219 lines
23 KiB
XML
219 lines
23 KiB
XML
<document id="0F5807B5BF621148CF17ABA0EFD9B914" ID-DOI="10.1649/0010-065X-73.1.127" ID-ISSN="1938-4394" ID-Zenodo-Dep="5333118" ID-ZooBank="urn:lsid:zoobank.org:pub:A5E82F63-4AC3-41BD-B1D7-78430C22A742" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630173649495" checkinUser="valdenar" docAuthor="López-García, Margarita M. & Deloya, Cuauhtémoc" docDate="2019" docId="6F6ED021D62CFFCBFD13FF2B7DA8FB68" docLanguage="en" docName="0010-065X-73.1.127.pdf" docOrigin="The Coleopterists Bulletin 73 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-73.1.127" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Tomarus maracaiboensis López-García & Deloya, 2019, new species" docType="treatment" docVersion="4" lastPageNumber="130" masterDocId="9357A859D628FFCEFFD0FFE87C4AFFFC" masterDocTitle="Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species" masterLastPageNumber="141" masterPageNumber="127" pageNumber="130" updateTime="1699174053468" updateUser="plazi">
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<mods:title id="492A583D8E3B9478969007B066B9C0E9">Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species</mods:title>
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<mods:namePart id="C4B163B6F21A240E282AE533AB59B4FB">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="82C30B4CC937B3538C4131967376CE03">Deloya, Cuauhtémoc</mods:namePart>
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<mods:title id="92F6D784DCFCDB0DD8E8C998741237F0">The Coleopterists Bulletin</mods:title>
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<mods:part id="DC9DB25262CCF2C1608E30EAC06F7267">
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<mods:date id="311EE44581F4A5A5DBACDFF93ED5C42F">2019</mods:date>
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<mods:number id="91E02F0E44D2A8D9B15155D00A52A00E">2019-03-25</mods:number>
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<mods:number id="02DC92606A1FD193F4A5C45B72EBB6F3">73</mods:number>
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<treatment id="6F6ED021D62CFFCBFD13FF2B7DA8FB68" ID-DOI="http://doi.org/10.5281/zenodo.5412051" ID-Zenodo-Dep="5412051" LSID="urn:lsid:plazi:treatment:6F6ED021D62CFFCBFD13FF2B7DA8FB68" httpUri="http://treatment.plazi.org/id/6F6ED021D62CFFCBFD13FF2B7DA8FB68" lastPageId="5" lastPageNumber="130" pageId="4" pageNumber="130">
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<subSubSection id="AFDD32BCD62CFFCAFD13FF2B7F9FFEBE" pageId="4" pageNumber="130" type="nomenclature">
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<paragraph id="E7786137D62CFFCAFD13FF2B784FFF0F" blockId="4.[689,1168,194,323]" pageId="4" pageNumber="130">
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<heading id="BC30D65BD62CFFCAFD13FF2B784FFF0F" centered="true" fontSize="9" level="2" pageId="4" pageNumber="130" reason="2">
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<emphasis id="D5B3BD25D62CFFCAFD13FF2B784FFF0F" bold="true" pageId="4" pageNumber="130">
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<taxonomicName id="20C71AB4D62CFFCAFD13FF2B7FF7FF24" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" box="[707,957,195,216]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="maracaiboensis">
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<emphasis id="D5B3BD25D62CFFCAFD13FF2B7FF7FF24" bold="true" box="[707,957,195,216]" italics="true" pageId="4" pageNumber="130">Tomarus maracaiboensis</emphasis>
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</taxonomicName>
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López-Garc´ıa and Deloya,
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<taxonomicNameLabel id="CE80005ED62CFFCAFC5FFF36784FFF0F" box="[911,1029,222,243]" pageId="4" pageNumber="130" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E7786137D62CFFCAFD3AFF107F9FFEBE" blockId="4.[689,1168,194,323]" pageId="4" pageNumber="130">
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Zoobank.org/
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urn:lsid:zoobank.org:act:
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<uuid id="93615BE2D62CFFCAFD61FEFB78DAFED4" box="[689,1168,275,296]" pageId="4" pageNumber="130">10BCDC71-EA26-4D32-9CAC-BABC1468FE9F</uuid>
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</uri>
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(
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<figureCitation id="7FFC7DB2D62CFFCAFCA3FEC57F84FEBE" box="[883,974,301,323]" captionStart="Figs" captionStartId="5.[197,236,987,1006]" captionTargetBox="[180,1188,196,966]" captionTargetId="figure-400@5.[175,1190,194,971]" captionTargetPageId="5" captionText="Figs. 7–9. Tomarus maracaiboensis. 7) Habitus; 8) Parameres, frontal view; 9) Parameres, lateral view. Scale bars = 2 mm (Fig. 7), 0.5 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.5333126" httpUri="https://zenodo.org/record/5333126/files/figure.png" pageId="4" pageNumber="130">Figs. 7–9</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="AFDD32BCD62CFFCAFD15FE8A7FB1FE50" pageId="4" pageNumber="130" type="materials_examined">
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<paragraph id="E7786137D62CFFCAFD15FE8A7FB1FE50" blockId="4.[685,1173,353,1730]" pageId="4" pageNumber="130">
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<emphasis id="D5B3BD25D62CFFCAFD15FE8A7F29FE8B" bold="true" box="[709,867,354,375]" pageId="4" pageNumber="130">Type Material.</emphasis>
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<materialsCitation id="57AF6B6AD62CFFCAFCA2FE8A7F3EFE6D" collectionCode="USNM" pageId="4" pageNumber="130" specimenCount="1" typeStatus="holotype">
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36.
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<typeStatus id="387CDF95D62CFFCAFC71FE8A7FB3FE8B" box="[929,1017,354,375]" pageId="4" pageNumber="130" type="holotype">Holotype</typeStatus>
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labeled “Maracaibo” (16
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<collectionCode id="81D6F9F2D62CFFCAFCF4FE947F27FE6D" box="[804,877,380,401]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="130" type="Museum">USNM</collectionCode>
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)
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</materialsCitation>
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||
.
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<materialsCitation id="57AF6B6AD62CFFCAFCACFE947FB2FE50" collectionCode="USNM, UNSM" pageId="4" pageNumber="130" specimenCount="1" typeStatus="paratype">
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<typeStatus id="387CDF95D62CFFCAFCACFE947F91FE6D" box="[892,987,380,401]" pageId="4" pageNumber="130" type="paratype">Paratypes</typeStatus>
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: Same label data as holotype (16
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<collectionCode id="81D6F9F2D62CFFCAFCE3FE7F7F30FE50" box="[819,890,407,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="130" type="Museum">USNM</collectionCode>
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, 16
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<collectionCode id="81D6F9F2D62CFFCAFC78FE7F7FBBFE50" box="[936,1009,407,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="4" pageNumber="130">UNSM</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="AFDD32BCD62CFFCAFD15FE5A7FFDFB52" pageId="4" pageNumber="130" type="description">
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<paragraph id="E7786137D62CFFCAFD15FE5A7F82FBA3" blockId="4.[685,1173,353,1730]" pageId="4" pageNumber="130">
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<emphasis id="D5B3BD25D62CFFCAFD15FE5A7FE4FE3B" bold="true" box="[709,942,434,455]" pageId="4" pageNumber="130">
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Description.
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<typeStatus id="387CDF95D62CFFCAFC9CFE5A7FE0FE3B" box="[844,938,434,455]" pageId="4" pageNumber="130" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Habitus as in
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<figureCitation id="7FFC7DB2D62CFFCAFB80FE5A78DAFE3B" box="[1104,1168,434,455]" captionStart="Figs" captionStartId="5.[197,236,987,1006]" captionTargetBox="[180,1188,196,966]" captionTargetId="figure-400@5.[175,1190,194,971]" captionTargetPageId="5" captionText="Figs. 7–9. Tomarus maracaiboensis. 7) Habitus; 8) Parameres, frontal view; 9) Parameres, lateral view. Scale bars = 2 mm (Fig. 7), 0.5 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.5333126" httpUri="https://zenodo.org/record/5333126/files/figure.png" pageId="4" pageNumber="130">Fig. 7</figureCitation>
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. Length
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<quantity id="203FCCD2D62CFFCAFD2CFE247F10FE1D" box="[764,858,460,481]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="4" pageNumber="130" unit="mm" value="23.3">23.3 mm</quantity>
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||
; width across humeri
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||
<quantity id="203FCCD2D62CFFCAFBEFFE2478DBFE1D" box="[1087,1169,460,481]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="130" unit="mm" value="2.2">2.2 mm</quantity>
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||
. Color brown to black, venter slightly lighter.
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<emphasis id="D5B3BD25D62CFFCAFB86FE0F78D9FE00" bold="true" box="[1110,1171,487,508]" pageId="4" pageNumber="130">Head:</emphasis>
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Frons coarsely rugose. Frontoclypeal region with 2 triangular tubercles separated by about 7 tubercle diameters. Clypeus narrowed towards apex, base 3 times as wide as apex. Clypeal teeth triangular, separated by 1 tooth diameter. Mandibles with 2 apical, acute teeth and a rounded basal lobe. Interocular distance equal to 3.3 times eye width. Antennal club short, subequal in length to antennomeres 2–7.
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<emphasis id="D5B3BD25D62CFFCAFCE6FD3E7FEEFD17" bold="true" box="[822,932,726,747]" pageId="4" pageNumber="130">Pronotum:</emphasis>
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Surface smooth on disc, with minute punctures on lateral margins. Apical tubercle small and rounded. Fovea shallow, striate, and about half as wide as interocular distance.
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<emphasis id="D5B3BD25D62CFFCAFD7DFCA87EBFFCA9" bold="true" box="[685,757,832,853]" pageId="4" pageNumber="130">Elytra:</emphasis>
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First interval punctate, punctures smaller than on other intervals.
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<emphasis id="D5B3BD25D62CFFCAFC44FCB37FB7FC8C" bold="true" box="[916,1021,859,880]" pageId="4" pageNumber="130">Pygidium:</emphasis>
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Surface rugose on basal fourth and lateral angles, disc with minute punctures. Slightly convex in lateral view. Apex regularly rounded.
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<emphasis id="D5B3BD25D62CFFCAFCB2FC427FD3FC43" bold="true" box="[866,921,938,959]" pageId="4" pageNumber="130">Legs:</emphasis>
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Protibia tridentate with an additional small basal tooth. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia not narrowed before apex, sides nearly parallel. Apex of metatibia crenulate, with 8 spinules.
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<emphasis id="D5B3BD25D62CFFCAFBCEFBFD78D9FBD6" bold="true" box="[1054,1171,1045,1066]" pageId="4" pageNumber="130">Parameres:</emphasis>
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With 2 pairs of dorsal lateral teeth, both long and upwardly arched (
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<figureCitation id="7FFC7DB2D62CFFCAFC8EFBA27FF6FBA3" box="[862,956,1097,1119]" captionStart="Figs" captionStartId="5.[197,236,987,1006]" captionTargetBox="[180,1188,196,966]" captionTargetId="figure-400@5.[175,1190,194,971]" captionTargetPageId="5" captionText="Figs. 7–9. Tomarus maracaiboensis. 7) Habitus; 8) Parameres, frontal view; 9) Parameres, lateral view. Scale bars = 2 mm (Fig. 7), 0.5 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.5333126" httpUri="https://zenodo.org/record/5333126/files/figure.png" pageId="4" pageNumber="130">Figs. 8–9</figureCitation>
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).
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</paragraph>
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<paragraph id="E7786137D62CFFCAFD15FB8C7FFDFB52" blockId="4.[685,1173,353,1730]" pageId="4" pageNumber="130">
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<emphasis id="D5B3BD25D62CFFCAFD15FB8C7F62FB85" bold="true" box="[709,808,1124,1145]" pageId="4" pageNumber="130">Variation.</emphasis>
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Male (
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<specimenCount id="F1C1AABED62CFFCAFCBDFB8C7F97FB85" box="[877,989,1124,1145]" pageId="4" pageNumber="130" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
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): Length
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<quantity id="203FCCD2D62CFFCAFBE7FB8C7E9EFB68" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.11" metricValueMin="2.09" pageId="4" pageNumber="130" unit="mm" value="21.0" valueMax="21.1" valueMin="20.9">20.9–21.1 mm</quantity>
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; width across humeri 10.0–
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<quantity id="203FCCD2D62CFFCAFC30FB977873FB68" box="[992,1081,1151,1172]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="4" pageNumber="130" unit="mm" value="12.3">12.3 mm</quantity>
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. Apex of metatibia with 8–9 spinules.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="AFDD32BCD62CFFCAFD15FB5C7F05FB18" pageId="4" pageNumber="130" type="etymology">
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||
<paragraph id="E7786137D62CFFCAFD15FB5C7F05FB18" blockId="4.[685,1173,353,1730]" pageId="4" pageNumber="130">
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||
<emphasis id="D5B3BD25D62CFFCAFD15FB5C7F72FB35" bold="true" box="[709,824,1204,1225]" pageId="4" pageNumber="130">Etymology.</emphasis>
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The name
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<taxonomicName id="20C71AB4D62CFFCAFC76FB5C7876FB35" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" box="[934,1084,1204,1225]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="maracaiboensis">
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||
<emphasis id="D5B3BD25D62CFFCAFC76FB5C7876FB35" box="[934,1084,1204,1225]" italics="true" pageId="4" pageNumber="130">maracaiboensis</emphasis>
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</taxonomicName>
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||
refers to the
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<typeStatus id="387CDF95D62CFFCAFD02FB277EB1FB18" box="[722,763,1231,1252]" pageId="4" pageNumber="130">type</typeStatus>
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locality.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="AFDD32BCD62CFFCAFD15FB017F89FACF" pageId="4" pageNumber="130" type="distribution">
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||
<paragraph id="E7786137D62CFFCAFD15FB017F89FACF" blockId="4.[685,1173,353,1730]" pageId="4" pageNumber="130">
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||
<emphasis id="D5B3BD25D62CFFCAFD15FB017F0CFB02" bold="true" box="[709,838,1257,1278]" pageId="4" pageNumber="130">Distribution.</emphasis>
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||
<taxonomicName id="20C71AB4D62CFFCAFC9BFB017873FB02" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" box="[843,1081,1257,1278]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="maracaiboensis">
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||
<emphasis id="D5B3BD25D62CFFCAFC9BFB017873FB02" box="[843,1081,1257,1278]" italics="true" pageId="4" pageNumber="130">Tomarus maracaiboensis</emphasis>
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</taxonomicName>
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is known only from Maracaibo, a municipality in northwestern
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<collectingCountry id="9FD021A7D62CFFCAFD2EFAF67F15FACF" box="[766,863,1310,1331]" name="Venezuela" pageId="4" pageNumber="130">Venezuela</collectingCountry>
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||
(
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<figureCitation id="7FFC7DB2D62CFFCAFCBFFAF67FFDFACF" box="[879,951,1310,1331]" captionStart="Fig" captionStartId="9.[197,228,918,937]" captionTargetBox="[177,1188,196,901]" captionTargetId="figure-402@9.[175,1190,194,903]" captionTargetPageId="9" captionText="Fig. 23. Known localities of five newly described species of Tomarus from Central and South America." figureDoi="http://doi.org/10.5281/zenodo.5333140" httpUri="https://zenodo.org/record/5333140/files/figure.png" pageId="4" pageNumber="130">Fig. 23</figureCitation>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="AFDD32BCD62CFFCAFD15FAD17FE1F9F4" pageId="4" pageNumber="130" type="diagnosis">
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||
<paragraph id="E7786137D62CFFCAFD15FAD17FE1F9F4" blockId="4.[685,1173,353,1730]" pageId="4" pageNumber="130">
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||
<emphasis id="D5B3BD25D62CFFCAFD15FAD17F67FAB2" bold="true" box="[709,813,1337,1358]" pageId="4" pageNumber="130">Diagnosis.</emphasis>
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This species can be recognized by the nearly impunctate pronotum, small pronotal tubercle and fovea, and tridentate protibia with an additional fourth basal tooth. It is similar to
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<taxonomicName id="20C71AB4D62CFFCAFB52FA617F44FA44" baseAuthorityName="Bates" baseAuthorityYear="1888" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="laevicollis">
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<emphasis id="D5B3BD25D62CFFCAFB52FA617F44FA44" italics="true" pageId="4" pageNumber="130">T. laevicollis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="20C71AB4D62CFFCAFCCEFA4B7FF8FA44" authorityName="Lopez-Garcia & Gasca-Alvarez" authorityYear="2014" box="[798,946,1443,1464]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="colombianus">
|
||
<emphasis id="D5B3BD25D62CFFCAFCCEFA4B7FF8FA44" box="[798,946,1443,1464]" italics="true" pageId="4" pageNumber="130">T. colombianus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="20C71AB4D62CFFCAFC3EFA4B7826FA44" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" box="[1006,1132,1443,1464]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="laticaudus">
|
||
<emphasis id="D5B3BD25D62CFFCAFC3EFA4B7826FA44" box="[1006,1132,1443,1464]" italics="true" pageId="4" pageNumber="130">T. laticaudus</emphasis>
|
||
</taxonomicName>
|
||
but can be differentiated by the parameres with a very short apex and the apical teeth similar in shape and as long as the basal tooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AFDD32BCD62CFFCBFD15F9E57DA8FB68" lastPageId="5" lastPageNumber="131" pageId="4" pageNumber="130" type="discussion">
|
||
<paragraph id="E7786137D62CFFCBFD15F9E57DA8FB68" blockId="4.[685,1173,353,1730]" lastBlockId="5.[173,659,1098,1172]" lastPageId="5" lastPageNumber="131" pageId="4" pageNumber="130">
|
||
<emphasis id="D5B3BD25D62CFFCAFD15F9E57F72F9DE" bold="true" box="[709,824,1549,1570]" pageId="4" pageNumber="130">Comments.</emphasis>
|
||
<taxonomicName id="20C71AB4D62CFFCAFC94F9E5784FF9DE" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" box="[836,1029,1549,1570]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="laticaudus">
|
||
<emphasis id="D5B3BD25D62CFFCAFC94F9E5784FF9DE" box="[836,1029,1549,1570]" italics="true" pageId="4" pageNumber="130">Tomarus laticaudus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="20C71AB4D62CFFCAFBC6F9E57E84F9C1" authorityName="Lopez-Garcia & Gasca-Alvarez" authorityYear="2014" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="colombianus">
|
||
<emphasis id="D5B3BD25D62CFFCAFBC6F9E57E84F9C1" italics="true" pageId="4" pageNumber="130">T. colombianus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="20C71AB4D62CFFCAFCD5F9C07FF9F9C1" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" box="[773,947,1576,1597]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="maracaiboensis">
|
||
<emphasis id="D5B3BD25D62CFFCAFCD5F9C07FF9F9C1" box="[773,947,1576,1597]" italics="true" pageId="4" pageNumber="130">T. maracaiboensis</emphasis>
|
||
</taxonomicName>
|
||
are found at elevations below
|
||
<specimenCount id="F1C1AABED62CFFCAFD3FF9AB7F7EF9A4" box="[751,820,1603,1624]" pageId="4" pageNumber="130" type="generic">
|
||
<quantity id="203FCCD2D62CFFCAFD3FF9AB7F7AF9A4" box="[751,816,1603,1624]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="130" unit="m" value="100.0">100 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
In particular,
|
||
<taxonomicName id="20C71AB4D62CFFCAFC6EF9AB781AF9A4" authorityName="Lopez-Garcia & Gasca-Alvarez" authorityYear="2014" box="[958,1104,1603,1624]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="colombianus">
|
||
<emphasis id="D5B3BD25D62CFFCAFC6EF9AB781AF9A4" box="[958,1104,1603,1624]" italics="true" pageId="4" pageNumber="130">T. colombianus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="20C71AB4D62CFFCAFB52F9AB7F09F98E" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="maracaiboensis">
|
||
<emphasis id="D5B3BD25D62CFFCAFB52F9AB7F09F98E" italics="true" pageId="4" pageNumber="130">T. maracaiboensis</emphasis>
|
||
</taxonomicName>
|
||
, which are the most similar in external appearance, are from coastal areas in South America. However, they occur in very distinct regions.
|
||
<taxonomicName id="20C71AB4D62CFFCAFD3CF9457FF0F93E" authorityName="Lopez-Garcia & Gasca-Alvarez" authorityYear="2014" box="[748,954,1709,1730]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="colombianus">
|
||
<emphasis id="D5B3BD25D62CFFCAFD3CF9457FF0F93E" box="[748,954,1709,1730]" italics="true" pageId="4" pageNumber="130">Tomarus colombianus</emphasis>
|
||
</taxonomicName>
|
||
lives in moist forest in the Biogeographic Chocó with extremely high levels of precipitation, whereas
|
||
<taxonomicName id="20C71AB4D62DFFCBFE4CFB8C7E0DFB85" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" box="[412,583,1124,1145]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="131" phylum="Arthropoda" rank="species" species="maracaiboensis">
|
||
<emphasis id="D5B3BD25D62DFFCBFE4CFB8C7E0DFB85" box="[412,583,1124,1145]" italics="true" pageId="5" pageNumber="131">T. maracaiboensis</emphasis>
|
||
</taxonomicName>
|
||
is from dry forest in northern
|
||
<collectingCountry id="9FD021A7D62DFFCBFEACFB977D94FB68" box="[380,478,1151,1172]" name="Venezuela" pageId="5" pageNumber="131">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |