227 lines
37 KiB
XML
227 lines
37 KiB
XML
<document id="E1F3E09EB043E5C8BD16B6BFA06E6A1F" ID-CLB-Dataset="8703" ID-DOI="10.11646/zootaxa.4825.1.1" ID-GBIF-Dataset="5ab5f5c8-481e-4d1a-8643-21e72c367278" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402145" ID-ZooBank="F230F199-1C94-4E2E-9CE4-5F56212C015F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609341789210" checkinUser="plazi" docAuthor="Schileyko, Arkady A., Vahtera, Varpu & Edgecombe, Gregory D." docDate="2020" docId="03DE092DFFFBD700FF13FAD12C39DEF4" docLanguage="en" docName="zootaxa.4825.1.1.pdf" docOrigin="Zootaxa 4825 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Kethopinae , Shelley 2002" docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="FFE77155FFF2D70AFF84FFE12D45DC1F" masterDocTitle="An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses" masterLastPageNumber="64" masterPageNumber="1" pageNumber="10" updateTime="1698852004925" updateUser="ExternalLinkService">
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<mods:mods id="7059841656F55C08B355E7073D813207" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FEB35C2A38C205D7D8B2FD23092AF900">
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<mods:title id="CD5975FE6E6353B12FDAD2AFE265A332">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:name id="28D488942FA2E3F7EE7027325CA74E0A" type="personal">
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<mods:roleTerm id="ACC97A11623ABD74957856FEEE557C7B">Author</mods:roleTerm>
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<mods:namePart id="1AEFA05C8B705CCAEB4A90C2D136896D">Schileyko, Arkady A.</mods:namePart>
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<mods:nameIdentifier id="868EA7FC80D93E849CEB41DFDC4894AD" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="2C8782571A5707D1E1AE06691510708D">Author</mods:roleTerm>
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<mods:namePart id="21C470055C86F33DA85CDCD4982ACFA2">Vahtera, Varpu</mods:namePart>
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<mods:nameIdentifier id="BF47AA9EF8D43A4AE7CE432E0E8DA696" type="email">varpu.vahtera@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F06CD9A298878D5671852F26F8B8FD8A">Edgecombe, Gregory D.</mods:namePart>
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<mods:nameIdentifier id="32D5765BA2A15697CA975ADBDDCFACDA" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:title id="F87A8D775C3DBECEC2BFCE43A65B5E6D">Zootaxa</mods:title>
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<mods:part id="98A7929F962AFF1BBC5B9912ADCFF5B5">
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<mods:date id="5D1DFC26B546EB56EFC0EF4DD10B5E7D">2020</mods:date>
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<mods:number id="DD90625EEDDBD7863FF45ABD58338594">2020-08-10</mods:number>
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<treatment id="03DE092DFFFBD700FF13FAD12C39DEF4" ID-DOI="http://doi.org/10.5281/zenodo.4455377" ID-GBIF-Taxon="177020702" ID-Zenodo-Dep="4455377" LSID="urn:lsid:plazi:treatment:03DE092DFFFBD700FF13FAD12C39DEF4" httpUri="http://treatment.plazi.org/id/03DE092DFFFBD700FF13FAD12C39DEF4" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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<subSubSection id="C36DEBB0FFFBD703FF13FAD12F1CD955" box="[151,601,1328,1354]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BC8B83BFFFBD703FF13FAD12F1CD955" blockId="9.[151,601,1328,1354]" box="[151,601,1328,1354]" pageId="9" pageNumber="10">
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<heading id="D0800F57FFFBD703FF13FAD12F1CD955" bold="true" box="[151,601,1328,1354]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="3">
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<emphasis id="B9036429FFFBD703FF13FAD12F1CD955" bold="true" box="[151,601,1328,1354]" pageId="9" pageNumber="10">
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Subfamily
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<taxonomicName id="4C77C3B8FFFBD703FE9BFAD12F1CD955" authority="Shelley, 2002" authorityName=", Shelley" authorityYear="2002" box="[287,601,1328,1354]" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Kethopinae">
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Kethopinae
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<bibRefCitation id="EFE6C5CAFFFBD703FE3CFAD12F1CD955" author="Shelley, R. M." box="[440,601,1328,1354]" pageId="9" pageNumber="10" pagination="1 - 108" refId="ref50598" refString="Shelley, R. M. (2002) A synopsis of the North American centipedes of the order Scolopendromorpha (Chilopoda). Virginia Museum of Natural History, 5, 1 - 108." type="journal article" year="2002">Shelley, 2002</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFFBD703FF13FA992985DAE5" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="8BC8B83BFFFBD703FF13FA992E1EDA91" blockId="9.[151,1437,1400,2039]" pageId="9" pageNumber="10">
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<emphasis id="B9036429FFFBD703FF13FA992C57D98D" bold="true" box="[151,274,1400,1426]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
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Antenna with normal trichoid sensilla, lacking collared sensilla. Anterior margin of forcipular trochantero-prefemur weakly sclerotized, without processes. LBS 7 without spiracles. Coxopleuron with a relatively short conical process. Ultimate legs cryptopiform sensu
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<bibRefCitation id="EFE6C5CAFFFBD703FD5AFA212EC4D9C4" author="Crabill, R. E. Jr." box="[734,897,1472,1499]" pageId="9" pageNumber="10" pagination="1 - 15" refId="ref46778" refString="Crabill, R. E. Jr. (1960) A new American genus of cryptopid centipedes, with an annotated key to the scolopendromorph genera of America North of Mexico. Proceedings of the United States National Museum, 111, 1 - 15." type="journal article" year="1960">Crabill (1960)</bibRefCitation>
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(in shape of a “pocket knife” sensu
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<bibRefCitation id="EFE6C5CAFFFBD703FAA8FA212D94D9E1" author="Schileyko, A." pageId="9" pageNumber="10" pagination="519 - 530" refId="ref50114" refString="Schileyko, A. (2009) Ectonocryptoides sandrops - a new scolopendromorph centipede from Belize. Soil Organisms, 81 (3), 519 - 530." type="journal article" year="2009">Schileyko 2009</bibRefCitation>
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) i.e. shortened tibia and tarsus 1 with characteristic saw teeth, both these podomeres capable of flexure against each other and tarsus 2 forming a kind of clasping apparatus (
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<figureCitation id="134CA4BEFFFBD703FCE7F9E92EADDA3D" box="[867,1000,1544,1570]" captionStart="FIGURES 7–13" captionStartId="11.[151,264,1641,1665]" captionTargetBox="[289,1300,194,1595]" captionTargetId="figure-17@11.[276,1312,181,1618]" captionTargetPageId="11" captionText="FIGURES 7–13. Scolopocryptops melanostoma Newport, 1845; Ad. Rc 7172 7 LBS 23 and ultimate legs, laterally; Ad. Rc 7503 8 anterior margin of forcipular coxosternite + maxillae 2, ventrally; Scolopocryptops ferrugineus (L., 1767); Peru specimen from Markus Koch collection 9 forcipular segment ventrally; Scolopocryptops miersii Newport, 1845; BMNH, BM 1889.4.13.3–5 10 apical part of maxilla 2 laterally; Kethops utahensis (Chamberlin, 1909) 11 figures 4–6 of Chamberlin (1912); Thalkethops grallatrix Crabill, 1960 12 figure 15 of Crabill (1960); Kethops utahensis (Chamberlin, 1909); USNM Mill Creek (Utah) specimen 13 apical part of maxilla 2 ventro-laterally; (20)—leg 20, (am)—anterior margin of forcipular coxosternite, (cp)—coxopleural process, (cst)—forcipular coxosternite, (dms)—dorso-medial spinous process of ultimate prefemur, (p2)— pretarsus of maxilla 2, (pa)—short projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (pla)—pretarsal lamellae, (pt)—process of forcipular trochantero-prefemur, (rl)—right[?] ultimate leg, (S)—? sternite, (st)—saw teeth-like small spinous processes, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)— ultimate tibia, (us)—ultimate LBS, (ust)—ultimate sternite, (vsp)—ventral spinous process of ultimate prefemur." figureDoi="http://doi.org/10.5281/zenodo.4402149" httpUri="https://zenodo.org/record/4402149/files/figure.png" pageId="9" pageNumber="10">Figs 11, 12</figureCitation>
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). According to
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<bibRefCitation id="EFE6C5CAFFFBD703FB18F9E92807DA3C" author="Lewis, J. G. E." box="[1180,1346,1544,1571]" pageId="9" pageNumber="10" pagination="21 - 31" refId="ref49313" refString="Lewis, J. G. E. (2016 b) On the consistency of some taxonomic characters in the Scolopendromorpha and comments on the scolopocryptopid subfamily Kethopinae (Myriapoda: Chilopoda). Acta Societatis Zoologicae Bohemicae, 80, 21 - 31." type="journal article" year="2016">Lewis (2016b)</bibRefCitation>
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at least ultimate tibia and tarsus 1 (in two of three species prefemur and femur as well) bear numerous, small and strongly curved spinous processes (
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<figureCitation id="134CA4BEFFFBD703FE42F9B12F5CDA75" box="[454,537,1616,1642]" captionStart="FIGURES 7–13" captionStartId="11.[151,264,1641,1665]" captionTargetBox="[289,1300,194,1595]" captionTargetId="figure-17@11.[276,1312,181,1618]" captionTargetPageId="11" captionText="FIGURES 7–13. Scolopocryptops melanostoma Newport, 1845; Ad. Rc 7172 7 LBS 23 and ultimate legs, laterally; Ad. Rc 7503 8 anterior margin of forcipular coxosternite + maxillae 2, ventrally; Scolopocryptops ferrugineus (L., 1767); Peru specimen from Markus Koch collection 9 forcipular segment ventrally; Scolopocryptops miersii Newport, 1845; BMNH, BM 1889.4.13.3–5 10 apical part of maxilla 2 laterally; Kethops utahensis (Chamberlin, 1909) 11 figures 4–6 of Chamberlin (1912); Thalkethops grallatrix Crabill, 1960 12 figure 15 of Crabill (1960); Kethops utahensis (Chamberlin, 1909); USNM Mill Creek (Utah) specimen 13 apical part of maxilla 2 ventro-laterally; (20)—leg 20, (am)—anterior margin of forcipular coxosternite, (cp)—coxopleural process, (cst)—forcipular coxosternite, (dms)—dorso-medial spinous process of ultimate prefemur, (p2)— pretarsus of maxilla 2, (pa)—short projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (pla)—pretarsal lamellae, (pt)—process of forcipular trochantero-prefemur, (rl)—right[?] ultimate leg, (S)—? sternite, (st)—saw teeth-like small spinous processes, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)— ultimate tibia, (us)—ultimate LBS, (ust)—ultimate sternite, (vsp)—ventral spinous process of ultimate prefemur." figureDoi="http://doi.org/10.5281/zenodo.4402149" httpUri="https://zenodo.org/record/4402149/files/figure.png" pageId="9" pageNumber="10">Fig. 12</figureCitation>
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) which are similar (and, apparently, homologous) to the typical cryptopid-type saw teeth of tibia and tarsus 1. Ultimate pretarsus claw-shaped.
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</paragraph>
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<paragraph id="8BC8B83BFFFBD703FF43F9792F67DAAC" blockId="9.[151,1437,1400,2039]" box="[199,546,1688,1715]" pageId="9" pageNumber="10">
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<emphasis id="B9036429FFFBD703FF43F9792CF1DAAD" bold="true" box="[199,436,1688,1714]" pageId="9" pageNumber="10">Number of subtaxa.</emphasis>
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2 genera.
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</paragraph>
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<paragraph id="8BC8B83BFFFBD703FF43F95D2F71DAC9" blockId="9.[151,1437,1400,2039]" box="[199,564,1724,1750]" pageId="9" pageNumber="10">
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<emphasis id="B9036429FFFBD703FF43F95D2CF2DAC9" bold="true" box="[199,439,1724,1750]" pageId="9" pageNumber="10">Sexual dimorphism.</emphasis>
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Unknown.
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</paragraph>
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<paragraph id="8BC8B83BFFFBD703FF43F9012985DAE5" blockId="9.[151,1437,1400,2039]" box="[199,1216,1760,1787]" pageId="9" pageNumber="10">
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<emphasis id="B9036429FFFBD703FF43F9012C5FDAE5" bold="true" box="[199,282,1760,1786]" pageId="9" pageNumber="10">Range.</emphasis>
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<materialsCitation id="3B1FB266FFFBD703FEA5F90129F9DAE5" ID-GBIF-Occurrence="3023417313" box="[289,1212,1760,1787]" country="United States of America" location="California" pageId="9" pageNumber="10" specimenCount="1" stateProvince="California">
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Central
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<collectingRegion id="49B376D9FFFBD703FEFFF9012CAADAE5" box="[379,495,1760,1786]" country="United States of America" name="California" pageId="9" pageNumber="10">California</collectingRegion>
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, Northern
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<collectingRegion id="49B376D9FFFBD703FDE2F9012FDADAE5" box="[614,671,1760,1786]" country="United States of America" name="Utah" pageId="9" pageNumber="10">Utah</collectingRegion>
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, Northern and Southern
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<collectingRegion id="49B376D9FFFBD703FC37F9002901DAE5" box="[947,1092,1760,1787]" country="United States of America" name="New Mexico" pageId="9" pageNumber="10">New Mexico</collectingRegion>
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(all
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<collectingCountry id="F360F8ABFFFBD703FBF3F90129F1DAE5" box="[1143,1204,1760,1786]" name="United States of America" pageId="9" pageNumber="10">USA</collectingCountry>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFFBD700FF43F8E52C56DDB2" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
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<paragraph id="8BC8B83BFFFBD700FF43F8E52E10DD02" blockId="9.[151,1437,1400,2039]" lastBlockId="10.[151,1437,151,285]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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<emphasis id="B9036429FFFBD703FF43F8E52C7EDB01" bold="true" box="[199,315,1796,1822]" pageId="9" pageNumber="10">Remarks.</emphasis>
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Treated as a subfamily in
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<bibRefCitation id="EFE6C5CAFFFBD703FDFCF8E5294EDB00" author="Edgecombe, G. D. & Bonato, L." box="[632,1035,1796,1823]" pageId="9" pageNumber="10" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 405)</bibRefCitation>
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,
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<bibRefCitation id="EFE6C5CAFFFBD703FB9DF8E5280DDB00" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[1049,1352,1796,1823]" pageId="9" pageNumber="10" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
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Vahtera
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<emphasis id="B9036429FFFBD703FBFDF8E429F0DB01" box="[1145,1205,1796,1822]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
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(2013: 595)
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</bibRefCitation>
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,
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<bibRefCitation id="EFE6C5CAFFFBD703FADCF8E52C67DB5C" author="Lewis, J. G. E." pageId="9" pageNumber="10" pagination="21 - 31" refId="ref49313" refString="Lewis, J. G. E. (2016 b) On the consistency of some taxonomic characters in the Scolopendromorpha and comments on the scolopocryptopid subfamily Kethopinae (Myriapoda: Chilopoda). Acta Societatis Zoologicae Bohemicae, 80, 21 - 31." type="journal article" year="2016">Lewis (2016b: 26)</bibRefCitation>
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.
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<bibRefCitation id="EFE6C5CAFFFBD703FEB4F8C92F85DB5C" author="Edgecombe, G. D. & Bonato, L." box="[304,704,1832,1859]" pageId="9" pageNumber="10" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 405)</bibRefCitation>
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erroneously stated that the ultimate prefemur in
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<taxonomicName id="4C77C3B8FFFBD703FB7AF8C9283ADB5D" authorityName=", Shelley" authorityYear="2002" box="[1278,1407,1832,1858]" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Kethopinae">Kethopinae</taxonomicName>
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is “unarmed” (i.e. bears no spinous processes), but saw teeth are borne on the prefemur of both
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<taxonomicName id="4C77C3B8FFFBD703FB4AF8AD2C31DB95" authority="(Chamberlin, 1909)" baseAuthorityName="Chamberlin" baseAuthorityYear="1909" class="Chilopoda" family="Scolopocryptopidae" genus="Kethops" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="utahensis">
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<emphasis id="B9036429FFFBD703FB4AF8AD28D9DB79" box="[1230,1436,1868,1894]" italics="true" pageId="9" pageNumber="10">Kethops utahensis</emphasis>
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(Chamberlin, 1909)
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</taxonomicName>
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and
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<taxonomicName id="4C77C3B8FFFBD703FE2DF8912E04DB95" authority="Crabill, 1960" authorityName="Crabill" authorityYear="1960" box="[425,833,1904,1930]" class="Chilopoda" family="Scolopocryptopidae" genus="Thalkethops" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="grallatrix">
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<emphasis id="B9036429FFFBD703FE2DF8912FE1DB95" box="[425,676,1904,1930]" italics="true" pageId="9" pageNumber="10">Thalkethops grallatrix</emphasis>
|
||
<bibRefCitation id="EFE6C5CAFFFBD703FD2FF8912E04DB95" author="Crabill, R. E. Jr." box="[683,833,1904,1930]" pageId="9" pageNumber="10" pagination="1 - 15" refId="ref46778" refString="Crabill, R. E. Jr. (1960) A new American genus of cryptopid centipedes, with an annotated key to the scolopendromorph genera of America North of Mexico. Proceedings of the United States National Museum, 111, 1 - 15." type="journal article" year="1960">Crabill, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
. The most recent account on this subfamily was given by
|
||
<bibRefCitation id="EFE6C5CAFFFBD703FF3AF8752CFADBB0" author="Lewis, J. G. E." box="[190,447,1940,1967]" pageId="9" pageNumber="10" pagination="21 - 31" refId="ref49313" refString="Lewis, J. G. E. (2016 b) On the consistency of some taxonomic characters in the Scolopendromorpha and comments on the scolopocryptopid subfamily Kethopinae (Myriapoda: Chilopoda). Acta Societatis Zoologicae Bohemicae, 80, 21 - 31." type="journal article" year="2016">Lewis (2016b: 26–28)</bibRefCitation>
|
||
who overviewed the scant available literature and wrote on page 28 that apart from
|
||
<bibRefCitation id="EFE6C5CAFFFBD703FF13F8592CC5DBCD" author="Chamberlin, R. V." box="[151,384,1976,2002]" pageId="9" pageNumber="10" pagination="141 - 172" refId="ref46640" refString="Chamberlin, R. V. (1912) New North American chilopods and diplopods. Annals of the Entomological Society of America, 5, 141 - 172." type="journal article" year="1912">Chamberlin’s (1912)</bibRefCitation>
|
||
“puzzling” drawing (
|
||
<figureCitation id="134CA4BEFFFBD703FDF6F8592F86DBCD" box="[626,707,1976,2002]" captionStart="FIGURES 7–13" captionStartId="11.[151,264,1641,1665]" captionTargetBox="[289,1300,194,1595]" captionTargetId="figure-17@11.[276,1312,181,1618]" captionTargetPageId="11" captionText="FIGURES 7–13. Scolopocryptops melanostoma Newport, 1845; Ad. Rc 7172 7 LBS 23 and ultimate legs, laterally; Ad. Rc 7503 8 anterior margin of forcipular coxosternite + maxillae 2, ventrally; Scolopocryptops ferrugineus (L., 1767); Peru specimen from Markus Koch collection 9 forcipular segment ventrally; Scolopocryptops miersii Newport, 1845; BMNH, BM 1889.4.13.3–5 10 apical part of maxilla 2 laterally; Kethops utahensis (Chamberlin, 1909) 11 figures 4–6 of Chamberlin (1912); Thalkethops grallatrix Crabill, 1960 12 figure 15 of Crabill (1960); Kethops utahensis (Chamberlin, 1909); USNM Mill Creek (Utah) specimen 13 apical part of maxilla 2 ventro-laterally; (20)—leg 20, (am)—anterior margin of forcipular coxosternite, (cp)—coxopleural process, (cst)—forcipular coxosternite, (dms)—dorso-medial spinous process of ultimate prefemur, (p2)— pretarsus of maxilla 2, (pa)—short projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (pla)—pretarsal lamellae, (pt)—process of forcipular trochantero-prefemur, (rl)—right[?] ultimate leg, (S)—? sternite, (st)—saw teeth-like small spinous processes, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)— ultimate tibia, (us)—ultimate LBS, (ust)—ultimate sternite, (vsp)—ventral spinous process of ultimate prefemur." figureDoi="http://doi.org/10.5281/zenodo.4402149" httpUri="https://zenodo.org/record/4402149/files/figure.png" pageId="9" pageNumber="10">Fig. 11</figureCitation>
|
||
), the only figures of the ultimate legs of
|
||
<taxonomicName id="4C77C3B8FFFBD703FB0FF8592849DBCD" authorityName=", Shelley" authorityYear="2002" box="[1163,1292,1976,2002]" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Kethopinae">Kethopinae</taxonomicName>
|
||
are
|
||
<bibRefCitation id="EFE6C5CAFFFBD703FAB8F8592D9FDBE8" author="Crabill, R. E. Jr." pageId="9" pageNumber="10" pagination="235 - 238" refId="ref46739" refString="Crabill, R. E. Jr. (1958) A new Kethops from New Mexico, with a key to its congeners. Proceedings of the Entomological Society of Washington, 60, 235 - 238." type="journal article" year="1958">Crabill’s (1958</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFFBD703FF62F83D2C5ADBE8" author="Crabill, R. E. Jr." box="[230,287,2012,2039]" pageId="9" pageNumber="10" pagination="1 - 15" refId="ref46778" refString="Crabill, R. E. Jr. (1960) A new American genus of cryptopid centipedes, with an annotated key to the scolopendromorph genera of America North of Mexico. Proceedings of the United States National Museum, 111, 1 - 15." type="journal article" year="1960">1960</bibRefCitation>
|
||
) figures of the tibia and tarsus 1 and 2 (
|
||
<figureCitation id="134CA4BEFFFBD703FD59F83D2E75DBE9" box="[733,816,2012,2038]" captionStart="FIGURES 7–13" captionStartId="11.[151,264,1641,1665]" captionTargetBox="[289,1300,194,1595]" captionTargetId="figure-17@11.[276,1312,181,1618]" captionTargetPageId="11" captionText="FIGURES 7–13. Scolopocryptops melanostoma Newport, 1845; Ad. Rc 7172 7 LBS 23 and ultimate legs, laterally; Ad. Rc 7503 8 anterior margin of forcipular coxosternite + maxillae 2, ventrally; Scolopocryptops ferrugineus (L., 1767); Peru specimen from Markus Koch collection 9 forcipular segment ventrally; Scolopocryptops miersii Newport, 1845; BMNH, BM 1889.4.13.3–5 10 apical part of maxilla 2 laterally; Kethops utahensis (Chamberlin, 1909) 11 figures 4–6 of Chamberlin (1912); Thalkethops grallatrix Crabill, 1960 12 figure 15 of Crabill (1960); Kethops utahensis (Chamberlin, 1909); USNM Mill Creek (Utah) specimen 13 apical part of maxilla 2 ventro-laterally; (20)—leg 20, (am)—anterior margin of forcipular coxosternite, (cp)—coxopleural process, (cst)—forcipular coxosternite, (dms)—dorso-medial spinous process of ultimate prefemur, (p2)— pretarsus of maxilla 2, (pa)—short projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (pla)—pretarsal lamellae, (pt)—process of forcipular trochantero-prefemur, (rl)—right[?] ultimate leg, (S)—? sternite, (st)—saw teeth-like small spinous processes, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)— ultimate tibia, (us)—ultimate LBS, (ust)—ultimate sternite, (vsp)—ventral spinous process of ultimate prefemur." figureDoi="http://doi.org/10.5281/zenodo.4402149" httpUri="https://zenodo.org/record/4402149/files/figure.png" pageId="9" pageNumber="10">Fig. 12</figureCitation>
|
||
). In fact, the real structure (and spinulation) of both the ultimate prefemur and femur of
|
||
<taxonomicName id="4C77C3B8FFF8D700FE79FF762F3BDCAE" authorityName=", Shelley" authorityYear="2002" box="[509,638,151,177]" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subFamily" subFamily="Kethopinae">Kethopinae</taxonomicName>
|
||
is ambiguous as there are no drawings of the ultimate legs among scant and schematic figures of
|
||
<bibRefCitation id="EFE6C5CAFFF8D700FE36FF5A2FC3DCCA" author="Shelley, R. M." box="[434,646,187,213]" pageId="10" pageNumber="11" pagination="1 - 108" refId="ref50598" refString="Shelley, R. M. (2002) A synopsis of the North American centipedes of the order Scolopendromorpha (Chilopoda). Virginia Museum of Natural History, 5, 1 - 108." type="journal article" year="2002">Shelley (2002: 78)</bibRefCitation>
|
||
. In 2008 the third author studied one of Chamberlin’s specimens of
|
||
<taxonomicName id="4C77C3B8FFF8D700FA07FF5A2C44DCE6" baseAuthorityName="Chamberlin" baseAuthorityYear="1909" class="Chilopoda" family="Scolopocryptopidae" genus="Kethops" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="utahensis">
|
||
<emphasis id="B9036429FFF8D700FA07FF5A2C44DCE6" italics="true" pageId="10" pageNumber="11">K. utahensis</emphasis>
|
||
</taxonomicName>
|
||
, lacking the ultimate legs, from which novel anatomical details were depicted by SEM (
|
||
<figureCitation id="134CA4BEFFF8D700FB58FF3E286BDCE6" box="[1244,1326,223,249]" captionStart="FIGURES 7–13" captionStartId="11.[151,264,1641,1665]" captionTargetBox="[289,1300,194,1595]" captionTargetId="figure-17@11.[276,1312,181,1618]" captionTargetPageId="11" captionText="FIGURES 7–13. Scolopocryptops melanostoma Newport, 1845; Ad. Rc 7172 7 LBS 23 and ultimate legs, laterally; Ad. Rc 7503 8 anterior margin of forcipular coxosternite + maxillae 2, ventrally; Scolopocryptops ferrugineus (L., 1767); Peru specimen from Markus Koch collection 9 forcipular segment ventrally; Scolopocryptops miersii Newport, 1845; BMNH, BM 1889.4.13.3–5 10 apical part of maxilla 2 laterally; Kethops utahensis (Chamberlin, 1909) 11 figures 4–6 of Chamberlin (1912); Thalkethops grallatrix Crabill, 1960 12 figure 15 of Crabill (1960); Kethops utahensis (Chamberlin, 1909); USNM Mill Creek (Utah) specimen 13 apical part of maxilla 2 ventro-laterally; (20)—leg 20, (am)—anterior margin of forcipular coxosternite, (cp)—coxopleural process, (cst)—forcipular coxosternite, (dms)—dorso-medial spinous process of ultimate prefemur, (p2)— pretarsus of maxilla 2, (pa)—short projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (pla)—pretarsal lamellae, (pt)—process of forcipular trochantero-prefemur, (rl)—right[?] ultimate leg, (S)—? sternite, (st)—saw teeth-like small spinous processes, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)— ultimate tibia, (us)—ultimate LBS, (ust)—ultimate sternite, (vsp)—ventral spinous process of ultimate prefemur." figureDoi="http://doi.org/10.5281/zenodo.4402149" httpUri="https://zenodo.org/record/4402149/files/figure.png" pageId="10" pageNumber="11">Fig. 13</figureCitation>
|
||
, fig. 1AB in
|
||
<bibRefCitation id="EFE6C5CAFFF8D700FF30FEE22C9EDD02" author="Edgecombe, G. D. & Koch, M." box="[180,475,259,285]" pageId="10" pageNumber="11" pagination="295 - 318" refId="ref47458" refString="Edgecombe, G. D. & Koch, M. (2009) The contribution of preoral chamber and foregut morphology to the phylogenetics of Scolopendromorpha (Chilopoda). Soil Organisms, 81, 295 - 318." type="journal article" year="2009">Edgecombe & Koch 2009</bibRefCitation>
|
||
, figs 2B, 6C in
|
||
<bibRefCitation id="EFE6C5CAFFF8D700FD0EFEE22E0CDD02" author="Koch, M. & Edgecombe, G. D. & Shelley, R. M." box="[650,841,259,285]" pageId="10" pageNumber="11" pagination="51 - 81" refId="ref48332" refString="Koch, M., Edgecombe, G. D. & Shelley, R. M. (2010) Anatomy of Ectonocryptoides (Scolopocryptopidae: Ectonocryptopinae) and the phylogeny of blind Scolopendromorpha (Chilopoda). International Journal of Myriapodology, 3, 51 - 81. https: // doi. org / 10.1163 / 187525410 X 12578602960344" type="journal article" year="2010">
|
||
Koch
|
||
<emphasis id="B9036429FFF8D700FD49FEE22FBADD02" box="[717,767,259,285]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFF8D700FF13FE8F2F43DD97" blockId="10.[151,518,366,429]" box="[151,518,366,393]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9036429FFF8D700FF13FE8F2F43DD97" bold="true" box="[151,518,366,393]" pageId="10" pageNumber="11">
|
||
(!)
|
||
<taxonomicName id="4C77C3B8FFF8D700FF38FE8F2F43DD97" authority="Chamberlin, 1912" authorityName="Chamberlin" authorityYear="1912" box="[188,518,366,393]" class="Chilopoda" family="Scolopocryptopidae" genus="Kethops" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFF8D700FF38FE8F2C5BDD96" bold="true" box="[188,286,366,393]" italics="true" pageId="10" pageNumber="11">Kethops</emphasis>
|
||
<bibRefCitation id="EFE6C5CAFFF8D700FEA1FE8F2F43DD97" author="Chamberlin, R. V." box="[293,518,366,392]" pageId="10" pageNumber="11" pagination="141 - 172" refId="ref46640" refString="Chamberlin, R. V. (1912) New North American chilopods and diplopods. Annals of the Entomological Society of America, 5, 141 - 172." type="journal article" year="1912">Chamberlin, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFF8D700FF13FE722C56DDB2" blockId="10.[151,518,366,429]" box="[151,275,403,429]" pageId="10" pageNumber="11">
|
||
<figureCitation id="134CA4BEFFF8D700FF13FE722C56DDB2" box="[151,275,403,429]" captionStart="FIGURES 7–13" captionStartId="11.[151,264,1641,1665]" captionTargetBox="[289,1300,194,1595]" captionTargetId="figure-17@11.[276,1312,181,1618]" captionTargetPageId="11" captionText="FIGURES 7–13. Scolopocryptops melanostoma Newport, 1845; Ad. Rc 7172 7 LBS 23 and ultimate legs, laterally; Ad. Rc 7503 8 anterior margin of forcipular coxosternite + maxillae 2, ventrally; Scolopocryptops ferrugineus (L., 1767); Peru specimen from Markus Koch collection 9 forcipular segment ventrally; Scolopocryptops miersii Newport, 1845; BMNH, BM 1889.4.13.3–5 10 apical part of maxilla 2 laterally; Kethops utahensis (Chamberlin, 1909) 11 figures 4–6 of Chamberlin (1912); Thalkethops grallatrix Crabill, 1960 12 figure 15 of Crabill (1960); Kethops utahensis (Chamberlin, 1909); USNM Mill Creek (Utah) specimen 13 apical part of maxilla 2 ventro-laterally; (20)—leg 20, (am)—anterior margin of forcipular coxosternite, (cp)—coxopleural process, (cst)—forcipular coxosternite, (dms)—dorso-medial spinous process of ultimate prefemur, (p2)— pretarsus of maxilla 2, (pa)—short projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (pla)—pretarsal lamellae, (pt)—process of forcipular trochantero-prefemur, (rl)—right[?] ultimate leg, (S)—? sternite, (st)—saw teeth-like small spinous processes, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)— ultimate tibia, (us)—ultimate LBS, (ust)—ultimate sternite, (vsp)—ventral spinous process of ultimate prefemur." figureDoi="http://doi.org/10.5281/zenodo.4402149" httpUri="https://zenodo.org/record/4402149/files/figure.png" pageId="10" pageNumber="11">Figs 11, 13</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFF8D700FF13FE362E85DDF0" box="[151,960,471,495]" pageId="10" pageNumber="11" type="type_taxon">
|
||
<paragraph id="8BC8B83BFFF8D700FF13FE362E85DDF0" blockId="10.[151,960,471,495]" box="[151,960,471,495]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9036429FFF8D700FF13FE362C66DDF0" bold="true" box="[151,291,471,495]" pageId="10" pageNumber="11">
|
||
<typeStatus id="54CC0699FFF8D700FF13FE362D89DDF0" box="[151,204,471,495]" pageId="10" pageNumber="11">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C77C3B8FFF8D700FEAEFE392FFDDDF0" authority="Chamberlin, 1909" authorityName="Chamberlin" authorityYear="1909" box="[298,696,471,495]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="utahensis">
|
||
<emphasis id="B9036429FFF8D700FEAEFE392CBEDDF0" box="[298,507,471,495]" italics="true" pageId="10" pageNumber="11">Newportia utahensis</emphasis>
|
||
Chamberlin, 1909
|
||
</taxonomicName>
|
||
(by original designation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFF8D700FF13FDFC2C80DE44" pageId="10" pageNumber="11" type="diagnosis">
|
||
<paragraph id="8BC8B83BFFF8D700FF13FDFC28C3DE27" blockId="10.[151,1437,541,747]" box="[151,1414,541,568]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9036429FFF8D700FF13FDFC2C57DE28" bold="true" box="[151,274,541,567]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
|
||
Sternites distinctly margined by lateral longitudinal sutures, with both median and transverse sutures.
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFF8D700FF43FDA02C80DE44" blockId="10.[151,1437,541,747]" box="[199,453,577,603]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9036429FFF8D700FF43FDA02CECDE44" bold="true" box="[199,425,577,603]" pageId="10" pageNumber="11">Number of species.</emphasis>
|
||
2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFF8D700FF43FD842C39DEF4" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="8BC8B83BFFF8D700FF43FD842C39DEF4" blockId="10.[151,1437,541,747]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9036429FFF8D700FF43FD842C7EDE60" bold="true" box="[199,315,613,639]" pageId="10" pageNumber="11">Remarks.</emphasis>
|
||
Treated as a genus in
|
||
<bibRefCitation id="EFE6C5CAFFF8D700FDBFFD842E81DE9F" author="Edgecombe, G. D. & Bonato, L." box="[571,964,613,640]" pageId="10" pageNumber="11" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 405)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFF8D700FC54FD8729FDDE9F" author="Bonato, L. & Chagas-Junior, A. & Edgecombe, G. D. & Lewis, J. G. E. & Minelli, A. & Pereira, L. A. & Shelley, R. M. & Stoev, P. & Zapparoli, M." box="[976,1208,613,640]" pageId="10" pageNumber="11" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. & Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
|
||
Bonato
|
||
<emphasis id="B9036429FFF8D700FBADFD872926DE60" box="[1065,1123,613,639]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFF8D700FB40FD8428DCDE9F" author="Lewis, J. G. E." box="[1220,1433,613,640]" pageId="10" pageNumber="11" pagination="21 - 31" refId="ref49313" refString="Lewis, J. G. E. (2016 b) On the consistency of some taxonomic characters in the Scolopendromorpha and comments on the scolopocryptopid subfamily Kethopinae (Myriapoda: Chilopoda). Acta Societatis Zoologicae Bohemicae, 80, 21 - 31." type="journal article" year="2016">Lewis (2016b: 26)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="EFE6C5CAFFF8D700FF13FD682CE1DEBB" author="Chamberlin, R. V." box="[151,420,649,676]" pageId="10" pageNumber="11" pagination="141 - 172" refId="ref46640" refString="Chamberlin, R. V. (1912) New North American chilopods and diplopods. Annals of the Entomological Society of America, 5, 141 - 172." type="journal article" year="1912">Chamberlin (1912: 156)</bibRefCitation>
|
||
wrote that sternites are with “Usually two[!] or more weaker and more indefinite transverse sulci”; his corresponding fig. 5 also demonstrates not one but two transverse sutures, which is very unusual for scolopendromorphs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |