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<document id="7B8D30114255DD7C07E20F2C82CBEEDE" ID="10.11646/zootaxa.4258.1.5SLASH10900" ID-CLB-Dataset="33163" ID-DOI="10.11646/zootaxa.4258.1.5" ID-GBIF-Dataset="bdf2db84-05b5-4c94-bfa7-7cf9274fda9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="569169" ID-ZooBank="D1095E5E-2C9F-40C6-BA10-EADBEA56681C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493196110549" checkinUser="plazi" docAuthor="Fomichev, Alexander A. &amp; Marusik, Yuri M." docDate="2017" docId="039C9E749666FFF5FF40FB98DCDAF8FA" docLanguage="en" docName="zootaxa.4258.1.5SLASH10900.pdf" docOrigin="Zootaxa 4258 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Berlandina litvinovi Fomichev &amp; Marusik, 2017, sp. n." docType="treatment" docVersion="13" lastPageNumber="75" masterDocId="FFA5E60C9662FFF3FFD7FFE8D87AFFD4" masterDocTitle="A survey of East Palaearctic Gnaphosidae (Araneae). 8. New data on Berlandina and Gnaphosa from Mongolia" masterLastPageNumber="80" masterPageNumber="69" pageNumber="73" updateTime="1698439081613" updateUser="ExternalLinkService">
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<mods:title id="474B62CB324F9AD23DFCD53030DC12C1">A survey of East Palaearctic Gnaphosidae (Araneae). 8. New data on Berlandina and Gnaphosa from Mongolia</mods:title>
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<mods:namePart id="9E46F6EF43E6A64FD47BDF149E981ACB">Fomichev, Alexander A.</mods:namePart>
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<mods:namePart id="AFCCDBA3C973D558F1C37C0606C7774F">Marusik, Yuri M.</mods:namePart>
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<mods:title id="946CD716B8EDFF1E63A826472C1B8385">Zootaxa</mods:title>
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<mods:date id="92E3240ABCC2044589920CD5281A81EE">2017</mods:date>
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<mods:number id="D2E38D89A72643B54182B0D58689E44A">4258</mods:number>
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<mods:identifier id="A50AE3A169DD6A25E4F29F2D1D21841F" type="CLB-Dataset">33163</mods:identifier>
<mods:identifier id="E3F9D803CFFC47C0715ACB7C4440A27C" type="DOI">10.11646/zootaxa.4258.1.5</mods:identifier>
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<treatment id="039C9E749666FFF5FF40FB98DCDAF8FA" ID-DOI="http://doi.org/10.5281/zenodo.6028587" ID-GBIF-Taxon="128699700" ID-Zenodo-Dep="6028587" LSID="urn:lsid:plazi:treatment:039C9E749666FFF5FF40FB98DCDAF8FA" httpUri="http://treatment.plazi.org/id/039C9E749666FFF5FF40FB98DCDAF8FA" lastPageId="6" lastPageNumber="75" pageId="4" pageNumber="73">
<subSubSection id="C32F7CE99666FFF7FF40FB98D94AFB78" pageId="4" pageNumber="73" type="nomenclature">
<paragraph id="8B8A2F629666FFF7FF40FB98D9A8FB5F" blockId="4.[151,466,1136,1196]" box="[151,466,1136,1163]" pageId="4" pageNumber="73">
<heading id="D0C2980E9666FFF7FF40FB98D9A8FB5F" bold="true" box="[151,466,1136,1163]" fontSize="11" level="1" pageId="4" pageNumber="73" reason="1">
<emphasis id="B941F3709666FFF7FF40FB98D9A8FB5F" bold="true" box="[151,466,1136,1163]" pageId="4" pageNumber="73">
<taxonomicName id="4C3554E19666FFF7FF40FB98D9F3FB5E" ID-CoL="68JT4" box="[151,393,1136,1162]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="litvinovi" status="sp. nov.">
<emphasis id="B941F3709666FFF7FF40FB98D9F3FB5E" bold="true" box="[151,393,1136,1162]" italics="true" pageId="4" pageNumber="73">Berlandina litvinovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2724E0B9666FFF7FE47FB99D9A8FB5F" box="[400,466,1137,1163]" pageId="4" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A2F629666FFF7FF40FB7BD94AFB78" blockId="4.[151,466,1136,1196]" box="[151,304,1171,1196]" pageId="4" pageNumber="73">
<figureCitation id="130E33E79666FFF7FF40FB7BD973FB78" box="[151,265,1171,1196]" captionStart="FIGURES 9 ‒ 19" captionStartId="5.[151,264,1873,1896]" captionTargetBox="[151,1432,202,1852]" captionTargetId="figure@5.[151,1436,195,1852]" captionTargetPageId="5" captionText="FIGURES 9 ‒ 19. Habitus and copulatory organs of Berlandina litvinovi sp. n. 9 ‒ 10 female and male habitus, dorsal; 11 ‒ 13 male palp, prolateral, ventral and retrolateral respectively; 14 ‒ 16 bulb, anterior, ventral and retrolateral; 17 intact epigyne, ventral; 18 ‒ 19 macerated epigyne, ventral and dorsal. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569171/files/figure.png" pageId="4" pageNumber="73">Figs 9‒19</figureCitation>
,
<figureCitation id="130E33E79666FFF7FEC3FB7BD94AFB78" box="[276,304,1171,1196]" captionStart="FIGURES 34 ‒ 35" captionStartId="10.[151,264,1665,1688]" captionTargetBox="[188,1398,1095,1643]" captionTargetId="figure@10.[188,1398,1095,1643]" captionTargetPageId="10" captionText="FIGURES 34 ‒ 35. Collecting localities of Berlandina and Gnaphosa species examined in this study. Solid symbol on Fig. 34 refers to new records, open symbols to the literature data, dotted square refers to the region shown on Fig. 35. Blue squares in Fig. 35 refer to the records of G. ustyuzhanini (= G. rasnitsyni). One symbol may designate more than one locality." httpUri="https://zenodo.org/record/569174/files/figure.png" pageId="4" pageNumber="73">35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99666FFF7FF40FB33D8AFFA50" pageId="4" pageNumber="73" type="description">
<paragraph id="8B8A2F629666FFF7FF40FB33D8AFFA50" blockId="4.[151,1437,1243,1916]" pageId="4" pageNumber="73">
<materialsCitation id="3B5D253F9666FFF7FF40FB33DA50FAE8" ID-GBIF-Occurrence="1493295190" collectingDate="2015-05-18" collectionCode="ISEA, ZMMU" country="Mongolia" elevation="1280" latitude="45.766666" location="Bodonchiyn-Gol River Valley" longLatPrecision="1128" longitude="92.2" municipality="Altai Village" pageId="4" pageNumber="73" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Khovd" typeStatus="holotype">
<emphasis id="B941F3709666FFF7FF40FB33D899FB20" bold="true" box="[151,227,1243,1268]" pageId="4" pageNumber="73">
<typeStatus id="548E91C09666FFF7FF40FB33D8A5FB20" box="[151,223,1243,1268]" pageId="4" pageNumber="73">Types</typeStatus>
.
</emphasis>
<collectingCountry id="F3226FF29666FFF7FF3DFB34D9FDFB20" box="[234,391,1244,1268]" name="Mongolia" pageId="4" pageNumber="73">MONGOLIA</collectingCountry>
:
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<emphasis id="B941F3709666FFF7FE47FB33D9A1FB20" bold="true" box="[400,475,1243,1268]" italics="true" pageId="4" pageNumber="73">Khovd</emphasis>
</collectingRegion>
Aimag:
<typeStatus id="548E91C09666FFF7FDEBFB33DAE4FB20" box="[572,670,1243,1268]" pageId="4" pageNumber="73" type="holotype">holotype</typeStatus>
<specimenCount id="9D33E4EB9666FFF7FD72FB34DAC0FB27" box="[677,698,1244,1267]" pageId="4" pageNumber="73" type="male">♂</specimenCount>
(
<collectionCode id="ED24B7A79666FFF7FD1FFB34DB73FB20" box="[712,777,1244,1268]" pageId="4" pageNumber="73">ISEA</collectionCode>
) and
<typeStatus id="548E91C09666FFF7FC91FB34DBCBFB20" box="[838,945,1244,1268]" pageId="4" pageNumber="73" type="paratype">paratypes</typeStatus>
<specimenCount id="9D33E4EB9666FFF7FC6FFB34DBA1FB20" box="[952,987,1244,1268]" pageId="4" pageNumber="73" type="male">2♂</specimenCount>
<specimenCount id="9D33E4EB9666FFF7FC36FB34DC7EFB20" box="[993,1028,1244,1268]" pageId="4" pageNumber="73" type="female">1♀</specimenCount>
(
<collectionCode id="ED24B7A79666FFF7FBC4FB34DC2EFB20" box="[1043,1108,1244,1268]" pageId="4" pageNumber="73">ISEA</collectionCode>
)
<specimenCount id="9D33E4EB9666FFF7FBB5FB34DCFFFB20" box="[1122,1157,1244,1268]" pageId="4" pageNumber="73" type="male">1♂</specimenCount>
<specimenCount id="9D33E4EB9666FFF7FB5BFB34DCD4FB20" box="[1164,1198,1244,1268]" pageId="4" pageNumber="73" type="female">1♀</specimenCount>
(
<collectionCode id="ED24B7A79666FFF7FB6AFB34DD6DFB20" box="[1213,1303,1244,1268]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="73">ZMMU</collectionCode>
)
<quantity id="4CCD82879666FFF7FAF2FB33DD16FB20" box="[1317,1388,1243,1268]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="73" unit="km" value="36.0">36 km</quantity>
SW from
<collectingMunicipality id="6BEEB5189666FFF7FF0FFB17D914FACC" box="[216,366,1279,1304]" pageId="4" pageNumber="73">Altai Village</collectingMunicipality>
,
<location id="8EEA79B99666FFF7FEAAFB17DAAFFACC" LSID="urn:lsid:plazi:treatment:039C9E749666FFF5FF40FB98DCDAF8FA:8EEA79B99666FFF7FEAAFB17DAAFFACC" box="[381,725,1279,1304]" country="Mongolia" latitude="45.766666" longLatPrecision="1128" longitude="92.2" municipality="Altai Village" name="Bodonchiyn-Gol River Valley" pageId="4" pageNumber="73" stateProvince="Khovd">Bodonchiyn-Gol River Valley</location>
,
<geoCoordinate id="EE0149A59666FFF7FD33FB17DB3DFACC" box="[740,839,1279,1304]" direction="north" orientation="latitude" pageId="4" pageNumber="73" precision="925" value="45.766666">45°46’N</geoCoordinate>
,
<geoCoordinate id="EE0149A59666FFF7FC81FAE8DBCCFACC" box="[854,950,1280,1304]" direction="east" orientation="longitude" pageId="4" pageNumber="73" precision="925" value="92.2">92°12’E</geoCoordinate>
, tugay (gallery forest), night, attracted by light,
<elevation id="0018C8519666FFF7FF0DFACCD948FAEF" box="[218,306,1316,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="4" pageNumber="73" unit="m" value="1280.0">
<quantity id="4CCD82879666FFF7FF0DFACCD948FAEF" box="[218,306,1316,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="4" pageNumber="73" unit="m" value="1280.0">1280 m</quantity>
</elevation>
, 17‒
<collectingDate id="EFCFF04A9666FFF7FEB0FACBD99DFAE8" box="[359,487,1315,1340]" pageId="4" pageNumber="73" value="2015-05-18">
<date id="FF8B09A29666FFF7FEB0FACBD99DFAE8" box="[359,487,1315,1340]" pageId="4" pageNumber="73" value="2015-05-18">18.05.2015</date>
</collectingDate>
(AF)
</materialsCitation>
;
<materialsCitation id="3B5D253F9666FFF7FDE0FACCDD68FAE8" ID-GBIF-Occurrence="1493295189" box="[567,1298,1315,1340]" collectingDate="2015-05-18" collectionCode="ISEA" country="Mongolia" elevation="1280" latitude="45.766666" location="Bodonchiyn-Gol River Valley" longLatPrecision="1128" longitude="92.166664" municipality="Altai Village" pageId="4" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Khovd" typeStatus="holotype">
<specimenCount id="9D33E4EB9666FFF7FDE0FACCDA20FAE8" box="[567,602,1316,1340]" pageId="4" pageNumber="73" type="female">1♀</specimenCount>
(
<collectionCode id="ED24B7A79666FFF7FDBCFACCDAD4FAE8" box="[619,686,1316,1340]" pageId="4" pageNumber="73">ISEA</collectionCode>
), same locality,
<geoCoordinate id="EE0149A59666FFF7FCB0FACBDBB0FAE8" box="[871,970,1315,1340]" direction="north" orientation="latitude" pageId="4" pageNumber="73" precision="925" value="45.766666">45°46’N</geoCoordinate>
,
<geoCoordinate id="EE0149A59666FFF7FC00FACCDC4DFAE8" box="[983,1079,1316,1340]" direction="east" orientation="longitude" pageId="4" pageNumber="73" precision="925" value="92.166664">92°10’E</geoCoordinate>
, 7–8.05.2012 (AF)
</materialsCitation>
;
<materialsCitation id="3B5D253F9666FFF7FAF7FACCD8AFFA50" ID-GBIF-Occurrence="1493295196" collectingDate="2012-05-16" collectionCode="ISEA" country="Mongolia" elevation="1850" latitude="46.35" location="Arshantyn-Nuruu Mt. Range" longLatPrecision="1124" longitude="91.21667" pageId="4" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Khovd" typeStatus="holotype">
<specimenCount id="9D33E4EB9666FFF7FAF7FACCDD38FAE8" box="[1312,1346,1316,1340]" pageId="4" pageNumber="73" type="female">1♀</specimenCount>
(
<collectionCode id="ED24B7A79666FFF7FA84FACCDDEEFAE8" box="[1363,1428,1316,1340]" pageId="4" pageNumber="73">ISEA</collectionCode>
)
<location id="8EEA79B99666FFF7FF40FAAFD99EFAB4" LSID="urn:lsid:plazi:treatment:039C9E749666FFF5FF40FB98DCDAF8FA:8EEA79B99666FFF7FF40FAAFD99EFAB4" box="[151,484,1351,1376]" country="Mongolia" latitude="46.35" longLatPrecision="1124" longitude="91.21667" name="Arshantyn-Nuruu Mt. Range" pageId="4" pageNumber="73" stateProvince="Khovd">Arshantyn-Nuruu Mt. Range</location>
,
<geoCoordinate id="EE0149A59666FFF7FE24FAAFDA2CFAB4" box="[499,598,1351,1376]" direction="north" orientation="latitude" pageId="4" pageNumber="73" precision="925" value="46.35">46°21’N</geoCoordinate>
,
<geoCoordinate id="EE0149A59666FFF7FDB2FAA0DABFFAB4" box="[613,709,1352,1376]" direction="east" orientation="longitude" pageId="4" pageNumber="73" precision="925" value="91.21667">91°13’E</geoCoordinate>
, mountain stony steppe with rocks,
<elevation id="0018C8519666FFF7FBB9FAA0DD75FA8B" box="[1134,1295,1352,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="4" pageNumber="73" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">
<quantity id="4CCD82879666FFF7FBB9FAA0DD75FA8B" box="[1134,1295,1352,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="4" pageNumber="73" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">1700‒2000 m</quantity>
</elevation>
,
<collectingDate id="EFCFF04A9666FFF7FAC9FAAFDDE6FAB4" box="[1310,1436,1351,1376]" pageId="4" pageNumber="73" value="2012-05-16">
<date id="FF8B09A29666FFF7FAC9FAAFDDE6FAB4" box="[1310,1436,1351,1376]" pageId="4" pageNumber="73" value="2012-05-16">16.05.2012</date>
</collectingDate>
(AF).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99666FFF7FF10FA67DBADFA18" pageId="4" pageNumber="73" type="etymology">
<paragraph id="8B8A2F629666FFF7FF10FA67DBADFA18" blockId="4.[151,1437,1243,1916]" pageId="4" pageNumber="73">
<emphasis id="B941F3709666FFF7FF10FA67D934FA7C" bold="true" box="[199,334,1423,1448]" pageId="4" pageNumber="73">Etymology.</emphasis>
The specific name is a patronym in honour of Anton N. Litvinov (Terekty,
<collectingCountry id="F3226FF29666FFF7FB41FA67DD67FA7C" box="[1174,1309,1423,1448]" name="Kazakhstan" pageId="4" pageNumber="73">Kazakhstan</collectingCountry>
), the friend of AF, who helped a lot in spider collecting and organization of field trips.
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99666FFF7FF10FA3FDBC5F938" pageId="4" pageNumber="73" type="diagnosis">
<paragraph id="8B8A2F629666FFF7FF10FA3FDBC5F938" blockId="4.[151,1437,1243,1916]" pageId="4" pageNumber="73">
<emphasis id="B941F3709666FFF7FF10FA3FD93BFA24" bold="true" box="[199,321,1495,1520]" pageId="4" pageNumber="73">Diagnosis.</emphasis>
Males of the new species differ from those of the similar species
<taxonomicName id="4C3554E19666FFF7FBE1FA31DCC7FA24" box="[1078,1213,1496,1520]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="B941F3709666FFF7FBE1FA31DCC7FA24" box="[1078,1213,1496,1520]" italics="true" pageId="4" pageNumber="73">B. koponeni</emphasis>
</taxonomicName>
by tegulum bulged prolaterally (
<emphasis id="B941F3709666FFF7FEFAFA14D936F9C7" box="[301,332,1532,1555]" italics="true" pageId="4" pageNumber="73">Bu</emphasis>
), by less curved (as viewed from above) median apophysis, absence of the ridge in embolic division and much longer tibial apophysis (cf.
<figureCitation id="130E33E79666FFF7FD7FF9F7DB5FF9EC" box="[680,805,1567,1592]" captionStart="FIGURES 9 ‒ 19" captionStartId="5.[151,264,1873,1896]" captionTargetBox="[151,1432,202,1852]" captionTargetId="figure@5.[151,1436,195,1852]" captionTargetPageId="5" captionText="FIGURES 9 ‒ 19. Habitus and copulatory organs of Berlandina litvinovi sp. n. 9 ‒ 10 female and male habitus, dorsal; 11 ‒ 13 male palp, prolateral, ventral and retrolateral respectively; 14 ‒ 16 bulb, anterior, ventral and retrolateral; 17 intact epigyne, ventral; 18 ‒ 19 macerated epigyne, ventral and dorsal. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569171/files/figure.png" pageId="4" pageNumber="73">Figs 11‒16</figureCitation>
and figs
<quantity id="4CCD82879666FFF7FC47F9F7DB8EF9EC" box="[912,1012,1567,1592]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" metricValueMax="9.144" metricValueMin="7.62" pageId="4" pageNumber="73" unit="in" value="33.0" valueMax="36.0" valueMin="30.0">30‒36 in</quantity>
<bibRefCitation id="EFA452939666FFF7FC2BF9F7DC96F9EC" author="Marusik" box="[1020,1260,1567,1592]" pageId="4" pageNumber="73" refString="Marusik, Y. M., Fomichev, A. A. &amp; Omelko, M. M. (2014 a) A survey of East Palaearctic Gnaphosidae (Araneae). 1. On the Berlandina Dalmas, 1922 (Gnaphosinae) from Mongolia and adjacent regions. Zootaxa, 3827 (2), 187 - 213. https: // doi. org / 10.11646 / zootaxa. 3827.2.4" type="journal article" year="2014" yearSuffix="a">
Marusik
<emphasis id="B941F3709666FFF7FBB6F9C9DCE0F9EC" box="[1121,1178,1568,1592]" italics="true" pageId="4" pageNumber="73">et al.</emphasis>
2014a
</bibRefCitation>
). Females of
<taxonomicName id="4C3554E19666FFF7FA53F9C9D889F988" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="litvinovi" status="sp. nov.">
<emphasis id="B941F3709666FFF7FA53F9C9D889F988" italics="true" pageId="4" pageNumber="73">B. litvinovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2724E0B9666FFF7FF2AF9ABD93BF988" box="[253,321,1603,1628]" pageId="4" pageNumber="73" rank="species">
<emphasis id="B941F3709666FFF7FF2AF9ABD93BF988" bold="true" box="[253,321,1603,1628]" pageId="4" pageNumber="73">sp. n.</emphasis>
</taxonomicNameLabel>
are similar to these of
<taxonomicName id="4C3554E19666FFF7FD8FF9ACDC40F988" authority="Marusik &amp; Logunov, 1995" authorityName="Marusik &amp; Logunov" authorityYear="1995" box="[600,1082,1603,1628]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="ubsunurica">
<emphasis id="B941F3709666FFF7FD8FF9ACDA83F988" box="[600,761,1604,1628]" italics="true" pageId="4" pageNumber="73">B. ubsunurica</emphasis>
Marusik &amp; Logunov, 1995
</taxonomicName>
and
<taxonomicName id="4C3554E19666FFF7FBAFF9ACD93CF954" authority="Ponomarev &amp; Tsvetkov, 2006" authorityName="Ponomarev &amp; Tsvetkov" authorityYear="2006" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="B941F3709666FFF7FBAFF9ACDC8AF988" box="[1144,1264,1604,1628]" italics="true" pageId="4" pageNumber="73">B. nenilini</emphasis>
Ponomarev &amp; Tsvetkov, 2006
</taxonomicName>
. They differ from two aforementioned species by wide fovea, less pronounced anterior protrusion on the edge of fovea and by less protruding outgrowths of lateral pockets (cf.
<figureCitation id="130E33E79666FFF7FC23F963DC0BF970" box="[1012,1137,1675,1700]" captionStart="FIGURES 9 ‒ 19" captionStartId="5.[151,264,1873,1896]" captionTargetBox="[151,1432,202,1852]" captionTargetId="figure@5.[151,1436,195,1852]" captionTargetPageId="5" captionText="FIGURES 9 ‒ 19. Habitus and copulatory organs of Berlandina litvinovi sp. n. 9 ‒ 10 female and male habitus, dorsal; 11 ‒ 13 male palp, prolateral, ventral and retrolateral respectively; 14 ‒ 16 bulb, anterior, ventral and retrolateral; 17 intact epigyne, ventral; 18 ‒ 19 macerated epigyne, ventral and dorsal. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569171/files/figure.png" pageId="4" pageNumber="73">Figs 17‒19</figureCitation>
and figs
<quantity id="4CCD82879666FFF7FB01F963DD43F970" box="[1238,1337,1675,1700]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="4" pageNumber="73" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10‒15 in</quantity>
<bibRefCitation id="EFA452939666FFF7FAE8F963D95BF91C" author="Marusik" pageId="4" pageNumber="73" refString="Marusik, Y. M., Fomichev, A. A. &amp; Omelko, M. M. (2014 a) A survey of East Palaearctic Gnaphosidae (Araneae). 1. On the Berlandina Dalmas, 1922 (Gnaphosinae) from Mongolia and adjacent regions. Zootaxa, 3827 (2), 187 - 213. https: // doi. org / 10.11646 / zootaxa. 3827.2.4" type="journal article" year="2014" yearSuffix="a">
Marusik
<emphasis id="B941F3709666FFF7FF40F959D8AAF91C" box="[151,208,1712,1736]" italics="true" pageId="4" pageNumber="73">et al.</emphasis>
2014a
</bibRefCitation>
). Both male and female of the new species can be distinguished from the closest species
<taxonomicName id="4C3554E19666FFF7FAC2F959DDE6F91C" box="[1301,1436,1712,1736]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="B941F3709666FFF7FAC2F959DDE6F91C" box="[1301,1436,1712,1736]" italics="true" pageId="4" pageNumber="73">B. koponeni</emphasis>
</taxonomicName>
by dark coloration (cf.
<figureCitation id="130E33E79666FFF7FE4FF93BD9A7F938" box="[408,477,1747,1772]" captionStart="FIGURES 1 ‒ 8" captionStartId="2.[151,264,1912,1935]" captionTargetBox="[214,1354,194,1876]" captionTargetId="figure@2.[212,1354,193,1876]" captionTargetPageId="2" captionText="FIGURES 1 ‒ 8. Habitus and epigyne of Berlandina koponeni (1 ‒ 4) and B. nakonechnyi (5 ‒ 8). 1, 5 habitus, dorsal; 2, 7 macerated epigyne, ventral; 3, 6 intact epigyne, ventral; 4, 8 macerated epigyne, dorsal. Scale = 0.2 mm if not otherwise indicated. Abbreviation. Op outgrowth of lateral pockets." httpUri="https://zenodo.org/record/569170/files/figure.png" pageId="4" pageNumber="73">Figs 1</figureCitation>
,
<figureCitation id="130E33E79666FFF7FE3CF93CDA59F938" box="[491,547,1748,1772]" captionStart="FIGURES 9 ‒ 19" captionStartId="5.[151,264,1873,1896]" captionTargetBox="[151,1432,202,1852]" captionTargetId="figure@5.[151,1436,195,1852]" captionTargetPageId="5" captionText="FIGURES 9 ‒ 19. Habitus and copulatory organs of Berlandina litvinovi sp. n. 9 ‒ 10 female and male habitus, dorsal; 11 ‒ 13 male palp, prolateral, ventral and retrolateral respectively; 14 ‒ 16 bulb, anterior, ventral and retrolateral; 17 intact epigyne, ventral; 18 ‒ 19 macerated epigyne, ventral and dorsal. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569171/files/figure.png" pageId="4" pageNumber="73">9‒10</figureCitation>
and fig.
<quantity id="4CCD82879666FFF7FD50F93CDABAF938" box="[647,704,1747,1772]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" pageId="4" pageNumber="73" unit="in" value="37.0">37 in</quantity>
<bibRefCitation id="EFA452939666FFF7FD10F93BDBCEF938" author="Marusik" box="[711,948,1747,1772]" pageId="4" pageNumber="73" refString="Marusik, Y. M., Fomichev, A. A. &amp; Omelko, M. M. (2014 a) A survey of East Palaearctic Gnaphosidae (Araneae). 1. On the Berlandina Dalmas, 1922 (Gnaphosinae) from Mongolia and adjacent regions. Zootaxa, 3827 (2), 187 - 213. https: // doi. org / 10.11646 / zootaxa. 3827.2.4" type="journal article" year="2014" yearSuffix="a">
Marusik
<emphasis id="B941F3709666FFF7FCFCF93CDB19F938" box="[811,867,1748,1772]" italics="true" pageId="4" pageNumber="73">et al.</emphasis>
2014a
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99666FFF5FF10F91FDD5AF96E" lastPageId="6" lastPageNumber="75" pageId="4" pageNumber="73" type="description">
<paragraph id="8B8A2F629666FFF7FF10F91FDD4DF8A8" blockId="4.[151,1437,1243,1916]" pageId="4" pageNumber="73">
<emphasis id="B941F3709666FFF7FF10F91FD923F8C4" bold="true" box="[199,345,1783,1808]" pageId="4" pageNumber="73">Description.</emphasis>
Male (
<typeStatus id="548E91C09666FFF7FE79F91FDA6DF8C4" box="[430,535,1783,1808]" pageId="4" pageNumber="73" type="holotype">holotype</typeStatus>
). Total length 5.4. Carapace: 2.25 long, 1.8 wide. Carapace brown with black margins, spots and radial stripes. Chelicerae and labium dark-brown. Maxillae light-brown. Sternum brown. Legs: femora, patellae and tibiae gray-brown with yellow spots and stripes, metatarsi and tarsi dirty-yellow. Palps yellow-gray. Abdomen gray dorsally with light longitudinal stripe, venter yellow. Spinnerets yellow-gray.
</paragraph>
<caption id="DF4A7FEA9667FFF6FF40F8B9DC16F871" httpUri="https://zenodo.org/record/569171/files/figure.png" pageId="5" pageNumber="74" targetBox="[151,1432,202,1852]" targetPageId="5">
<paragraph id="8B8A2F629667FFF6FF40F8B9DC16F871" blockId="5.[151,1435,1873,1987]" pageId="5" pageNumber="74">
<emphasis id="B941F3709667FFF6FF40F8B9D933F8B3" bold="true" box="[151,329,1873,1896]" pageId="5" pageNumber="74">FIGURES 9‒19.</emphasis>
Habitus and copulatory organs of
<taxonomicName id="4C3554E19667FFF6FD65F8BADB05F8B3" box="[690,895,1874,1895]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="litvinovi" status="sp. nov.">
<emphasis id="B941F3709667FFF6FD65F8BADB05F8B3" box="[690,895,1874,1895]" italics="true" pageId="5" pageNumber="74">Berlandina litvinovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2724E0B9667FFF6FC50F8BADBB8F8B3" box="[903,962,1874,1895]" pageId="5" pageNumber="74" rank="species">
<emphasis id="B941F3709667FFF6FC50F8BADBB8F8B3" bold="true" box="[903,962,1874,1895]" pageId="5" pageNumber="74">sp. n.</emphasis>
</taxonomicNameLabel>
9‒10 female and male habitus, dorsal; 11‒13 male palp, prolateral, ventral and retrolateral respectively; 14‒16 bulb, anterior, ventral and retrolateral; 17 intact epigyne, ventral; 18‒19 macerated epigyne, ventral and dorsal. Scale = 0.2 mm if not otherwise indicated.
</paragraph>
</caption>
<paragraph id="8B8A2F629667FFF6FF40F845DADDF817" blockId="5.[151,1435,1873,1987]" box="[151,679,1965,1987]" pageId="5" pageNumber="74">
Abbreviations.
<emphasis id="B941F3709667FFF6FEE4F846D935F817" box="[307,335,1966,1987]" italics="true" pageId="5" pageNumber="74">Bu</emphasis>
tegular bulge,
<emphasis id="B941F3709667FFF6FE30F846D985F817" box="[487,511,1966,1987]" italics="true" pageId="5" pageNumber="74">Cf</emphasis>
cymbial furrow.
</paragraph>
<paragraph id="8B8A2F629664FFF5FF40FF70D908FF7B" blockId="6.[151,370,152,176]" box="[151,370,152,176]" pageId="6" pageNumber="75">Leg measurements:</paragraph>
<paragraph id="8B8A2F629664FFF5FEE5FF25D942FE33" blockId="6.[159,1296,204,391]" lastBlockId="6.[151,312,462,487]" pageId="6" pageNumber="75">Fe Pt Ti Mt Ta Total I 1.7 0.9 1.25 1.1 0.9 5.85 II 1.55 0.85 1.05 1.05 1.0 5.5 III 1.45 0.75 0.9 1.35 0.9 5.35 IV 2.15 1.0 1.5 2.25 1.1 8.0 Leg spination:</paragraph>
<paragraph id="8B8A2F629664FFF5FF35FDEBDC47FDCC" blockId="6.[159,1340,514,702]" box="[226,1085,514,536]" pageId="6" pageNumber="75">Fe Pt Ti Mt</paragraph>
<paragraph id="8B8A2F629664FFF5FF48FDC5DC18FD96" blockId="6.[159,1340,514,702]" box="[159,1122,556,578]" pageId="6" pageNumber="75">
I d1-1-0
<date id="FF8B09A29664FFF5FD4AFDC4DAB1FD96" box="[669,715,556,578]" pageId="6" pageNumber="75">v2-1</date>
-
<date id="FF8B09A29664FFF5FD05FDC4DC56FD96" box="[722,1068,556,578]" pageId="6" pageNumber="75">2 v</date>
0-2-2
</paragraph>
<paragraph id="8B8A2F629664FFF5FF48FDBEDC18FDBF" blockId="6.[159,1340,514,702]" box="[159,1122,597,619]" pageId="6" pageNumber="75">
II d1-1-0 p0-0-1 p0-0-
<date id="FF8B09A29664FFF5FD00FDBDDA8DFDBF" box="[727,759,597,619]" pageId="6" pageNumber="75">1 v</date>
1-1-
<date id="FF8B09A29664FFF5FCCCFDBDDC56FDBF" box="[795,1068,597,619]" pageId="6" pageNumber="75">2 v</date>
0-2-2
</paragraph>
<paragraph id="8B8A2F629664FFF5FF48FD97DB7BFCBE" blockId="6.[159,1340,514,702]" lastBlockId="6.[151,1437,777,913]" pageId="6" pageNumber="75">
III d1-1-1 p0-1-1 r0-1-1 p1-0 r1-0 d0-1-0 p1-1-1 r0-1-
<date id="FF8B09A29664FFF5FCB1FD96DBFEFD40" box="[870,900,638,660]" pageId="6" pageNumber="75">1 v</date>
2-2-2 d0-1-0 p1-1-0 r1-1-0 v0-2-
<date id="FF8B09A29664FFF5FAF8FD96D8C0FD6A" pageId="6" pageNumber="75">2 IV</date>
d1-1-0 p0-0-1 r0-0-1 r1-0 d1-0-1 p1-1-1 r1-1-
<date id="FF8B09A29664FFF5FCB2FD40DBFEFD6A" box="[869,900,680,702]" pageId="6" pageNumber="75">1 v</date>
2-2-2 d1-2-0 p1-1-0 r1-1-0 v0-2-2 Palp as in
<figureCitation id="130E33E79664FFF5FEE9FCE1D9C4FCF6" box="[318,446,777,802]" captionStart="FIGURES 9 ‒ 19" captionStartId="5.[151,264,1873,1896]" captionTargetBox="[151,1432,202,1852]" captionTargetId="figure@5.[151,1436,195,1852]" captionTargetPageId="5" captionText="FIGURES 9 ‒ 19. Habitus and copulatory organs of Berlandina litvinovi sp. n. 9 ‒ 10 female and male habitus, dorsal; 11 ‒ 13 male palp, prolateral, ventral and retrolateral respectively; 14 ‒ 16 bulb, anterior, ventral and retrolateral; 17 intact epigyne, ventral; 18 ‒ 19 macerated epigyne, ventral and dorsal. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569171/files/figure.png" pageId="6" pageNumber="75">Figs 11‒16</figureCitation>
; tibial apophysis large, sharply pointed, equal in length to tibia; cymbium with slanting prolateral furrow (
<emphasis id="B941F3709664FFF5FEA1FCC5D9EAFC91" box="[374,400,813,837]" italics="true" pageId="6" pageNumber="75">Cf</emphasis>
) against anterior edge of tibial apophysis; tegulum bulged (
<emphasis id="B941F3709664FFF5FBAEFCC6DCE2FC91" box="[1145,1176,814,837]" italics="true" pageId="6" pageNumber="75">Bu</emphasis>
) prolaterally; median apophysis longer than embolus; embolus with bifid tip.
</paragraph>
<paragraph id="8B8A2F629664FFF5FF10FC90DC41FC45" blockId="6.[151,1437,777,913]" box="[199,1083,888,913]" pageId="6" pageNumber="75">Female. Total length 7.4. Carapace: 2.4 long, 1.95 wide. Coloration as in male.</paragraph>
<paragraph id="8B8A2F629664FFF5FF40FC29D908FC0C" blockId="6.[151,370,961,985]" box="[151,370,961,985]" pageId="6" pageNumber="75">Leg measurements:</paragraph>
<paragraph id="8B8A2F629664FFF5FEF9FC1DD942FAC4" blockId="6.[159,1285,1012,1200]" lastBlockId="6.[151,312,1271,1296]" pageId="6" pageNumber="75">Fe Pt Ti Mt Ta Total I 1.8 0.95 1.2 1.1 0.95 6.0 II 1.55 0.85 1.05 1.05 0.9 5.4 III 1.45 0.8 0.9 1.3 0.9 5.35 IV 2.15 1.05 1.55 2.15 1.15 8.05 Leg spination:</paragraph>
<paragraph id="8B8A2F629664FFF5FF35FAC4DC3AFA95" blockId="6.[159,1343,1323,1510]" box="[226,1088,1323,1345]" pageId="6" pageNumber="75">Fe Pt Ti Mt</paragraph>
<paragraph id="8B8A2F629664FFF5FF48FABDDC1CFABE" blockId="6.[159,1343,1323,1510]" box="[159,1126,1364,1386]" pageId="6" pageNumber="75">
I d1-1-0 p0-0-
<date id="FF8B09A29664FFF5FEB1FABCDAD4FABE" box="[358,686,1364,1386]" pageId="6" pageNumber="75">1 v</date>
2-1-
<date id="FF8B09A29664FFF5FD03FABCDC4AFABE" box="[724,1072,1364,1386]" pageId="6" pageNumber="75">2 v</date>
0-2-2
</paragraph>
<paragraph id="8B8A2F629664FFF5FF48FA97DC1FFA40" blockId="6.[159,1343,1323,1510]" box="[159,1125,1406,1428]" pageId="6" pageNumber="75">
II d1-1-0 p0-0-
<date id="FF8B09A29664FFF5FEB1FA96DAD5FA40" box="[358,687,1406,1428]" pageId="6" pageNumber="75">1 v</date>
1-1-
<date id="FF8B09A29664FFF5FD04FA96DC55FA40" box="[723,1071,1406,1428]" pageId="6" pageNumber="75">2 v</date>
2-0-2
</paragraph>
<paragraph id="8B8A2F629664FFF5FF48FA40DC36F9BA" blockId="6.[159,1343,1323,1510]" lastBlockId="6.[151,1436,1585,1838]" pageId="6" pageNumber="75">
III d1-1-1 p0-1-1 r0-1-1 p1-0 r1-0 d1-0-0 p1-1-1 r0-1-
<date id="FF8B09A29664FFF5FCBFFA4FDBFCFA69" box="[872,902,1447,1469]" pageId="6" pageNumber="75">1 v</date>
2-2-2 d0-1-0 p1-1-0 r1-1-0
<date id="FF8B09A29664FFF5FB2CFA4FDD6CFA69" box="[1275,1302,1447,1469]" pageId="6" pageNumber="75">v2</date>
-0-
<date id="FF8B09A29664FFF5FAE5FA4FD8C0FA32" pageId="6" pageNumber="75">2 IV</date>
d1-1-0 p0-0-1 r0-0-1 r1-0 d1-0-1 p1-1-1 r1-1-
<date id="FF8B09A29664FFF5FCB0FA38DBFCFA32" box="[871,902,1488,1510]" pageId="6" pageNumber="75">1 v</date>
2-2-2 d1-2-0 p1-1-0 r1-1-0 v0-2-2 Epigyne as in
<figureCitation id="130E33E79664FFF5FEACF9D9DA78F99E" box="[379,514,1585,1610]" captionStart="FIGURES 9 ‒ 19" captionStartId="5.[151,264,1873,1896]" captionTargetBox="[151,1432,202,1852]" captionTargetId="figure@5.[151,1436,195,1852]" captionTargetPageId="5" captionText="FIGURES 9 ‒ 19. Habitus and copulatory organs of Berlandina litvinovi sp. n. 9 ‒ 10 female and male habitus, dorsal; 11 ‒ 13 male palp, prolateral, ventral and retrolateral respectively; 14 ‒ 16 bulb, anterior, ventral and retrolateral; 17 intact epigyne, ventral; 18 ‒ 19 macerated epigyne, ventral and dorsal. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569171/files/figure.png" pageId="6" pageNumber="75">Figs 17‒19</figureCitation>
; atrium oval shaped with weakly developed septum, twice wider than high; receptacles round, located posteriorly from atrium, spaced by less than one diameter.
</paragraph>
<paragraph id="8B8A2F629664FFF5FF10F995DD5AF96E" blockId="6.[151,1436,1585,1838]" pageId="6" pageNumber="75">
Size variations. Males vary from 5.0 to
<quantity id="4CCD82879664FFF5FD41F995DAA3F942" box="[662,729,1661,1686]" metricMagnitude="-1" metricUnit="m" metricValue="1.3716" pageId="6" pageNumber="75" unit="in" value="5.4">5.4 in</quantity>
the body length, carapace 2.05–2.25 long and 1.65–1.9 wide (n=4). Females vary from 5.0 to
<quantity id="4CCD82879664FFF5FDD2F94ADA3EF96E" box="[517,580,1697,1722]" metricMagnitude="-1" metricUnit="m" metricValue="1.8796" pageId="6" pageNumber="75" unit="in" value="7.4">7.4 in</quantity>
the body length, carapace 2.0–2.4 long and 1.6–1.95 wide (n=3).
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99664FFF5FF10F921DA8BF936" box="[199,753,1737,1762]" pageId="6" pageNumber="75" type="distribution">
<paragraph id="8B8A2F629664FFF5FF10F921DA8BF936" blockId="6.[151,1436,1585,1838]" box="[199,753,1737,1762]" pageId="6" pageNumber="75">
<emphasis id="B941F3709664FFF5FF10F921D91AF936" bold="true" box="[199,352,1737,1762]" pageId="6" pageNumber="75">Distribution.</emphasis>
Known from the
<typeStatus id="548E91C09664FFF5FDF0F922DA2DF936" box="[551,599,1738,1762]" pageId="6" pageNumber="75">type</typeStatus>
locality only.
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99664FFF5FF10F919DCDAF8FA" pageId="6" pageNumber="75" type="discussion">
<paragraph id="8B8A2F629664FFF5FF10F919DCDAF8FA" blockId="6.[151,1436,1585,1838]" pageId="6" pageNumber="75">
<emphasis id="B941F3709664FFF5FF10F919D935F8DE" bold="true" box="[199,335,1777,1802]" pageId="6" pageNumber="75">Comments.</emphasis>
Unlike most of
<taxonomicName id="4C3554E19664FFF5FDCEF919DAEDF8DD" box="[537,663,1777,1801]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B941F3709664FFF5FDCEF919DAEDF8DD" box="[537,663,1777,1801]" italics="true" pageId="6" pageNumber="75">Berlandina</emphasis>
</taxonomicName>
species, which inhabit arid habitats (steppe and various
<typeStatus id="548E91C09664FFF5FAE9F91ADD03F8DE" box="[1342,1401,1778,1802]" pageId="6" pageNumber="75">types</typeStatus>
of deserts), the new species is found also in the wet riparian forest subject to periodic flooding.
</paragraph>
</subSubSection>
</treatment>
</document>
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