134 lines
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134 lines
14 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.468.8506" ID-GBIF-Dataset="df3f2d10-3b5e-480f-936c-368e01ffd7b1" ID-PMC="PMC4296520" ID-Pensoft-Pub="1313-2970-468-1" ID-PubMed="25610341" ID-ZBK="AE5AB793FDC74DCD8A47AE96A141E2AD" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-468-1" ModsDocOrigin="ZooKeys 468" ModsDocTitle="Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1451244902156" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo, Panjaitan, Rawati, Menufandu, Herlina & Balke, Michael" docDate="2014" docId="4A41F2FA37DFB6D6E3A464130963814F" docLanguage="en" docName="ZooKeys 468: 1-83" docOrigin="ZooKeys 468" docSource="http://dx.doi.org/10.3897/zookeys.468.8506" docTitle="Exocelina gorokaensis Shaverdo & Balke, sp. n." docType="treatment" docUuid="C87417FC-F7D6-4395-9897-02628EB0DF59" docUuidSource="ZooBank" docVersion="4" lastPageNumber="8" masterDocId="FFE9FFA5117AFFD17452FFB5FF94FFE3" masterDocTitle="Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="8" updateTime="1668159774671" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panjaitan, Rawati</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Menufandu, Herlina</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>468</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.468.8506</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.468.8506</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-468-1</mods:identifier>
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<mods:identifier type="ZBK">AE5AB793FDC74DCD8A47AE96A141E2AD</mods:identifier>
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<mods:identifier type="ZooBank">AE5AB793FDC74DCD8A47AE96A141E2AD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152057381" LSID="urn:lsid:zoobank.org:act:C87417FC-F7D6-4395-9897-02628EB0DF59" httpUri="http://treatment.plazi.org/id/4A41F2FA37DFB6D6E3A464130963814F" lastPageNumber="8" pageId="7" pageNumber="8">
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<subSubSection pageId="7" pageNumber="8" type="multiple">
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<paragraph pageId="7" pageNumber="8">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="nomenclature">
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<paragraph pageId="7" pageNumber="8">
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4.
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<taxonomicName LSID="http://zoobank.org/C87417FC-F7D6-4395-9897-02628EB0DF59" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina gorokaensis" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gorokaensis">Exocelina gorokaensis Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="7" pageNumber="8">sp. n.</taxonomicNameLabel>
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Figs 14, 42
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="reference_group">
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<paragraph pageId="7" pageNumber="8">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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undescribed sp. MB1307:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="45" pageNumber="46" pagination="1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity." url="10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
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: Supplementary figs 1-4, Tab. 2.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="type locality">
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<paragraph pageId="7" pageNumber="8">Type locality.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Papua New Guinea: Eastern Highlands Province, 37 km S Goroka, Hogave village, Mt. Michael,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-6.3746667">06°22.48'S</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="145.25433">145°15.26'E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="type material">
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<paragraph pageId="7" pageNumber="8">Type material.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Holotype: male "Papua New Guinea: Eastern Highlands, 37 km S Goroka, Hogave vill., Mt. Michael, 2179-2800m, 9.-15vii.2009, 06.22.479S 145.15.256E, Sagata (PNG 230)" (ZSM). Paratypes: Eastern Highlands: 23 males, 39 females with the same label as the holotype, 1 male additionally with a green label "DNA M.Balke 4038" (NHMW, ZSM). 5 males, 4 females "Papua New Guinea: Eastern Highlands, Goroka, Mt. Gahavisuka, 2200m, 8.iv.2006, 06.00.896S 145.24.753E, Balke & Sagata (PNG 35)" (NHMW, ZSM). 1 male, 1 female "Papua New Guinea: Eastern Highlands, Hogu, Mt. Barola, 1900m, 9.v.2006, 06.17.556S 145.45.036E, Balke & Sagata (PNG 56)" (ZSM). 10 males, 5 females "Papua New Guinea: Eastern Highlands, Goroka, below Mt. Otto, 2000m, 11.v.2006, 06.01.687S 145.26.493E, Balke (PNG 57)", one male additionally with a green label "DNA M.Balke 1305" (NHMW, ZSM). 21 males, 14 females "Papua New Guinea: Eastern Highlands, Goroka, Daulo Pass, 2500m, 19.v.2006, 06.02.432S 145.13.333E, John & Balke (PNG 67)", one male additionally with a green label "DNA M.Balke 1307" (NHMW, ZSM). 2 males, 1 female "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 2200m, 23.xi.2006, 05.56.801S 145.22.238E, Balke & Kinibel (PNG 106)" (ZSM). 1 female "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 2400m, 23.xi.2006, 05.55.615S 145.22.699E, Balke & Kinibel (PNG 107)", "DNA M.Balke 1518" [green] (ZSM). Simbu: 8 males, 5 females "Papua New Guinea Simbu prov L. Cizek lgt.", "Kundiawa, Mu vill.
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="145.03334">145°02'E</geoCoordinate>
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-4.7">4°42'S</geoCoordinate>
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[
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-6.0833335">6°05'S</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="145.03334">145°02'E</geoCoordinate>
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] III.2001, 1900m" (ZSM). Western Highlands: 9 males, 4 females "Papua New Guinea: Western Highlands, Mondmill,5 Km SE Minj, small pools near creek, 1741m, 12.vi.2006, 05.56.801S 144.39.898E, John (PNG 77)", one male additionally with a green label "DNA M.Balke 1343" (NHMW, ZSM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="diagnosis">
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<paragraph pageId="7" pageNumber="8">Diagnosis.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Beetle medium-sized, dark brown to piceous, with paler clypeus, vertex, and pronotal sides, submatt; pronotum with distinct lateral bead; male antennomeres 3-5 evidently enlarged, slightly rounded, almost equal in size, antennomeres 6 and 7 somewhat enlarged; male protarsomere 4 with medium-sized, slender, evidently curved anterolateral hook-like seta; median lobe with very weak submedian constriction and apex very slightly concave in ventral view and with apex slightly pointed and broadened in lateral view; paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, more or less long, thin setae. The species is similar to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina jimiensis" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jimiensis">Exocelina jimiensis</taxonomicName>
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sp. n., from which differs with evident punctation of the dorsal surface and shape of the median lobe. Also it is similar to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina vovai" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="vovai">Exocelina vovai</taxonomicName>
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sp. n. and
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina bismarckensis" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bismarckensis">Exocelina bismarckensis</taxonomicName>
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sp. n., from which differs with weaker punctation of the dorsal surface, paramere with distinct notch on the dorsal side, and shape of the median lobe.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="description">
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<paragraph pageId="7" pageNumber="8">Description.</paragraph>
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<paragraph pageId="7" pageNumber="8">Size and shape: Beetle medium-sized (TL-H 3.7-4.3 mm, TL 4.1-4.8 mm, MW 1.95-2.4 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Dorsal surface more or less uniform dark brown to piceous, paler on clypeus, vertex, pronotal sides, and along elytral suture; head appendages and legs yellowish red to dark reddish, legs darker distally (Fig. 42). Teneral specimens paler.</paragraph>
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<paragraph pageId="7" pageNumber="8">Surface sculpture: Head with very dense, coarse punctation (spaces between punctures 1-2 times size of punctures). Pronotum with punctation finer than on head. Elytra with punctation sparser than on pronotum. Pronotum and elytra with relatively strongly impressed microreticulation, dorsal surface submatt. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal figs with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal ventrites.</paragraph>
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<paragraph pageId="7" pageNumber="8">Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process and neck of prosternal process with distinct ridge, smooth and not rounded anteriorly, with small anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct lateral bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 broadly rounded or slightly truncate apically.</paragraph>
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<paragraph pageId="7" pageNumber="8">Male: Antennomeres 3-5 evidently enlarged, slightly rounded, almost equal in size, antennomeres 6 and 7 somewhat enlarged (Fig. 14A), antennomeres 3-7 rugose ventrally. Protarsomere 4 with medium-sized, slender, evidently curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 17 and posterior row of 6 short setae (Fig. 14B). Abdominal ventrite 6 with 11-14 lateral striae on each side, broadly rounded or slightly truncate apically. Median lobe with very weak submedian constriction and apex very slightly concave in ventral view and with apex slightly pointed and broadened in lateral view (Fig. 14C, D). Paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, more or less long, thin setae (Fig. 14E).</paragraph>
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<paragraph pageId="7" pageNumber="8">Holotype: TL-H 4.0 mm, TL 4.5 mm, MW 2.15 mm.</paragraph>
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<paragraph pageId="7" pageNumber="8">Female: Antennae simple, abdominal ventrite 6 broadly rounded apically, without striae.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="distribution">
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<paragraph pageId="7" pageNumber="8">Distribution.</paragraph>
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<paragraph pageId="7" pageNumber="8">Papua New Guinea: Eastern Highlands, Simbu, and Western Highlands Provinces. The species is known mainly from the area around Goroka and Kundiawa (Fig. 53).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="etymology">
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<paragraph pageId="7" pageNumber="8">Etymology.</paragraph>
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<paragraph pageId="7" pageNumber="8">The species is named after Goroka, where it was mostly collected. The name is an adjective in the nominative singular.</paragraph>
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</subSubSection>
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</treatment>
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</document> |