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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1136.72744" ID-Pensoft-Pub="1313-2970-1136-1" ID-Pensoft-UUID="5D0E6613B1D05E4293ABEE4BFD1CA18A" ID-ZooBank="11C6BBFB339A4672AE662CE2B1E6321E" ModsDocID="1313-2970-1136-1" checkinTime="1671287389350" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2022" docId="504C87B041C55B09B60FF95FA9368BBA" docLanguage="en" docName="ZooKeys 1136: 1-56" docOrigin="ZooKeys 1136" docPubDate="2022-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1136.72744" docTitle="Desmopachria impunctata Miller 2022, sp. nov." docType="treatment" docUuid="8248D6E7-C6F1-43F0-B298-704E02A9F4D4" docUuidSource="ZooBank" docVersion="2" id="5D0E6613B1D05E4293ABEE4BFD1CA18A" lastPageNumber="1" masterDocId="5D0E6613B1D05E4293ABEE4BFD1CA18A" masterDocTitle="Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="1" updateTime="1671287691396" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Kelly B.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
<mods:nameIdentifier type="email">kbmiller@unm.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1136</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>56</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1136.72744</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1136.72744</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1136-1</mods:identifier>
<mods:identifier type="ZooBank">11C6BBFB339A4672AE662CE2B1E6321E</mods:identifier>
<mods:identifier type="Pensoft-UUID">5D0E6613B1D05E4293ABEE4BFD1CA18A</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:8248D6E7-C6F1-43F0-B298-704E02A9F4D4" httpUri="http://treatment.plazi.org/id/504C87B041C55B09B60FF95FA9368BBA" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/8248D6E7-C6F1-43F0-B298-704E02A9F4D4" authority="Miller, 2022" authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria impunctata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="impunctata" status="sp. nov.">Desmopachria impunctata</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">Figures 60-62</figureCitation>
<figureCitation captionStart="Figures 79, 80" captionStartId="F9" captionText="Figures 79, 80. Desmopachria species distributions. 79 Desmopachria nitida species group and Desmopachria portmanni-aldessa species group 80 Desmopachria portmanni-aldessa species group." figureDoi="10.3897/zookeys.1136.72744.figures79-80" httpUri="https://binary.pensoft.net/fig/782461" pageId="0" pageNumber="1">, 79</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Suriname, Sipaliwini District, Raleighfallen Nature Reserve, Voltzberg Trail,
<geoCoordinate degrees="04" direction="north" minutes="40.910" orientation="latitude" precision="1" value="4.6818333">04°40.910'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="11.138" orientation="longitude" precision="1" value="-56.185635">56°11.138'W</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species includes medium-sized
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria</emphasis>
</taxonomicName>
specimens in this group (TL = 1.8-1.9 mm). The elytra are more finely and indistinctly punctate than many species and punctures are of only a single, fine size. Also, the elytra are only slightly iridescent. The male genitalia are most similar to
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria emarginata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria emarginata</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">54-56</figureCitation>
), but the apical emargination in the median lobe in
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria impunctata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="impunctata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria impunctata</emphasis>
</taxonomicName>
is relatively shallower and in lateral aspect the apex is distinctly hooked ventrad (Figs
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">60</figureCitation>
,
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">61</figureCitation>
). Also, specimens of
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria impunctata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="impunctata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria impunctata</emphasis>
</taxonomicName>
are smaller (TL = 1.8-1.9 mm) than
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria emarginata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria emarginata</emphasis>
</taxonomicName>
(TL = 2.0-2.2 mm).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Measurements.</emphasis>
TL = 1.8-1.9 mm, GW = 1.2-1.3 mm, PW = 1.0-1.1 mm, HW = 0.6-0.7 mm, EW = 0.4-0.5 mm, TL/GW = 1.5-1.6, HW/EW = 1.7-1.8. Body very broadly oval, laterally broadly curved, lateral margins slightly discontinuous between pronotum and elytron.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration.</emphasis>
Head orange to orange yellow. Pronotum evenly yellow, in most specimens paler in color than head or elytron. Elytron evenly brownish orange, iridescent, especially laterally and apically. Ventral surface of head, prosternum, head appendages, and pro- and mesolegs yellow, other ventral surfaces and metalegs orange.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sculpture and structure.</emphasis>
Head broad, short; anterior margin of clypeus broadly curved, flattened, finely margined with conspicuous, continuous narrow bead; surface of head shiny, extremely finely and sparsely punctate; eyes moderately large (HW/EW = 1.7-1.8); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, antennomere XI elongate, apically pointed. Pronotum short, lateral margins short, slightly curved with continuous marginal bead; surface shiny, extremely finely and sparsely punctate, posterior margin sinuate. Elytron broad, laterally broadly curved and rounded; surface shiny, extremely minutely punctate, punctures of even size, somewhat denser along elytral suture. Prosternum extremely short, longitudinally compressed, medially carinate; prosternal process in male very slender anteriorly, with low, indistinct medial tubercle, bifid apically with deep medial pit, in female slender anteriorly, with distinctive, small medial tubercle, apically short and broad, medially slightly carinate, apically acutely pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, impunctate medially, shallowly and minutely punctate laterally and posteromedially; metaventrite wings extremely slender. Metacoxa with medial portion short, &lt;1/2 length of metaventrite medially, metacoxal lines slightly sinuate, divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, very finely punctate. Metatrochanter very large, subequal to length of ventral margin of metafemur; legs otherwise not noticeably modified. Abdomen with surfaces shiny and smooth, very finely and sparsely punctate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia.</emphasis>
Male median bilaterally symmetrical, in lateral aspect elongate, evenly broad, evenly and broadly curved on both dorsal and ventral margins, apex slightly curved ventrad, pointed (Fig.
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">60</figureCitation>
); in ventral aspect moderately broad basally, apically expanded, lateral margins broadly rounded, apex broad, medially moderately deeply emarginate, apical branches each apically broadly triangular (Fig.
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">61</figureCitation>
). Lateral lobe very broad medially, irregularly and broadly narrowed to pointed apex, with series of long setae along apicodorsal margin (Fig.
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">62</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sexual dimorphism.</emphasis>
Male pro- and mesotarsomeres I-III slightly more broadly expanded and with ventral adhesive setae. Male and female prosternal processes different as in all
<taxonomicName baseAuthorityName="Clark" baseAuthorityYear="1862" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria portmanni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="portmanni">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria portmanni</emphasis>
</taxonomicName>
group species.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Variation.</emphasis>
There is variation in intensity and degree of coloration across specimens. Some specimens are more evenly colored, some have greater disparity between the pale color of the pronotum and the head and elytron. Other specific variation was not detected.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is named
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria impunctata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="impunctata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria impunctata</emphasis>
</taxonomicName>
, Latin for not punctate, for the relatively less punctate dorsal surface than in many species in the group.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from localities in Sipaliwini District, Suriname and
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
State, Venezuela (Fig.
<figureCitation captionStart="Figures 79, 80" captionStartId="F9" captionText="Figures 79, 80. Desmopachria species distributions. 79 Desmopachria nitida species group and Desmopachria portmanni-aldessa species group 80 Desmopachria portmanni-aldessa species group." figureDoi="10.3897/zookeys.1136.72744.figures79-80" httpUri="https://binary.pensoft.net/fig/782461" pageId="0" pageNumber="1">79</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype in NZCS, male labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="04" direction="north" minutes="40.910" orientation="latitude" precision="1" value="4.6818333">04°40.910'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="11.138" orientation="longitude" precision="1" value="-56.185635">56°11.138'W</geoCoordinate>
, 78 m Raleighfallen [sic] Nature Reserve Voltzberg trail; detrital pools along stream; leg. A. Short, C. McIntosh 30.vii.2012; SR12-0730-01B/ SEMC1113574 KUNHM-ENT [barcode label]/ HOLOTYPE
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria impunctata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="impunctata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria impunctata</emphasis>
</taxonomicName>
Miller, 2021 [red label with black line border].&quot; Paratypes, 232 specimens in MIZA, MSBA NCZS, and SEMC each with paratype label,
<normalizedToken originalValue="“…">&quot;...</normalizedToken>
/ PARATYPE
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria impunctata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="impunctata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria impunctata</emphasis>
</taxonomicName>
Miller, 2021 [blue label with black line border]&quot; and different barcode labels (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Barcode label data for Desmopachria species." httpUri="http://table.plazi.org/id/B5FF9C5283064B11CB6B8D1966FB9A48" pageId="0" pageNumber="1" tableUuid="B5FF9C5283064B11CB6B8D1966FB9A48">1</tableCitation>
); 23 labeled same as holotype; 24 in NZCS and SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.942" orientation="latitude" precision="1" value="3.8990333">3°53.942'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.849" orientation="longitude" precision="1" value="-56.180817">56°10.849'W</geoCoordinate>
, 733m CSNR: Tafelberg Summit, nr Caiman Creek Camp, pools in forest; leg. Short &amp; Bloom 19.viii.2013; SR13-0819-05B/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 15 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.942" orientation="latitude" precision="1" value="3.8990333">3°53.942'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.849" orientation="longitude" precision="1" value="-56.180817">56°10.849'W</geoCoordinate>
, 733m CSNR: Tafelberg Summit, nr Caiman Creek Camp, forest detrital pools; leg. Short &amp; Bloom 19.viii.2013; SR13-0819-05C/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 11 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.942" orientation="latitude" precision="1" value="3.8990333">3°53.942'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.849" orientation="longitude" precision="1" value="-56.180817">56°10.849'W</geoCoordinate>
, 733m CSNR: Tafelberg Summit, nr Caiman Creek Camp, small streams with lots of plants &amp; leaf litter; leg. Short &amp; Bloom 18.viii.2013; SR13-0818-03A/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 1 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.359" orientation="latitude" precision="1" value="3.8893166">3°53.359'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.052" orientation="longitude" precision="1" value="-56.167534">56°10.052'W</geoCoordinate>
, 879m CSNR: Tafelberg Summit, near South Rim, pool in rock leg. Short &amp; Bloom; 20.viii.2013 SR13-0820-01C/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 5 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="04" direction="north" minutes="40.910" orientation="latitude" precision="1" value="4.6818333">04°40.910'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="11.138" orientation="longitude" precision="1" value="-56.185635">56°11.138'W</geoCoordinate>
, 78 m Raleighfallen [sic] Nature Reserve Voltzberg trail; margin of stream leg. C. Maier, V. Kadosoe 30.vii.2012; sr12-0730-01A/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 69 in NZCA and SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.942" orientation="latitude" precision="1" value="3.8990333">3°53.942'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.849" orientation="longitude" precision="1" value="-56.180817">56°10.849'W</geoCoordinate>
, 733m CSNR: Tafelberg Summit, nr. Caiman Crk. Camp; dipnetting by stream margins; leg. Bloom 20.viii.2013, SR13-0820-02A/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 3 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.942" orientation="latitude" precision="1" value="3.8990333">3°53.942'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.849" orientation="longitude" precision="1" value="-56.180817">56°10.849'W</geoCoordinate>
, 733m CSNR: Tafelberg Summit, near Caiman Creek Camp, forest detrital pools; leg. Short &amp; Bloom 19.viii.2013; SR13-0819-05C/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 69 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.942" orientation="latitude" precision="1" value="3.8990333">3°53.942'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.849" orientation="longitude" precision="1" value="-56.180817">56°10.849'W</geoCoordinate>
, 733m CSNR: Tafelberg Summit, nr Caiman Creek Camp, pools in forest; leg. Short &amp; Bloom 19.viii.2013; SR13-0819-05B/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 3 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.942" orientation="latitude" precision="1" value="3.8990333">3°53.942'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.849" orientation="longitude" precision="1" value="-56.180817">56°10.849'W</geoCoordinate>
, 733m CSNR: Tafelberg Summit, near Caiman Creek Camp, pooled up detrital forest stream leg. Short &amp; Bloom; 19.viii.2013 SR13-0819-05A/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 4 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="55.600" orientation="latitude" precision="1" value="3.9266667">3°55.600'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="11.300" orientation="longitude" precision="1" value="-56.18833">56°11.300'W</geoCoordinate>
, 600m CSNR: Tafelberg Summit, nr Augustus Creek Camp, pond on trail into Arrowhead basin leg. Short &amp; Bloom; 16.viii.2013 SR13-0816-02A/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 1 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="55.600" orientation="latitude" precision="1" value="3.9266667">3°55.600'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="11.300" orientation="longitude" precision="1" value="-56.18833">56°11.300'W</geoCoordinate>
, 600m CSNR: Tafelberg Summit, nr Augustus Creek Camp, pools &amp; creeks on trail into Arrowhead basin; leg. Short &amp; Bloom 17.viii.2013; SR13-0817-01A/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 3 in SEMC labeled, &quot;SURINAME: Sipaliwini District
<geoCoordinate degrees="3" direction="north" minutes="53.942" orientation="latitude" precision="1" value="3.8990333">3°53.942'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="west" minutes="10.849" orientation="longitude" precision="1" value="-56.180817">56°10.849'W</geoCoordinate>
, 733m CSNR: Tafelberg Summit, near Caiman Creek Camp, stream margins: legs. Short &amp; Bloom 18.viii.2013; SR13-0818-01A/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
; 1 in MIZA labeled, &quot;VENEZUELA:
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
State
<geoCoordinate degrees="6" direction="north" minutes="35.617" orientation="latitude" precision="1" value="6.5936165">6°35.617'N</geoCoordinate>
,
<geoCoordinate degrees="66" direction="west" minutes="49.238" orientation="longitude" precision="1" value="-66.82063">66°49.238'W</geoCoordinate>
, 80m Los Pijiguaos: outcrop/morichal 12.i.2009; leg Miller &amp; Short VZ09-0112-01C: detrial [sic] pools/
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
</paragraph>
</subSubSection>
</treatment>
</document>
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