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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.690.13751" ID-GBIF-Dataset="356a4c29-470b-4ca5-8503-76191b4fef6f" ID-PMC="PMC5672587" ID-Pensoft-Pub="1313-2970-690-1" ID-PubMed="29118595" ID-ZBK="C1B8D7C059E54C3A944F69F4FDE96B20" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-690-1" ModsDocOrigin="ZooKeys 690" ModsDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" checkinTime="1502920743207" checkinUser="pensoft" docAuthor="Shpeley, Danny, Hunting, Wesley &amp; Ball, George E." docDate="2017" docId="23E824BC79ABB09D74928424D2A6EE90" docLanguage="en" docName="ZooKeys 690: 1-195" docOrigin="ZooKeys 690" docSource="http://dx.doi.org/10.3897/zookeys.690.13751" docTitle="Selenophorus fabricii Shpeley, Hunting &amp; Ball, 2017, sp. n." docType="treatment" docUuid="EFCD958C-8523-4164-8112-0799CDFE173E" docUuidSource="ZooBank" docVersion="6" lastPageNumber="64" masterDocId="FE026465FF86843AE44A7B6EFF8DC422" masterDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" masterLastPageNumber="195" masterPageNumber="1" pageNumber="61" updateTime="1668164690231" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shpeley, Danny</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunting, Wesley</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ball, George E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>690</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>195</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.690.13751</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.690.13751</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-690-1</mods:identifier>
<mods:identifier type="ZBK">C1B8D7C059E54C3A944F69F4FDE96B20</mods:identifier>
<mods:identifier type="ZooBank">C1B8D7C059E54C3A944F69F4FDE96B20</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="132785207" LSID="urn:lsid:zoobank.org:act:EFCD958C-8523-4164-8112-0799CDFE173E" httpUri="http://treatment.plazi.org/id/23E824BC79ABB09D74928424D2A6EE90" lastPageId="63" lastPageNumber="64" pageId="60" pageNumber="61">
<subSubSection pageId="60" pageNumber="61" type="nomenclature">
<paragraph pageId="60" pageNumber="61">
<taxonomicName LSID="http://zoobank.org/EFCD958C-8523-4164-8112-0799CDFE173E" class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus fabricii" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="fabricii">Selenophorus fabricii</taxonomicName>
<taxonomicNameLabel pageId="60" pageNumber="61">sp. n.</taxonomicNameLabel>
Figs 39A, 41A, 42
<normalizedToken originalValue="A–C">A-C</normalizedToken>
, 45
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="specific epithet">
<paragraph pageId="61" pageNumber="62">
<pageBreakToken pageId="61" pageNumber="62" start="start">Specific</pageBreakToken>
epithet.
</paragraph>
<paragraph pageId="61" pageNumber="62">
A Latinized eponym, genitive case, based on the surname of Johann Christian Fabricius, who described
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus integer" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="integer">Carabus integer</taxonomicName>
, the species with which this one has been confused.
</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="63" type="type material">
<paragraph pageId="62" pageNumber="63">
<pageBreakToken pageId="62" pageNumber="63" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="62" pageNumber="63">Total of 283 specimens collected on the Greater Antillean island of Hispaniola, 156 males and 127 females. HOLOTYPE male, labelled: &quot;DOMINICAN REPUBLIC:/ Pedernales, Cabo Rojo/ 10 m.17-55N, 71-39W/ 26-27 September 1991&quot;; &quot;C. Young, S. Thompson,/ R.Davidson, J.Rawlins/ Coastal desert&quot; (CMNH). PARATYPES 282, sex and label data as follows. 53 males, 35 females, labelled same as holotype (CMNH). 5 males, 5 females, labelled same as holotype (UASM). 1 male, 1 female, labelled same as holotype (CASC). 50 males, 45 females, labelled: &quot;DOM.REP.:Prov.Pedernales/ Cabo Rojo, 0-10 m/ 10 SEP 1988, at light/ M. A. Ivie, TK. Philips/ &amp; K. A. Johnson colrs.&quot; (WIBF). 5 males, 5 females, labelled same as previous (UASM). 25 males, 25 females, labelled: &quot;DOM.REP:/ Prov.La Romana/ La Romana IX.18.1976/ E.Folch blacklight trap/ in sugar cane field&quot; (FSCA). 6 males, 1 female, labelled: &quot;DOM.REP:Dajabon Prov/ Rio Massacre, 40m.,/ Balneario Don Miguel/ 7 km sw. Dajabon/ 26 May 1973/ Don &amp; Mignon Davis (USNM). 4 males, 3 females, labelled: Rio Massacre, Balneario Don Miguel, 40 m, 7 km SW Dajabon, V.20.1973, D &amp; M Davis (USNM). 3 males, labelled: &quot;DOMINICANREP/ San Cristobal&quot;; &quot;8/9-VI-1969/ Flint &amp;Gomez&quot; (USNM). 1 male, 1 female labelled: &quot;DOMINICANREP/ San Cristobal&quot;; &quot;8-9-VI-1969/ Flint&amp;Gomez&quot; (USNM). 2 females, labelled: &quot;DOMINICANREP/ San Cristobal&quot;; &quot;8-9-VI-1969/ Flint &amp;Gomez&quot; (USNM). 2 females, labelled: &quot;DOMINICAN REP./ Los Hidalgos&quot;; &quot;4-5 VI 1969/ Flint&amp;Gomez&quot; (USNM). 2 females, labelled: &quot;DOMINICAN REP./ Los Hidalgos/ 4-5-June 1969/ Flint &amp; Gomez&quot; (USNM). 1 male, labelled: &quot;DOMINICAN REP./ Jarabacoa/ 3-4 June 1969/ Flint&amp;Gomez&quot; (USNM). 1 male, labelled: &quot;DOMINICAN REP./ Cachon de la Rubia/ nr.Central Ozama/ 10 June 1969 Flint &amp; Gomez (USNM).</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="63" type="type locality">
<paragraph pageId="62" pageNumber="63">Type locality.</paragraph>
<paragraph pageId="62" pageNumber="63">Cabo Rojo, Pedernales province, Dominican Republic.</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="63" type="diagnosis">
<paragraph pageId="62" pageNumber="63">Diagnosis.</paragraph>
<paragraph pageId="62" pageNumber="63">This species is readily separated from the other members of the opalinus species group by the very wide striae in the preapical portion of the elytron, relative to the width of the striae on the elytral disc.</paragraph>
</subSubSection>
<subSubSection lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="descriptive notes">
<paragraph pageId="62" pageNumber="63">Descriptive notes.</paragraph>
<paragraph pageId="62" pageNumber="63">
Data for SBL in Table 1. Habitus as in Fig. 39A. Clypeus and labrum with anterior margin of each shallowly concave. Antennae and mouthparts testaceous to rufo-testaceous. Legs rufo-testaceous to dark brunneous, femur slightly darker than remainder of leg. Dorsal and ventral surfaces rufo-brunneous to nearly piceous. Elytra with moderate to brilliant iridescence, varying with angles to light source. Ventral surface with faint to moderate iridescence. Head, pronotum and elytra shiny, without microlines visible at 100
<normalizedToken originalValue="×">x</normalizedToken>
. Pronotum with posteriolateral angles rounded; posteriolateral impressions and laterally near the bead finely punctate, each puncture bearing a short, fine seta. Base of elytra, intervals 8 and 9 and apical portion of elytra with short, fine setae. Elytral striae impunctate, except the standard setigerous punctures in striae 2, 5 and 7. Elytral striae widened preapically, about as wide as adjacent interval, markedly wider than on elytral disc (Fig. 41A). Intervals with fine micro-punctures. Males with two terminal setae and females with four terminal setae near the posterior margin on sternum VII.
</paragraph>
<paragraph lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
Male genitalia. Fig. 42
<normalizedToken originalValue="A–C">A-C</normalizedToken>
. Apical portion of phallic median lobe moderately long, narrowly tapered, symmetrically broadly rounded in dorsal/ventral aspect, ex
<pageBreakToken pageId="63" pageNumber="64" start="start">treme</pageBreakToken>
apex curved ventrad; endophallus with one darkened microtrichial field, about medial, in right lateral aspect; without lamina; ostium anopic. Ventral surface of distal 1/3 of shaft with two sharp ridges to apex.
</paragraph>
<paragraph pageId="63" pageNumber="64">
Ovipositor and female reproductive tract. Very similar to those of
<taxonomicName lsidName="S. opalinus" pageId="63" pageNumber="64" rank="species" species="opalinus">S. opalinus</taxonomicName>
, Fig. 44B. For details, see this topic for
<taxonomicName lsidName="S. opalinus" pageId="63" pageNumber="64" rank="species" species="opalinus">S. opalinus</taxonomicName>
, below.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="distribution">
<paragraph pageId="63" pageNumber="64">Geographical distribution.</paragraph>
<paragraph pageId="63" pageNumber="64">Fig. 45. The known range of this species extends from Puerto Rico westward to Hispaniola, and then south-westward to Jamaica, the Caymans, the Swan Islands, and north-westward from Hispaniola to the Bahamas and the Key Islands off the coast of Florida.</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="chorological affinities and relationships">
<paragraph pageId="63" pageNumber="64">Chorological affinities and relationships.</paragraph>
<paragraph pageId="63" pageNumber="64">
Within the opalinus species group, the range of this species is overlapped by the ranges of
<taxonomicName lsidName="S. flavilabris" pageId="63" pageNumber="64" rank="species" species="flavilabris">S. flavilabris</taxonomicName>
(sensu lato),
<taxonomicName lsidName="S. integer" pageId="63" pageNumber="64" rank="species" species="integer">S. integer</taxonomicName>
,
<taxonomicName lsidName="S. opalinus" pageId="63" pageNumber="64" rank="species" species="opalinus">S. opalinus</taxonomicName>
, and
<taxonomicName lsidName="S. propinquus" pageId="63" pageNumber="64" rank="species" species="propinquus">S. propinquus</taxonomicName>
. Relationships of
<taxonomicName lsidName="S. fabricii" pageId="63" pageNumber="64" rank="species" species="fabricii">S. fabricii</taxonomicName>
are not postulated beyond species group membership.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="materials_examined">
<paragraph pageId="63" pageNumber="64">Material examined.</paragraph>
<paragraph pageId="63" pageNumber="64">In addition to type material, we have seen a total of 1,633 specimens (843 males, 790 females). See Appendix for details.</paragraph>
<caption pageId="63" pageNumber="64">
<paragraph pageId="63" pageNumber="64">
Figure 39. Habitus digital images of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus opalinus" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="opalinus">Selenophorus opalinus</taxonomicName>
species group, in part, dorsal aspect. A
<taxonomicName lsidName="S. fabricii" pageId="63" pageNumber="64" rank="species" species="fabricii">S. fabricii</taxonomicName>
sp. n. B
<taxonomicName lsidName="S. flavilabris" pageId="63" pageNumber="64" rank="subSpecies" species="flavilabris" subSpecies="cubanus">S. flavilabris cubanus</taxonomicName>
Darlington C
<taxonomicName lsidName="S. flavilabris" pageId="63" pageNumber="64" rank="subSpecies" species="flavilabris" subSpecies="flavilabris">S. flavilabris flavilabris</taxonomicName>
Dejean D
<taxonomicName lsidName="S. flavilabris" pageId="63" pageNumber="64" rank="subSpecies" species="flavilabris" subSpecies="ubancus">S. flavilabris ubancus</taxonomicName>
Ball &amp; Shpeley. Scale bars: A 10 mm;
<normalizedToken originalValue="B–D">B-D</normalizedToken>
5 mm.
</paragraph>
</caption>
<caption pageId="63" pageNumber="64">
<paragraph pageId="63" pageNumber="64">
Figure 40. Habitus digital images of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus opalinus" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="opalinus">Selenophorus opalinus</taxonomicName>
species group, in part, dorsal aspect. A
<taxonomicName lsidName="S. integer" pageId="63" pageNumber="64" rank="species" species="integer">S. integer</taxonomicName>
(Fabricius) B
<taxonomicName lsidName="S. opalinus" pageId="63" pageNumber="64" rank="species" species="opalinus">S. opalinus</taxonomicName>
LeConte C
<taxonomicName lsidName="S. propinquus" pageId="63" pageNumber="64" rank="species" species="propinquus">S. propinquus</taxonomicName>
Putzeys. Scale bars: A 10 mm; B, C 5 mm.
</paragraph>
</caption>
<caption pageId="63" pageNumber="64">
<paragraph pageId="63" pageNumber="64">
Figure 41. Digital images of apical portion of elytra of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">Selenophorus</taxonomicName>
species, tilted dorsal aspect. A
<taxonomicName lsidName="S. fabricii" pageId="63" pageNumber="64" rank="species" species="fabricii">S. fabricii</taxonomicName>
sp. n. B
<taxonomicName lsidName="S. integer" pageId="63" pageNumber="64" rank="species" species="integer">S. integer</taxonomicName>
(Fabricius). Scale bars 3 mm.
</paragraph>
</caption>
<caption pageId="63" pageNumber="64">
<paragraph pageId="63" pageNumber="64">
Figure 42. Digital images of male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus opalinus" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="opalinus">Selenophorus opalinus</taxonomicName>
species group, in part. A, D right lateral aspect B, E dorsal aspect C, F left lateral aspect.
<normalizedToken originalValue="A–CS">A-CS</normalizedToken>
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori fabricii" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="fabricii">fabricii</taxonomicName>
sp. n.
<normalizedToken originalValue="D–F">D-F</normalizedToken>
<taxonomicName lsidName="S. flavilabris" pageId="63" pageNumber="64" rank="subSpecies" species="flavilabris" subSpecies="ubancus">S. flavilabris ubancus</taxonomicName>
Ball &amp; Shpeley. Scale bars 1 mm.
</paragraph>
</caption>
<caption pageId="63" pageNumber="64">
<paragraph pageId="63" pageNumber="64">
Figure 43. Digital images of male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus opalinus" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="opalinus">Selenophorus opalinus</taxonomicName>
species group, in part. A, D, G right lateral aspect B, E, H dorsal aspect C, F, I left lateral aspect.
<normalizedToken originalValue="A–C">A-C</normalizedToken>
<taxonomicName lsidName="S. integer" pageId="63" pageNumber="64" rank="species" species="integer">S. integer</taxonomicName>
(Fabricius)
<normalizedToken originalValue="D–F">D-F</normalizedToken>
<taxonomicName lsidName="S. opalinus" pageId="63" pageNumber="64" rank="species" species="opalinus">S. opalinus</taxonomicName>
LeConte
<normalizedToken originalValue="G–I">G-I</normalizedToken>
<taxonomicName lsidName="S. propinquus" pageId="63" pageNumber="64" rank="species" species="propinquus">S. propinquus</taxonomicName>
Putzeys. Scale bars 1 mm.
</paragraph>
</caption>
<caption pageId="63" pageNumber="64">
<paragraph pageId="63" pageNumber="64">
Figure 44. Line drawings of female reproductive tract of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus opalinus" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="opalinus">Selenophorus opalinus</taxonomicName>
species group, in part, ventral aspect. A
<taxonomicName lsidName="S. flavilabris" pageId="63" pageNumber="64" rank="subSpecies" species="flavilabris" subSpecies="ubancus">S. flavilabris ubancus</taxonomicName>
Ball &amp; Shpeley B
<taxonomicName lsidName="S. opalinus" pageId="63" pageNumber="64" rank="species" species="opalinus">S. opalinus</taxonomicName>
LeConte. Legend: bc bursa copulatrix co common oviduct gc1 gonocoxite 1 gc2 gonocoxite 2 sp spermatheca spg spermathecal gland spgd spermathecal gland duct v valvifer. Scale bar 1 mm.
</paragraph>
</caption>
<caption pageId="63" pageNumber="64">
<paragraph pageId="63" pageNumber="64">
Figure 45. Map of West Indies showing known localities for species of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus opalinus" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="opalinus">Selenophorus opalinus</taxonomicName>
species group, in part.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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