0" pageNumber="175">Belisana yuhaoi</emphasis>
</taxonomicName>
sp. nov., holotype male
<emphasis id="0A6B70163FAF351815F737792A252FF1" bold="true" pageId="0" pageNumber="175">A, B</emphasis>
palp:
<emphasis id="F62C7A78EDB0272C3B442DEE3F4382B8" bold="true" pageId="0" pageNumber="175">A</emphasis>
prolateral view
<emphasis id="58DF738A6142400628AA7812909F7089" bold="true" pageId="0" pageNumber="175">B</emphasis>
retrolateral view
<emphasis id="39890D1EF488051D09FE0486FFCA1CAC" bold="true" pageId="0" pageNumber="175">C, D</emphasis>
distal part of procursus:
<emphasis id="B786D0377038BFCA20CCA0263C8D3485" bold="true" pageId="0" pageNumber="175">C</emphasis>
prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis
<emphasis id="2E6B098FC77B5F177B7ED2C9065FD427" bold="true" pageId="0" pageNumber="175">D</emphasis>
retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (
<emphasis id="0D0C76471682D9D5233C39D51434FC39" bold="true" pageId="0" pageNumber="175">A, B</emphasis>
); 0.02 mm (
<emphasis id="7A0CC51236BC9AA5E6712809FA04DAF1" bold="true" pageId="0" pageNumber="175">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="20EFF6D19FE162C1A261350E8635D2AB" doi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175" start="Figure 4" startId="F4">
<paragraph id="11907870EED351C0EB440E400AFAE023" pageId="0" pageNumber="175">
<emphasis id="C66D9D9C547D06B187E69B755382E3B2" bold="true" pageId="0" pageNumber="175">Figure 4.</emphasis>
<taxonomicName id="A8CFF6DC1C39D8AA2D260ED19602CA35" authorityName="Yang &amp; Yao" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana yuhaoi" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="yuhaoi">
<emphasis id="44757C8E4195250F5D62631E347F0A8C" italics="true" pageId="0" pageNumber="175">Belisana yuhaoi</emphasis>
</taxonomicName>
sp. nov., holotype male (
<emphasis id="CEDF8E64521BEF1F1AB22C388166314A" bold="true" pageId="0" pageNumber="175">C-F</emphasis>
) and paratype female (
<emphasis id="C94FA6179F5ED6F13752D03440ED6126" bold="true" pageId="0" pageNumber="175">A, B, G, H</emphasis>
)
<emphasis id="736A11C7834D91E35F00E7BFABA718F5" bold="true" pageId="0" pageNumber="175">A</emphasis>
epigyne, ventral view
<emphasis id="6FDAC62143DC599A768EE6864D5505C8" bold="true" pageId="0" pageNumber="175">B</emphasis>
vulva, dorsal view
<emphasis id="F9539F734E638B0C58C17B789E675950" bold="true" pageId="0" pageNumber="175">C</emphasis>
bulbal apophyses, prolateral view
<emphasis id="E5BF0818BCDBD7D4F2401807F5BF3E9B" bold="true" pageId="0" pageNumber="175">D</emphasis>
chelicerae, frontal view
<emphasis id="731E5A1B96CC7C850D0CD31A731232C3" bold="true" pageId="0" pageNumber="175">E-H</emphasis>
habitus:
<emphasis id="0336242F098B9F70E859FEF1EE24F904" bold="true" pageId="0" pageNumber="175">E, G</emphasis>
dorsal view
<emphasis id="B468AA35DEC2456BD6AABD5E767C9019" bold="true" pageId="0" pageNumber="175">F</emphasis>
lateral view
<emphasis id="7745F1FBAB579C8FA1771A1E1641C4BF" bold="true" pageId="0" pageNumber="175">H</emphasis>
ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (
<emphasis id="D9C16CFE8C2DB7015ED3B71F5A65BBB9" bold="true" pageId="0" pageNumber="175">A-D</emphasis>
); 0.20 mm (
<emphasis id="421ED837C962322ABACBE30F995313CD" bold="true" pageId="0" pageNumber="175">E-H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="BCDB3B5163F3B10A7378068110B20E7D" pageId="0" pageNumber="175" type="description">
<paragraph id="D25E08DA64F3827AEC09B074BA0C847D" pageId="0" pageNumber="175">Description.</paragraph>
<paragraph id="120D4135C0BE468848232F1F2557492E" pageId="0" pageNumber="175">
<emphasis id="2218226CC3D5E84B76778E3ABFB6069A" bold="true" pageId="0" pageNumber="175">Male</emphasis>
(holotype, SYNU-Ar00301): total length 1.93 (2.03 with clypeus), prosoma 0.74 long, 0.78 wide, opisthosoma 1.19 long, 0.96 wide. Legs I and IV missing, femur II: 3.92 (other segments missing), leg III: 10.18 (2.97, 0.33, 2.48, 3.56, 0.84). Eye interdistances and diameters: PME-PME 0.13, PME 0.10, PME-ALE 0.04, AME absent. Sternum width/length: 0.65/0.62. Habitus as in Fig.
<figureCitation id="2C2A5F324A3ECBD8EE680839ABF889AE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4E, F</figureCitation>
. Dorsal shield of prosoma yellowish, with large, brown radiating marks; ocular area and clypeus yellowish, with brown marks; sternum yellowish, with triangular posterior brown marks. Legs whitish, without darker rings. Opisthosoma yellowish, with dorsal and lateral brown spots. Ocular area not elevated. Thoracic furrow absent. Clypeus unmodified. Chelicerae as in Fig.
<figureCitation id="70B50F9C75B0A22BAA09741EB993ECC4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4D</figureCitation>
, with a pair of proximo-lateral apophyses and a pair of curved distal apophyses (distance between tips of distal apophyses: 0.21). Palp as in Fig.
<figureCitation id="A3A98E7131D1EF3C12F2FDE816400CB3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3A, B</figureCitation>
; trochanter with short retrolatero-ventral apophysis; femur with small retrolatero-proximal protrusion; procursus simple proximally but complex distally, with prolatero-subdistal sclerite (arrow 1 in Fig.
<figureCitation id="76AA3E1C994FF0862D9383BE0D633AE5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3C</figureCitation>
), sclerotized prolatero-ventral lamella (arrow 2 in Fig.
<figureCitation id="5E2AC09439A93CC14C1A465587E52D3D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3C</figureCitation>
), distal membranous lamella (arrow 3 in Fig.
<figureCitation id="2FFE85C546DF279AF70C373A062EC2CD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3C</figureCitation>
), curved distal spine (arrow 4 in Fig.
<figureCitation id="B342B45D0B66AE8F871B13F87D68529A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3C</figureCitation>
), sclerotized dorsal apophysis (arrow 5 in Fig.
<figureCitation id="9AAC58F9441E765E20EDA94BD836DFF3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3C</figureCitation>
), and angular retrolateral membranous flap (f in Fig.
<figureCitation id="ABA77815A31C2A9760D20DD0DA84BE1D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana yuhaoi sp. nov., holotype male A, B palp: A prolateral view B retrolateral view C, D distal part of procursus: C prolateral view, arrow 1 indicates prolatero-subdistal sclerite, arrow 2 indicates sclerotized prolatero-ventral lamella, arrow 3 indicates distal membranous lamella, arrow 4 indicates curved distal spine, arrow 5 indicates sclerotized dorsal apophysis D retrolateral view. Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1186.105736.figure3" httpUri="https://binary.pensoft.net/fig/952211" pageId="0" pageNumber="175">3D</figureCitation>
); bulb (Fig.
<figureCitation id="05CF174C6785633F099B5FE6CF23E357" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4C</figureCitation>
) with hooked apophysis and simple embolus.
</paragraph>
<paragraph id="97A892B3D3D80EA8771896AFB5506974" pageId="0" pageNumber="175">
<emphasis id="CEA99B704F5C3FCC69EA42669B5F808F" bold="true" pageId="0" pageNumber="175">Female</emphasis>
(paratype, SYNU-Ar00304): similar to male, habitus as in Fig.
<figureCitation id="79FB0CCE2A8E2BECFFAEAC7C731C66A1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4G, H</figureCitation>
. Total length 2.03 (2.13 with clypeus), prosoma 0.83 long, 0.91 wide, opisthosoma 1.20 long, 1.04 wide; tibia I: 4.02; tibia I L/d: 50. Eye interdistances and diameters: PME-PME 0.12, PME 0.08, PME-ALE 0.03, AME absent. Sternum width/length: 0.58/0.51. Epigyne (Fig.
<figureCitation id="A87B5D6951CA143A4FB3A42EF3ADCCEE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4A</figureCitation>
) simple and flat, with brown marks and a pair of posterior pockets, 0.17 apart (ep in Fig.
<figureCitation id="03E3D24CB485D2C131BB68810BFA7299" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4A, B</figureCitation>
). Vulva (Fig.
<figureCitation id="CC32C1FCCAD7C3DC297D8CDEFFEBA1D5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana yuhaoi sp. nov., holotype male (C-F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulbal apophyses, prolateral view D chelicerae, frontal view E-H habitus: E, G dorsal view F lateral view H ventral view. Abbreviations: b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A-D); 0.20 mm (E-H)." figureDoi="10.3897/zookeys.1186.105736.figure4" httpUri="https://binary.pensoft.net/fig/952212" pageId="0" pageNumber="175">4B</figureCitation>
) with ridge-shaped anterior arch and a pair of nearly triangular pore plates.
</paragraph>
</subSubSection>
<subSubSection id="1FA625DC6E2ECFFB4EE369BA89FFAB83" pageId="0" pageNumber="175" type="variation">
<paragraph id="ACEA706B2D5F6B70503CA6FF420D732A" pageId="0" pageNumber="175">Variation.</paragraph>
<paragraph id="46541F315F787C8B58E9B22719369BA3" pageId="0" pageNumber="175">In one male paratype (SYNU-Ar00302), leg I: 23.28 (6.02, 0.39, 5.71, 9.36, 1.80); tibia I L/d: 60. Retrolateral trichobothrium of tibia I at 6% proximally; legs with short vertical setae on metatarsi, without spines and curved setae; tarsus I with 22 distinct pseudosegments. Tibia I in another male paratype (SYNU-Ar00303): 5.38. Tibia I in the other two female paratypes (SYNU-Ar00305, Ar00306): 3.28, 3.75 (leg I missing in SYNU-Ar00307).</paragraph>
</subSubSection>
<subSubSection id="0E3FC7737BFE3A55BC227E3CF24EEAEB" pageId="0" pageNumber="175" type="habitat">
<paragraph id="1DEB6CFBAA8D4087C0F2C0283D4D09E0" pageId="0" pageNumber="175">Habitat.</paragraph>
<paragraph id="9977CA880EEB47ADD184F61A038EDAF0" pageId="0" pageNumber="175">The species was found inside cave.</paragraph>
</subSubSection>
<subSubSection id="1CC8D3B112122A939D51ECD401EA40F6" pageId="0" pageNumber="175" type="distribution">
<paragraph id="4E82205DCBCA245572D7223544EC55C8" pageId="0" pageNumber="175">Distribution.</paragraph>
<paragraph id="2F6B2210BBC97CBB32E958583AC2BE41" pageId="0" pageNumber="175">
China (Xiuwen County in Guizhou; type locality, Fig.
<figureCitation id="6DC6FBAC34732ACBC4B9E9045590FFBB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Pholcidae treated in this paper 1 Belisana yuhaoi sp. nov. 2 Leptopholcus tanikawai 3 Pholcus spilis 4 Spermophora senoculata." figureDoi="10.3897/zookeys.1186.105736.figure1" httpUri="https://binary.pensoft.net/fig/952209" pageId="0" pageNumber="175">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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