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<document id="95353705077B6301271A699C421B66E6" ID-DOI="10.5852/ejt.2023.915.2379" ID-ISSN="2118-9773" ID-Zenodo-Dep="10410126" ID-ZooBank="78346F83-1177-4EAE-AFFD-D4DDCC1BFB83" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1703073859257" checkinUser="plazi" docAuthor="Balkenohl, Michael" docDate="2023" docId="4A6487ECCF2EFFD3FDC3FB01FCA12F61" docLanguage="en" docName="EJT.2023.915.1-57.pdf" docOrigin="European Journal of Taxonomy 915 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2379/10451" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sulciclivina sagittaria subsp. sagittaria" docType="treatment" docVersion="3" lastPageNumber="45" masterDocId="B65DFF94CF07FFFFFFC5FFC1FF8F2A5D" masterDocTitle="Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="42" updateTime="1704213011641" updateUser="julia" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="C8796B9C87E68F765638267281068859" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="58989FC06C97A0A496B83BA19BE5ABF5">Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini)</mods:title>
</mods:titleInfo>
<mods:name id="DA92A0696AEBA3B6240C66BEC2AECD96" type="personal">
<mods:role id="0306425CCF445AAD6B1C66AB4F926713">
<mods:roleTerm id="FD9A5E5BF6239A66F5504AA64212A80D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BB2890AB662D2ACEDBC99692C1B27CE2">Balkenohl, Michael</mods:namePart>
<mods:nameIdentifier id="C789FA1D1C58B0CFE1C136232C1A39C3" type="LSID">urn:lsid:zoobank.org:author:D2154C49-8334-405E-B02D-DD265A0B3BF6</mods:nameIdentifier>
<mods:affiliation id="DC3BC7AAF15F00D5E284CCC29BBA08CB">Ligusterweg 9, CH- 8906 Bonstetten, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="687BC2AA0066A11682B45CDF2ADBDBF7" type="email">Email: mike.balkenohl@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="079B872E50EC301598DC10CFF568250C">text</mods:typeOfResource>
<mods:relatedItem id="66AC81076876B92A8D68666821FBFED2" type="host">
<mods:titleInfo id="6AD37B81D282BDF3068F1DF808DAC436">
<mods:title id="81C862EC1FB7E0FB90CF145EE5FB4B2D">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="E670D2605769AB0598F59F48614A1A7A">
<mods:date id="4B802B6858DFD9BEF8DF19E09EA8FAA4">2023</mods:date>
<mods:detail id="CE98407D33E503C78C710F7E757833C7" type="pubDate">
<mods:number id="4E5F0E7712AB2353C063D91EE5ADA304">2023-12-20</mods:number>
</mods:detail>
<mods:detail id="3E639C6BE8F9C5E1D0E691A0DEA693C1" type="volume">
<mods:number id="DBECDC8ABF2C34B17A4CFC2EF24EF205">915</mods:number>
</mods:detail>
<mods:detail id="36F954031990E7073228573F9D4BFC89" type="issue">
<mods:number id="DB086302AF08F46D9B901E0C63401664">1</mods:number>
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<mods:start id="2F9226A601969E552E85F8CF77B0D343">1</mods:start>
<mods:end id="9A96A36DC93EB19F1D27E9AAE43909FB">57</mods:end>
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<mods:location id="9450DD061AE6831C58C2CE16C85E7AE8">
<mods:url id="F711422069A5F67522C81EC42E558C01">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2379/10451</mods:url>
</mods:location>
<mods:classification id="001C49D4E09E4F1F5C88FEA3D1EC4DCC">journal article</mods:classification>
<mods:identifier id="9FD8FE32E23E19E37A0DE478CA5CBEE8" type="DOI">10.5852/ejt.2023.915.2379</mods:identifier>
<mods:identifier id="EF5665313ED2C537FEA6062FE148FAF8" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="D9025684019875E45C2450F9061A0005" type="Zenodo-Dep">10410126</mods:identifier>
<mods:identifier id="2A93EFF0B5A71AA679EB5074333D3EC4" type="ZooBank">78346F83-1177-4EAE-AFFD-D4DDCC1BFB83</mods:identifier>
</mods:mods>
<treatment id="4A6487ECCF2EFFD3FDC3FB01FCA12F61" LSID="urn:lsid:plazi:treatment:4A6487ECCF2EFFD3FDC3FB01FCA12F61" httpUri="http://treatment.plazi.org/id/4A6487ECCF2EFFD3FDC3FB01FCA12F61" lastPageId="44" lastPageNumber="45" pageId="41" pageNumber="42">
<subSubSection id="8AD76571CF2EFFD6FDC3FB01FBA22E86" box="[518,1069,1216,1243]" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="C27236FACF2EFFD6FDC3FB01FBA22E86" blockId="41.[518,1069,1216,1279]" box="[518,1069,1216,1243]" pageId="41" pageNumber="42">
<heading id="993A8196CF2EFFD6FDC3FB01FBA22E86" box="[518,1069,1216,1243]" centered="true" fontSize="11" level="2" pageId="41" pageNumber="42" reason="2">
<taxonomicName id="05CD4D79CF2EFFD6FDC3FB01FBA22E86" authority="(Bates, 1892)" baseAuthorityName="Bates" baseAuthorityYear="1892" box="[518,1069,1216,1243]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="sagittaria" subSpecies="sagittaria">
<emphasis id="F0B9EAE8CF2EFFD6FDC3FB01FC072E86" bold="true" box="[518,904,1216,1243]" italics="true" pageId="41" pageNumber="42">Sulciclivina sagittaria sagittaria</emphasis>
(
<bibRefCitation id="A65C4B0BCF2EFFD6FC5DFB00FBAA2E86" author="Bates H. W." box="[920,1061,1217,1243]" pageId="41" pageNumber="42" pagination="267 - 428" refId="ref38229" refString="Bates H. W. 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. Annali del Museo Civico di Genova 12: 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF2EFFD6FD58FB25FC192EA2" box="[669,918,1252,1279]" pageId="41" pageNumber="42" type="description">
<paragraph id="C27236FACF2EFFD6FD58FB25FC192EA2" blockId="41.[518,1069,1216,1279]" box="[669,918,1252,1279]" pageId="41" pageNumber="42">
<figureCitation id="5AF62A7FCF2EFFD6FD58FB25FD682EA3" box="[669,743,1252,1278]" captionStart="Figs 1–3" captionStartId="2.[189,240,1879,1905]" captionTargetBox="[195,1393,947,1816]" captionTargetId="figure-245@2.[189,1398,843,1844]" captionTargetPageId="2" captionText="Figs 1–3.Sulciclivina Balkenohl, 2022, head in dorsal view, showing the specific terms used. 1. Sulciclivina attenuata attenuata (Herbst, 1806), ♀, lectotype (MFNB). 2. S. splendida sp. nov., ♂, holotype (NHMB). 3. S. sagittaria sagittaria (Bates, 1892), ♂, paralectotype (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.10410128" httpUri="https://zenodo.org/record/10410128/files/figure.png" pageId="41" pageNumber="42">Figs 3</figureCitation>
,
<figureCitation id="5AF62A7FCF2EFFD6FD30FB25FC9A2EA3" box="[757,789,1252,1278]" captionStart="Figs 20–23" captionStartId="43.[189,242,1928,1954]" captionTargetBox="[209,1340,1340,1816]" captionTargetId="figure-373@43.[189,1398,1308,1893]" captionTargetPageId="43" captionText="Figs 20–23. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 20. S. sagittaria sagittaria (Bates, 1892), ♂, paralectotype (NHMUK). 21. S. sagittaria singaporensis subsp. nov., ♀, holotype (NHMUK). 22. S. oculiangusta sp. nov., ♀, holotype (NHMUK). 23. S. andrewesi sp. nov., ♂, holotype (NHMB)." figureDoi="http://doi.org/10.5281/zenodo.10410140" httpUri="https://zenodo.org/record/10410140/files/figure.png" pageId="41" pageNumber="42">20</figureCitation>
, 34, 50, 53
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF2EFFD6FF78FAE7FAD62FB1" pageId="41" pageNumber="42" type="reference_group">
<paragraph id="C27236FACF2EFFD6FF78FAE7FCED2F1D" blockId="41.[189,866,1318,1345]" box="[189,866,1318,1345]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="E2DD11D4CF2EFFD6FF78FAE7FCED2F1D" box="[189,866,1318,1345]" pageId="41" pageNumber="42">
<treatmentCitation id="436C10EBCF2EFFD6FF78FAE7FDD52F1C" author="Bates H. W." box="[189,602,1318,1345]" page="275" pageId="41" pageNumber="42" year="1892">
<taxonomicName id="05CD4D79CF2EFFD6FF78FAE7FDD52F1C" ID-CoL="5ZCD6" authority="Bates, 1892: 275" authorityName="Bates H. E. Andrewes" authorityPageNumber="275" authorityYear="1892" box="[189,602,1318,1345]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="F0B9EAE8CF2EFFD6FF78FAE7FE022F1D" box="[189,397,1318,1344]" italics="true" pageId="41" pageNumber="42">Clivina sagittaria</emphasis>
<bibRefCitation id="A65C4B0BCF2EFFD6FE51FAE6FDD52F1C" author="Bates H. W." box="[404,602,1318,1345]" pageId="41" pageNumber="42" pagination="267 - 428" refId="ref38229" refString="Bates H. W. 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. Annali del Museo Civico di Genova 12: 267 - 428." type="journal article" year="1892">Bates, 1892: 275</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="C27236FACF2EFFD6FF78FAA9FD682F95" blockId="41.[189,1400,1383,1517]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="E2DD11D4CF2EFFD6FF78FAA9FD682F95" pageId="41" pageNumber="42">
<taxonomicName id="05CD4D79CF2EFFD6FF78FAA9FE1D2FDC" ID-CoL="5ZCD6" authorityName="Bates H. E. Andrewes" authorityYear="1892" box="[189,402,1384,1410]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="F0B9EAE8CF2EFFD6FF78FAA9FE1D2FDC" box="[189,402,1384,1410]" italics="true" pageId="41" pageNumber="42">Clivina sagittaria</emphasis>
</taxonomicName>
–
<treatmentCitation id="436C10EBCF2EFFD6FE71FAA9FDF92FDF" author="Csiki E." box="[436,630,1383,1410]" page="510" pageId="41" pageNumber="42" year="1927">
<bibRefCitation id="A65C4B0BCF2EFFD6FE71FAA9FDF92FDF" author="Csiki E." box="[436,630,1383,1410]" pageId="41" pageNumber="42" refId="ref38372" refString="Csiki E. 1927. Coleopterorum Catalogus auspiciis et auxilio. W. Junk editus a S. Schenkling. 91: Carabidae: Carabinae I. W. Junk, Berlin." type="book" year="1927">Csiki 1927: 510</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="436C10EBCF2EFFD6FCFEFAA9FBB92FDF" author="Andrewes H. E." box="[827,1078,1383,1410]" page="358" pageId="41" pageNumber="42" year="1929">
<bibRefCitation id="A65C4B0BCF2EFFD6FCFEFAA9FBB92FDF" author="Andrewes H. E." box="[827,1078,1383,1410]" pageId="41" pageNumber="42" refId="ref37824" refString="Andrewes H. E. 1929. The Fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. I., Carabinae. Taylor &amp; Francis, London." type="book" year="1929">Andrewes 1929: 358</bibRefCitation>
</treatmentCitation>
(redescription); 1930: 116 (catalogue). —
<treatmentCitation id="436C10EBCF2EFFD6FE60FA4AFD012FF8" author="Balkenohl M." box="[421,654,1419,1445]" page="16" pageId="41" pageNumber="42" year="2001">
<bibRefCitation id="A65C4B0BCF2EFFD6FE60FA4AFD012FF8" author="Balkenohl M." box="[421,654,1419,1445]" pageId="41" pageNumber="42" refId="ref37929" refString="Balkenohl M. 2001. Key and Catalogue of the Tribe Clivinini from the Oriental Realm, with Revisions of the Genera Thliboclivina KULT, and Trilophidius JEANNEL (Insecta, Coleoptera, Carabidae, Scarititae, Clivinini). Pensoft Series Faunistica 21. Pensoft, Sofia, Moscow." type="book" year="2001">Balkenohl 2001: 16</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="436C10EBCF2EFFD6FC8AFA4AFC7B2FFB" author="Kult K." box="[847,1012,1419,1446]" page="25" pageId="41" pageNumber="42" year="1951">
<bibRefCitation id="A65C4B0BCF2EFFD6FC8AFA4AFC7B2FFB" author="Kult K." box="[847,1012,1419,1446]" pageId="41" pageNumber="42" pagination="16 - 32" refId="ref38483" refString="Kult K. 1951. Revision of the genus Clivina, Latr., from Oriental region. (Col. Carabidae). Casopis Ceskoslovenske Spolecnosti Entomologicke 48: 16 - 32." type="journal article" year="1951">Kult 1951: 25</bibRefCitation>
</treatmentCitation>
(enumeration). —
<treatmentCitation id="436C10EBCF2EFFD6FB1DFA4DFE982F94" author="Lorenz W." page="134" pageId="41" pageNumber="42" year="2005">
<bibRefCitation id="A65C4B0BCF2EFFD6FB1DFA4DFE982F94" author="Lorenz W." pageId="41" pageNumber="42" refId="ref38542" refString="Lorenz W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae), Second Edition 2005. Printed by the author, Tutzing." type="book" year="2005">Lorenz 2005: 134</bibRefCitation>
</treatmentCitation>
(catalogue); 2022: 5ZCD6 (catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="C27236FACF2EFFD6FF78FA13FAD62FB1" blockId="41.[189,1400,1383,1517]" box="[189,1369,1490,1517]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="E2DD11D4CF2EFFD6FF78FA13FAD62FB1" box="[189,1369,1490,1517]" pageId="41" pageNumber="42">
<taxonomicName id="05CD4D79CF2EFFD6FF78FA13FE4E2FB1" authority="Balkenohl 2022" box="[189,449,1490,1516]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="F0B9EAE8CF2EFFD6FF78FA13FE4E2FB1" box="[189,449,1490,1516]" italics="true" pageId="41" pageNumber="42">Sulciclivina sagittaria</emphasis>
</taxonomicName>
–
<treatmentCitation id="436C10EBCF2EFFD6FE1BFA13FD592FB1" author="Balkenohl M." box="[478,726,1490,1516]" page="108" pageId="41" pageNumber="42" year="2022">
<bibRefCitation id="A65C4B0BCF2EFFD6FE1BFA13FD592FB1" author="Balkenohl M." box="[478,726,1490,1516]" pageId="41" pageNumber="42" pagination="99 - 126" refId="ref38100" refString="Balkenohl M. 2022. On the female gonocoxites of Asian, Arabian, and African Clivinini with realignment of the Thliboclivina-, Physoclivina-, and Eoclivina- species groups, and notes on the higher systematics of the tribe Clivinini Rafinesque, 1815 (Coleoptera, Carabidae). Zootaxa 5190 (1): 99 - 126. https: // doi. org / 10.11646 / zootaxa. 5190.1.4" type="journal article" year="2022">Balkenohl 2022: 108</bibRefCitation>
</treatmentCitation>
, fig. 46 (figures of gonocoxites and new combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF2EFFD6FF78F9D2FCC52C9A" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="C27236FACF2EFFD6FF78F9D2FEB92C70" blockId="41.[189,310,1555,1581]" box="[189,310,1555,1581]" pageId="41" pageNumber="42">
<heading id="993A8196CF2EFFD6FF78F9D2FEB92C70" bold="true" box="[189,310,1555,1581]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="F0B9EAE8CF2EFFD6FF78F9D2FEB92C70" bold="true" box="[189,310,1555,1581]" pageId="41" pageNumber="42">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF2EFFD6FF78F982FCC52C9A" blockId="41.[187,1399,1602,1735]" pageId="41" pageNumber="42">
A small-sized species with the pronotum as long as wide or slightly longer than wide and distinct clypeal wings. Mainly distinguished from the other members of the
<emphasis id="F0B9EAE8CF2EFFD6FCA3F9A6FC6F2CDD" box="[870,992,1639,1664]" italics="true" pageId="41" pageNumber="42">sagittaria-</emphasis>
species group by the clypeal wings, which are developed as sharp teeth. In addition, the lateral margin of the pronotum is more attenuated anteriorly as in the other members of the species group.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF2EFFD5FF78F92EFD032E4F" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="C27236FACF2EFFD6FF78F92EFEE32D55" blockId="41.[189,364,1774,1801]" box="[189,364,1774,1801]" pageId="41" pageNumber="42">
<heading id="993A8196CF2EFFD6FF78F92EFEE32D55" bold="true" box="[189,364,1774,1801]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="F0B9EAE8CF2EFFD6FF78F92EFEE32D55" bold="true" box="[189,364,1774,1801]" pageId="41" pageNumber="42">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="72A53CA7CF2EFFD6FF2FF8DCFE052DB4" collectionCode="NHMUK" collectorName="H. E. Andrewes" country="Myanmar" location="MYANMAR" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<paragraph id="C27236FACF2EFFD6FF2FF8DCFD062D6A" blockId="41.[188,1398,1821,2025]" box="[234,649,1821,1848]" pageId="41" pageNumber="42">
<emphasis id="F0B9EAE8CF2EFFD6FF2FF8DCFEE92D6A" bold="true" box="[234,358,1821,1847]" pageId="41" pageNumber="42">
<typeStatus id="1D768858CF2EFFD6FF2FF8DCFEE92D6A" box="[234,358,1821,1847]" pageId="41" pageNumber="42" type="lectotype">Lectotype</typeStatus>
</emphasis>
(by present designation)
</paragraph>
<paragraph id="C27236FACF2EFFD6FF78F880FE052DB4" blockId="41.[188,1398,1821,2025]" pageId="41" pageNumber="42">
<collectingCountry id="BADA766ACF2EFFD6FF78F880FED62D06" box="[189,345,1857,1883]" name="Myanmar" pageId="41" pageNumber="42">MYANMAR</collectingCountry>
•
<specimenCount id="D4CBFD73CF2EFFD6FEBDF880FE1E2D06" box="[376,401,1857,1883]" count="1" pageId="41" pageNumber="42" type="male">♂</specimenCount>
; with labels and data: white, black framed, printed and handwritten in black ink “Rangoon Birmania Fea
<date id="B673103ACF2EFFD6FE25F8A4FDCC2D23" box="[480,579,1892,1919]" pageId="41" pageNumber="42" value="1887-06">VI 1887</date>
” / white, handwritten in black ink “
<taxonomicName id="05CD4D79CF2EFFD6FC27F8A5FBF62D23" box="[994,1145,1892,1918]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sagittaria">Cl. sagittaria</taxonomicName>
es. Tip. Bat.” / circle, green framed “Co-type” / white, printed “Bought from Janson,1917.” / “
<collectorName id="6F38532CCF2EFFD6FBC5F846FB202DFC" box="[1024,1199,1927,1953]" pageId="41" pageNumber="42">H.E.Andrewes</collectorName>
Coll. B.M.1945- 97.” / red, black printed “LECTOTYPE
<taxonomicName id="05CD4D79CF2EFFD6FD4AF86AFB2E2D98" authority="(Bates, 1892)" baseAuthorityName="Bates" baseAuthorityYear="1892" box="[655,1185,1963,1989]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="sagittaria" subSpecies="sagittaria">
Sulciclivina sagittaria sagittaria (
<bibRefCitation id="A65C4B0BCF2EFFD6FBC8F86AFB162D98" author="Bates H. W." box="[1037,1177,1963,1989]" pageId="41" pageNumber="42" pagination="267 - 428" refId="ref38229" refString="Bates H. W. 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. Annali del Museo Civico di Genova 12: 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
)
</taxonomicName>
des. M.Balkenohl 2023”;
<collectionCode id="A4DCAE3FCF2EFFD6FED6F80EFE092DB4" box="[275,390,1999,2025]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="41" pageNumber="42" type="Museum">NHMUK</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="C27236FACF2DFFD5FF2FFEC8FE2D2B7E" blockId="42.[187,1400,265,697]" box="[234,418,265,291]" pageId="42" pageNumber="43">
<emphasis id="F0B9EAE8CF2DFFD5FF2FFEC8FE2D2B7E" bold="true" box="[234,418,265,291]" pageId="42" pageNumber="43">
<typeStatus id="1D768858CF2DFFD5FF2FFEC8FE2D2B7E" box="[234,418,265,291]" pageId="42" pageNumber="43" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="C27236FACF2DFFD5FF78FEEAFDA228E4" blockId="42.[187,1400,265,697]" pageId="42" pageNumber="43">
<materialsCitation id="72A53CA7CF2DFFD5FF78FEEAFAA22BF7" collectingDate="1887-06" collectionCode="NHMUK" collectorName="F. van Emden Bequest" country="Myanmar" location="Birmania Fea" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Rangoon" typeStatus="paralectotype">
<collectingCountry id="BADA766ACF2DFFD5FF78FEEAFED62B18" box="[189,345,299,325]" name="Myanmar" pageId="42" pageNumber="43">MYANMAR</collectingCountry>
•
<specimenCount id="D4CBFD73CF2DFFD5FEB0FEEBFE2B2B18" box="[373,420,298,325]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
; with labels and data: white, black framed, printed and handwritten in black ink “
<collectingRegion id="0009F818CF2DFFD5FF0DFE8DFEBA2B3B" box="[200,309,332,358]" country="Myanmar" name="Yangon" pageId="42" pageNumber="43">Rangoon</collectingRegion>
<location id="C7126021CF2DFFD5FEFBFE8DFE6F2B3A" LSID="urn:lsid:plazi:treatment:4A6487ECCF2EFFD3FDC3FB01FCA12F61:C7126021CF2DFFD5FEFBFE8DFE6F2B3A" box="[318,480,332,359]" country="Myanmar" name="Birmania Fea" pageId="42" pageNumber="43" stateProvince="Rangoon">Birmania Fea</location>
<date id="B673103ACF2DFFD5FE2DFE8CFDC22B3B" box="[488,589,332,359]" pageId="42" pageNumber="43" value="1887-06">
<collectingDate id="A637E9D2CF2DFFD5FE2DFE8CFDC22B3B" box="[488,589,332,359]" pageId="42" pageNumber="43" value="1887-06">VI 1887</collectingDate>
</date>
” / circle, green framed “Co-type” / white, handwritten in black ink and printed “
<taxonomicName id="05CD4D79CF2DFFD5FE96FEAFFC982BD5" authority="Bates H. E. Andrewes" authorityName="Bates H. E. Andrewes" authorityYear="1892" box="[339,791,366,393]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sagittaria">Clivina sagittaria Bates H.E.Andrewes</taxonomicName>
det.” / white, handwritten and black printed “Gesch. 8. 1938 von Andrewes” / white, black printed “F. van Emden Bequest B.M. 1960-129.”;
<collectionCode id="A4DCAE3FCF2DFFD5FB7BFE51FAA22BF7" box="[1214,1325,400,426]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="42" pageNumber="43" type="Museum">NHMUK</collectionCode>
</materialsCitation>
•
<materialsCitation id="72A53CA7CF2DFFD5FA86FE51FE85284D" collectingDate="1887-06" collectionCode="ETHZ" collectorName="Co-type" country="Myanmar" location="Birmania Fea" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Rangoon" typeStatus="paralectotype">
<specimenCount id="D4CBFD73CF2DFFD5FA86FE51FAFE2BF7" box="[1347,1393,400,426]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
; with labels and data: white, black framed, printed and handwritten in black ink “
<collectingRegion id="0009F818CF2DFFD5FBA0FE73FB5D2B91" box="[1125,1234,434,460]" country="Myanmar" name="Yangon" pageId="42" pageNumber="43">Rangoon</collectingRegion>
<location id="C7126021CF2DFFD5FB1DFE73FAF92B91" LSID="urn:lsid:plazi:treatment:4A6487ECCF2EFFD3FDC3FB01FCA12F61:C7126021CF2DFFD5FB1DFE73FAF92B91" box="[1240,1398,434,460]" country="Myanmar" name="Birmania Fea" pageId="42" pageNumber="43" stateProvince="Rangoon">Birmania Fea</location>
<date id="B673103ACF2DFFD5FF78FE15FEAD2BB3" box="[189,290,468,494]" pageId="42" pageNumber="43" value="1887-06">
<collectingDate id="A637E9D2CF2DFFD5FF78FE15FEAD2BB3" box="[189,290,468,494]" pageId="42" pageNumber="43" value="1887-06">VI 1887</collectingDate>
</date>
” / circle, green framed “Co-type” / beige, handwritten in black ink “Civina
<taxonomicName id="05CD4D79CF2DFFD5FB79FE12FAEB2BB3" authority="Bts." authorityName="Bates H. E. Andrewes" authorityYear="1892" box="[1212,1380,467,494]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sagittaria">sagittaria Bts.</taxonomicName>
”;
<collectionCode id="A4DCAE3FCF2DFFD5FF7AFE37FE85284D" box="[191,266,502,528]" collectionName="Switzerland, Zurich, Erdgenoessische Technische Hochschule-Zentrum" country="Switzerland" httpUri="http://grbio.org/cool/3kp9-a263" name="Eidgenoessische Technische Hochschule-Zentrum" pageId="42" pageNumber="43">ETHZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="72A53CA7CF2DFFD5FEE7FE34FB7D2809" collectingDate="1887-06" collectionCode="MFNB" collectorName="Zool. Mus. Berlin" country="Myanmar" determinerName="Balkenohl" location="Birmania Fea" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Rangoon" typeStatus="paralectotype">
<specimenCount id="D4CBFD73CF2DFFD5FEE7FE34FEDE284D" box="[290,337,501,528]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
; with labels and data: white, black framed, printed and handwritten in black ink “
<collectingRegion id="0009F818CF2DFFD5FACEFE34FAF72852" box="[1291,1400,501,527]" country="Myanmar" name="Yangon" pageId="42" pageNumber="43">Rangoon</collectingRegion>
<location id="C7126021CF2DFFD5FF78FDD6FED4286F" LSID="urn:lsid:plazi:treatment:4A6487ECCF2EFFD3FDC3FB01FCA12F61:C7126021CF2DFFD5FF78FDD6FED4286F" box="[189,347,535,562]" country="Myanmar" name="Birmania Fea" pageId="42" pageNumber="43" stateProvince="Rangoon">Birmania Fea</location>
<date id="B673103ACF2DFFD5FEA7FDD9FE4B286C" box="[354,452,535,562]" pageId="42" pageNumber="43" value="1887-06">
<collectingDate id="A637E9D2CF2DFFD5FEA7FDD9FE4B286C" box="[354,452,535,562]" pageId="42" pageNumber="43" value="1887-06">VI 1887</collectingDate>
</date>
” / white, handwritten in black ink
<taxonomicName id="05CD4D79CF2DFFD5FC9DFDD6FB94286C" authority="Bates" authorityName="Bates H. E. Andrewes" authorityYear="1892" box="[856,1051,535,561]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sagittaria">“sagittaria Bates</taxonomicName>
” / white, black printed “Zool. Mus. Berlin” / white, black printed “
<taxonomicName id="05CD4D79CF2DFFD5FDA0FDF8FC1F280E" authority="BATES" authorityName="Bates H. E. Andrewes" authorityYear="1892" box="[613,912,569,595]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sagittaria">Clivina sagittaria BATES</taxonomicName>
det.
<determinerName id="37AEDA3ECF2DFFD5FC05FDF8FBB1280E" box="[960,1086,569,595]" pageId="42" pageNumber="43">Balkenohl</determinerName>
, 1996”;
<collectionCode id="A4DCAE3FCF2DFFD5FB5AFDFBFB7D2809" box="[1183,1266,570,596]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="42" pageNumber="43">MFNB</collectionCode>
</materialsCitation>
•
<materialsCitation id="72A53CA7CF2DFFD5FACFFDF8FDA628E4" collectingDate="1887-06" collectionCode="RBINS" country="Myanmar" location="Birmania Fea" pageId="42" pageNumber="43" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rangoon" typeStatus="paralectotype">
<specimenCount id="D4CBFD73CF2DFFD5FACFFDF8FAB82809" box="[1290,1335,569,596]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
,
<specimenCount id="D4CBFD73CF2DFFD5FA86FDF8FAE02809" box="[1347,1391,569,596]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
; on the same pin with labels and data: white, black framed, printed and handwritten in black ink “
<collectingRegion id="0009F818CF2DFFD5FACEFD9AFAF82828" box="[1291,1399,603,629]" country="Myanmar" name="Yangon" pageId="42" pageNumber="43">Rangoon</collectingRegion>
<location id="C7126021CF2DFFD5FF7EFDBCFED428CA" LSID="urn:lsid:plazi:treatment:4A6487ECCF2EFFD3FDC3FB01FCA12F61:C7126021CF2DFFD5FF7EFDBCFED428CA" box="[187,347,637,663]" country="Myanmar" name="Birmania Fea" pageId="42" pageNumber="43" stateProvince="Rangoon">Birmania Fea</location>
<date id="B673103ACF2DFFD5FEA6FDBCFE4828CA" box="[355,455,637,663]" pageId="42" pageNumber="43" value="1887-06">
<collectingDate id="A637E9D2CF2DFFD5FEA6FDBCFE4828CA" box="[355,455,637,663]" pageId="42" pageNumber="43" value="1887-06">VI 1887</collectingDate>
</date>
” / white, handwritten in black ink “
<taxonomicName id="05CD4D79CF2DFFD5FCAAFDBCFB9228CA" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[879,1053,637,663]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sagittaria" status="sp. nov.">Cliv. sagittaria</taxonomicName>
<taxonomicNameLabel id="EB8A5793CF2DFFD5FBE0FDBCFBD228CA" box="[1061,1117,637,663]" pageId="42" pageNumber="43" rank="species">n.sp.</taxonomicNameLabel>
” / white, black framed, red printed “Syntype”;
<collectionCode id="A4DCAE3FCF2DFFD5FE0AFD5EFDA628E4" box="[463,553,671,697]" collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="42" pageNumber="43" type="Museum">RBINS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C27236FACF2DFFD5FF78FD1FFE2328A4" blockId="42.[189,428,734,761]" box="[189,428,734,761]" pageId="42" pageNumber="43">
<heading id="993A8196CF2DFFD5FF78FD1FFE2328A4" bold="true" box="[189,428,734,761]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="F0B9EAE8CF2DFFD5FF78FD1FFE2328A4" bold="true" box="[189,428,734,761]" pageId="42" pageNumber="43">Taxonomic remark</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF2DFFD5FF78FCCDFD032E4F" blockId="42.[189,1376,780,1043]" pageId="42" pageNumber="43">
For the selection of the
<typeStatus id="1D768858CF2DFFD5FE16FCCDFDB0297B" box="[467,575,780,806]" pageId="42" pageNumber="43" type="lectotype">lectotype</typeStatus>
out of the series of six available cotypes, two considerations were applied. On the one hand, a complete male specimen, in good condition and not rotten inside, was selected. In addition, this specimen is the only one bearing the additional label in Italian language by Leonardo Fea “
<taxonomicName id="05CD4D79CF2DFFD5FE52FCB0FDA129D6" authorityName="Balkenohl" authorityYear="2023" box="[407,558,881,907]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sagittaria">Cl. sagittaria</taxonomicName>
es. tip. Bat.”, which is an abbreviation of ‘esempio tipo Bates’ and means ‘type specimen of Bates’. The material of
<taxonomicName id="05CD4D79CF2DFFD5FD3BFC52FC0229F0" authorityName="Balkenohl" authorityYear="2023" box="[766,909,915,941]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="F0B9EAE8CF2DFFD5FD3BFC52FC0229F0" box="[766,909,915,941]" italics="true" pageId="42" pageNumber="43">S. sagittaria</emphasis>
</taxonomicName>
collected by L. Fea was distributed and even sold to several places (written information from M.V.L. Barclay,
<collectionCode id="A4DCAE3FCF2DFFD5FBEDFC74FB132992" box="[1064,1180,949,975]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="42" pageNumber="43" type="Museum">NHMUK</collectionCode>
). The type series including the
<typeStatus id="1D768858CF2DFFD5FE9BFC16FE4529AC" box="[350,458,983,1009]" pageId="42" pageNumber="43" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="05CD4D79CF2DFFD5FE34FC16FD7729AC" authority="(Bates)" baseAuthorityName="Bates" box="[497,760,983,1009]" class="Insecta" family="Carabidae" genus="Eoclivina" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="F0B9EAE8CF2DFFD5FE34FC16FD1629AC" box="[497,665,983,1009]" italics="true" pageId="42" pageNumber="43">S. bhamoensis</emphasis>
(Bates)
</taxonomicName>
, also collected by L. Fea in
<collectingCountry id="BADA766ACF2DFFD5FB85FC16FB3D29AC" box="[1088,1202,983,1009]" name="Myanmar" pageId="42" pageNumber="43">Myanmar</collectingCountry>
was deposited in
<collectionCode id="A4DCAE3FCF2DFFD5FF1EFC38FEC52E4E" box="[219,330,1017,1043]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="42" pageNumber="43" type="Museum">NHMUK</collectionCode>
where Bates often worked.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF2DFFD3FF78FBF9FD202E94" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43" type="description">
<paragraph id="C27236FACF2DFFD5FF78FBF9FEE42E0F" blockId="42.[189,363,1080,1106]" box="[189,363,1080,1106]" pageId="42" pageNumber="43">
<heading id="993A8196CF2DFFD5FF78FBF9FEE42E0F" bold="true" box="[189,363,1080,1106]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="F0B9EAE8CF2DFFD5FF78FBF9FEE42E0F" bold="true" box="[189,363,1080,1106]" pageId="42" pageNumber="43">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF2DFFD5FF78FBA7FB5D2E9E" blockId="42.[188,1399,1126,1220]" pageId="42" pageNumber="43">
MEASUREMENTS.
<typeStatus id="1D768858CF2DFFD5FE47FBA7FE732EDD" box="[386,508,1126,1152]" pageId="42" pageNumber="43" type="lectotype">Lectotype</typeStatus>
: body length
<quantity id="05359B1FCF2DFFD5FD50FBA7FD702EDD" box="[661,767,1126,1152]" metricMagnitude="-3" metricUnit="m" metricValue="4.87" pageId="42" pageNumber="43" unit="mm" value="4.87">4.87 mm</quantity>
, width
<quantity id="05359B1FCF2DFFD5FC97FBA7FC332EDD" box="[850,956,1126,1152]" metricMagnitude="-3" metricUnit="m" metricValue="1.34" pageId="42" pageNumber="43" unit="mm" value="1.34">1.34 mm</quantity>
; L/W of pronotum 1.08; L/W of elytra 1.97. All material: body length
<quantity id="05359B1FCF2DFFD5FDEAFB49FD542EFF" box="[559,731,1160,1186]" metricMagnitude="-3" metricUnit="m" metricValue="4.83" metricValueMax="4.98" metricValueMin="4.68" pageId="42" pageNumber="43" unit="mm" value="4.83" valueMax="4.98" valueMin="4.68">4.68–4.98 mm</quantity>
(ẍ: =
<quantity id="05359B1FCF2DFFD5FCEDFB49FC172EFF" box="[808,920,1160,1186]" metricMagnitude="-3" metricUnit="m" metricValue="4.83" pageId="42" pageNumber="43" unit="mm" value="4.83">4.83 mm</quantity>
*), width
<quantity id="05359B1FCF2DFFD5FBC0FB46FB3F2EFF" box="[1029,1200,1159,1186]" metricMagnitude="-3" metricUnit="m" metricValue="1.305" metricValueMax="1.36" metricValueMin="1.25" pageId="42" pageNumber="43" unit="mm" value="1.305" valueMax="1.36" valueMin="1.25">1.25–1.36 mm</quantity>
(ẍ: =
<quantity id="05359B1FCF2DFFD5FB39FB49FAD22EFF" box="[1276,1373,1160,1186]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="42" pageNumber="43" unit="mm" value="1.3">1.3 mm</quantity>
*), L/W of pronotum 1.03–1.11 (ẍ: = 1.08*), L/W of elytra 1.95–2.01 (ẍ: = 1.99*); (*n = 6).
</paragraph>
<paragraph id="C27236FACF2DFFD5FF78FB2BFE912F1A" blockId="42.[187,1398,1257,1351]" pageId="42" pageNumber="43">COLOUR. Glossy. Piceous. Mouthparts, elytron, intermediate and hind leg, tarsomeres of front leg fuscous, antenna and palpomeres more pale fuscous. Anterior part of supraantennal plate at the margin translucent-fuscous.</paragraph>
<paragraph id="C27236FACF2DFFD4FF78FAAFFB682B1B" blockId="42.[187,1399,1389,2025]" lastBlockId="43.[188,1398,265,327]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
HEAD. A quarter narrower than pronotum. Clypeus with central part distinctly bilobed anteriorly; lateral lobe developed as acute teeth, slightly more projecting than central lobes, clypeal wings well developed, sharp, not as far projecting anteriorly as lateral teeth of clypeus, distinctly separated from clypeus by notch; supraantennal plate laterally moderately convex, laterally nearly as far projecting as eye; supraantennal plate finely reflexed margined. Supraantennal plate overlapping eye anteriorly by about a quarter (dorsal view), with the supraorbital carina keeled posteriorly and extended up to mid-eye level, separated from wing by sharp notch. Clypeus transverse, separated from frons by sharp carina in form like an inverted nearly right-angled V. Frons moderately convex, with another but smaller carina like an inverted V, with supraorbital carina short, extending up to posterior eye-level. Clypeus smooth, area between V-like carinae smooth, frons with two to three longitudinal rugae at middle to posterior eye-level, clypeus and frons separated from supraantennal plate and supraorbital carina by deep and moderately wide furrow; with two supraorbital setigerous punctures at middle of the eye and end of gena-level. With some moderately sized punctures laterally at level of neck, punctures less dense or missing at middle. Eye small, encased, moderately convex in dorsal view, main part situated ventrally where it is more distinctly convex. Gena moderately convex, covering around a sixth of posterior eye in ventral view. Eye somewhat constricted due to the overlapping supraantennal plate and the genae. Antenna moderately long, reaching distinctly over middle of pronotum, antennomeres five to ten distinctly longer than wide (L/W around 1.23). Labrum straight anteriorly, with isodiametric-irregular reticulation, six setose. Mandibles sub-falcate slightly bent up dorsally.
<collectingRegion id="0009F818CF2DFFD5FE88F80EFE3F2DB4" box="[333,432,1999,2025]" country="Myanmar" name="Chin" pageId="42" pageNumber="43">Mentum</collectingRegion>
with isodiametric reticulation at middle, with lobe flattened, with few fine carinae on surface, shape of lobe asymmetric slightly longitudinal oval, with sharp median carina. Median tooth moderately wide, not as projecting anteriorly as lobe, rounded and hollowed out anteriorly.
</paragraph>
<paragraph id="C27236FACF2CFFD4FF78FEAFFAF82899" blockId="43.[187,1399,366,709]" pageId="43" pageNumber="44">PRONOTUM. Disk slightly convex in lateral and distinctly convex in frontal view. Slightly longer than wide, peltate. Reflexed lateral margin nearly smooth, straight in anterior half, widest behind middle; anterior angle distinct, slightly projecting, posterior angle indistinct; lateral channel moderately wide, crenulated, narrower before and behind posterior setigerous puncture, slightly curved upward at posterior setigerous puncture with small tooth (lateral view). Median line moderately deep, sharp, subcrenulated, complete. Anterior transverse line crenulated, slightly wider than median line, joining median line, joining declivity at anterior margin. Surface glossy, with micro-punctures, with paralateral impression basally formed by impressed punctures, with paramedian cloud-like longitudinal group of punctures of moderate size, with few transverse wrinkles laterally and basally. Base distinctly marked, channel two times as wide as lateral channel, width about a fifth of width of pronotum. Proepisternum moderately well visible in dorsal view.</paragraph>
<paragraph id="C27236FACF2CFFD4FF78FD2DFD372E1E" blockId="43.[188,1399,748,1091]" pageId="43" pageNumber="44">ELYTRON. Shape long-oval, indistinctly clavate. Disk flattened in anterior half in lateral view, distinctly and regularly convex in frontal view. Nearly two times longer than wide, with maximum width distinctly behind middle. Reflexed lateral margin smooth, setigerous punctures in lateral channel with long yellow setae, interrupted at middle by one puncture. Scutellar striole missing; setigerous tubercle at base of first stria, with distinct tubercle at base of third interval, basal declivity with isodiametric reticulation. Humeral tooth situated at base of fourth interval. All striae distinctly deepened, punctuate-striate, one to three free at base, four reaching humerus, one and two, three and four, and five and six joining apically. Intervals distinctly convex, more convex laterally. Setigerous punctures missing on intervals. Surface of all intervals glossy, interval seven and eight with indistinct reticulation at humerus, interval eight at apex subcarinate and with indistinct reticulation.</paragraph>
<paragraph id="C27236FACF2CFFD4FF78FBAAFC142E97" blockId="43.[189,1399,1130,1227]" pageId="43" pageNumber="44">
HIND WINGS. In the
<typeStatus id="1D768858CF2CFFD4FE62FBABFD992ED9" box="[423,534,1130,1156]" pageId="43" pageNumber="44" type="lectotype">lectotype</typeStatus>
, the alae are reduced to half of the length and width of the elytron looking like a longitudinal spatula. In
<specimenCount id="D4CBFD73CF2CFFD4FDDCFB4FFD462EFA" box="[537,713,1165,1192]" count="2" pageId="43" pageNumber="44" type="generic">two specimens</specimenCount>
, there are no alae visible and in another two the alae are as in the
<typeStatus id="1D768858CF2CFFD4FEC3FB71FEFA2E97" box="[262,373,1200,1226]" pageId="43" pageNumber="44" type="lectotype">lectotype</typeStatus>
. In
<specimenCount id="D4CBFD73CF2CFFD4FE65FB70FDCB2E97" box="[416,580,1200,1227]" count="1" pageId="43" pageNumber="44" type="generic">one specimen</specimenCount>
, the alae are fully developed.
</paragraph>
<caption id="96B26672CF2CFFD4FF78F849FAE62DB5" ID-DOI="http://doi.org/10.5281/zenodo.10410140" ID-Zenodo-Dep="10410140" httpUri="https://zenodo.org/record/10410140/files/figure.png" pageId="43" pageNumber="44" startId="43.[189,242,1928,1954]" targetBox="[209,1340,1340,1816]" targetPageId="43">
<paragraph id="C27236FACF2CFFD4FF78F849FAE62DB5" blockId="43.[189,1399,1928,2025]" pageId="43" pageNumber="44">
<emphasis id="F0B9EAE8CF2CFFD4FF78F849FEC02DFF" bold="true" box="[189,335,1928,1954]" pageId="43" pageNumber="44">Figs 20–23.</emphasis>
<taxonomicName id="05CD4D79CF2CFFD4FE9CF849FD312DFF" authority="Balkenohl, 2022" authorityName="Balkenohl" authorityYear="2022" box="[345,702,1928,1954]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="F0B9EAE8CF2CFFD4FE9CF849FE682DFF" box="[345,487,1928,1954]" italics="true" pageId="43" pageNumber="44">Sulciclivina</emphasis>
<bibRefCitation id="A65C4B0BCF2CFFD4FE37F849FD312DFF" author="Balkenohl M." box="[498,702,1928,1954]" pageId="43" pageNumber="44" pagination="99 - 126" refId="ref38100" refString="Balkenohl M. 2022. On the female gonocoxites of Asian, Arabian, and African Clivinini with realignment of the Thliboclivina-, Physoclivina-, and Eoclivina- species groups, and notes on the higher systematics of the tribe Clivinini Rafinesque, 1815 (Coleoptera, Carabidae). Zootaxa 5190 (1): 99 - 126. https: // doi. org / 10.11646 / zootaxa. 5190.1.4" type="journal article" year="2022">Balkenohl, 2022</bibRefCitation>
</taxonomicName>
, habitus, dorsal view. 20.
<taxonomicName id="05CD4D79CF2CFFD4FBC6F849FE8B2D98" authority="(Bates, 1892)" baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="sagittaria" subSpecies="sagittaria">
<emphasis id="F0B9EAE8CF2CFFD4FBC6F849FA972DFF" box="[1027,1304,1928,1954]" italics="true" pageId="43" pageNumber="44">S. sagittaria sagittaria</emphasis>
(
<bibRefCitation id="A65C4B0BCF2CFFD4FAEEF849FF732D98" author="Bates H. W." pageId="43" pageNumber="44" pagination="267 - 428" refId="ref38229" refString="Bates H. W. 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. Annali del Museo Civico di Genova 12: 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
)
</taxonomicName>
, ♂, paralectotype (NHMUK).
<emphasis id="F0B9EAE8CF2CFFD4FDA3F86AFD0C2D98" bold="true" box="[614,643,1963,1989]" pageId="43" pageNumber="44">21</emphasis>
.
<taxonomicName id="05CD4D79CF2CFFD4FD54F86AFC482D98" box="[657,967,1963,1989]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="sagittaria" subSpecies="singaporensis">
<emphasis id="F0B9EAE8CF2CFFD4FD54F86AFC482D98" box="[657,967,1963,1989]" italics="true" pageId="43" pageNumber="44">S. sagittaria singaporensis</emphasis>
</taxonomicName>
subsp. nov., ♀, holotype (NHMUK).
<emphasis id="F0B9EAE8CF2CFFD4FF78F80FFF552DB5" bold="true" box="[189,218,1998,2024]" pageId="43" pageNumber="44">22</emphasis>
.
<taxonomicName id="05CD4D79CF2CFFD4FF2CF80EFE112DB5" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[233,414,1998,2025]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="oculiangusta" status="sp. nov.">
<emphasis id="F0B9EAE8CF2CFFD4FF2CF80EFE112DB5" box="[233,414,1998,2025]" italics="true" pageId="43" pageNumber="44">S. oculiangusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB8A5793CF2CFFD4FE60F80EFD8D2DB4" box="[421,514,1999,2025]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype (NHMUK).
<emphasis id="F0B9EAE8CF2CFFD4FCF7F80FFCC02DB5" bold="true" box="[818,847,1998,2024]" pageId="43" pageNumber="44">23</emphasis>
.
<taxonomicName id="05CD4D79CF2CFFD4FC9BF80EFC7C2DB5" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[862,1011,1998,2025]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="andrewesi" status="sp. nov.">
<emphasis id="F0B9EAE8CF2CFFD4FC9BF80EFC7C2DB5" box="[862,1011,1998,2025]" italics="true" pageId="43" pageNumber="44">S. andrewesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB8A5793CF2CFFD4FC3FF80EFBD82DB4" box="[1018,1111,1999,2025]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (NHMB).
</paragraph>
</caption>
<paragraph id="C27236FACF2BFFD3FF78FEC8FC1C2B36" blockId="44.[187,1399,265,364]" pageId="44" pageNumber="45">LOWER SURFACE. Proepisternum covered with dense moderately sized punctures, laterally with some irregular reticulation and with dense transverse wrinkles.All sternites of abdomen covered with moderately large punctures, with slightly less density at middle at middle.</paragraph>
<paragraph id="C27236FACF2BFFD3FF78FE55FAF62B8E" blockId="44.[189,1401,404,467]" pageId="44" pageNumber="45">LEGS. Protibia covered with longitudinal reticulation, sulcate dorsally at base, with three spines of moderate length, apical spine distinctly arcuate, movable spur robust, in apical half moderately arcuate.</paragraph>
<paragraph id="C27236FACF2BFFD3FF78FE3AFBC428FB" blockId="44.[187,1399,506,678]" pageId="44" pageNumber="45">MALE GENITALIA (Fig. 34). Median lobe moderately slender in dorsal view, relatively short in comparison to body size, regularly arcuate at middle, with moderate dorsal bulge, apex with moderately long, flattened, slightly widened, and distorted spatula, with few microtrichia on the surface. Endophallus at middle with longitudinal group of bristles. Ventral paramere with somewhat widened velum-like apophysis, apex with two setae. Dorsal paramere robust, attenuated to apex, with three apical setae.</paragraph>
<paragraph id="C27236FACF2BFFD3FF78FD0FFBDD296D" blockId="44.[188,1399,718,817]" pageId="44" pageNumber="45">FEMALE GONOCOXITE AND EPIPLEURITE (Fig. 50). Gonocoxite conical-shaped, flattened dorsally and ventrally, with three nematiform setae positioned at the median side, with one preapical long nematiform seta. Epipleurite semi-rectangular, with conspicuously widened angulated rod.</paragraph>
<paragraph id="C27236FACF2BFFD3FF78FC98FEBC292E" blockId="44.[189,307,857,883]" box="[189,307,857,883]" pageId="44" pageNumber="45">
<heading id="993A8196CF2BFFD3FF78FC98FEBC292E" bold="true" box="[189,307,857,883]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="F0B9EAE8CF2BFFD3FF78FC98FEBC292E" bold="true" box="[189,307,857,883]" pageId="44" pageNumber="45">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF2BFFD3FF78FC49FBD42E0A" blockId="44.[188,1399,904,1112]" pageId="44" pageNumber="45">On the frons of the head, the posterior smaller carina is anteriorly not closed in some of the specimens. In about half of the specimens, there is a deep impression posterior to the smaller carina (not developed in the LT). On the pronotum, the lateral channel varies with the development of the crenulation. In addition, the number of punctures on the disk varies. In a third of the specimens, the lateral margin of the elytron is indistinctly crenulated behind the humerus, and in a few specimens stria one and two are not joining at apex respectively at one side, whereas the striae are joining at the other side.</paragraph>
<paragraph id="C27236FACF2BFFD3FF78FB41FE3F2EC7" blockId="44.[189,432,1152,1178]" box="[189,432,1152,1178]" pageId="44" pageNumber="45">
<heading id="993A8196CF2BFFD3FF78FB41FE3F2EC7" bold="true" box="[189,432,1152,1178]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="F0B9EAE8CF2BFFD3FF78FB41FE3F2EC7" bold="true" box="[189,432,1152,1178]" pageId="44" pageNumber="45">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF2BFFD3FF78FB6EFD202E94" blockId="44.[189,687,1199,1226]" box="[189,687,1199,1226]" pageId="44" pageNumber="45">Secondary sexual characters not observed.</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF2BFFD3FF78FB33FCA12F61" box="[189,814,1266,1340]" pageId="44" pageNumber="45" type="distribution">
<paragraph id="C27236FACF2BFFD3FF78FB33FED92F51" blockId="44.[189,342,1266,1292]" box="[189,342,1266,1292]" pageId="44" pageNumber="45">
<heading id="993A8196CF2BFFD3FF78FB33FED92F51" bold="true" box="[189,342,1266,1292]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="F0B9EAE8CF2BFFD3FF78FB33FED92F51" bold="true" box="[189,342,1266,1292]" pageId="44" pageNumber="45">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF2BFFD3FF78FAE3FCA12F61" blockId="44.[189,814,1313,1340]" box="[189,814,1313,1340]" pageId="44" pageNumber="45">
Known from the
<typeStatus id="1D768858CF2BFFD3FE43FAE3FE362F61" box="[390,441,1314,1340]" pageId="44" pageNumber="45">type</typeStatus>
locality in
<collectingRegion id="0009F818CF2BFFD3FDFBFAE3FD242F61" box="[574,683,1314,1340]" country="Myanmar" name="Yangon" pageId="44" pageNumber="45">Rangoon</collectingRegion>
,
<collectingCountry id="BADA766ACF2BFFD3FD73FAE3FCA52F61" box="[694,810,1314,1340]" name="Myanmar" pageId="44" pageNumber="45">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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