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<document id="8758F9FF50264CAF2C6F46561E4FC28C" ID-CLB-Dataset="8352" ID-DOI="10.5281/zenodo.4526404" ID-GBIF-Dataset="3cea39ec-86b6-419b-8cdb-5baeedc4cc43" ID-ISSN="0253-116X" ID-Zenodo-Dep="4526404" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612224402507" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03C587F68410FFAAFF123905FDCAFA74" docLanguage="en" docName="LinzerbiolBeitr.49.2.1029-1039.pdf" docOrigin="Linzer biologische Beiträge 49 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Quedius (Raphirus) boops" docType="treatment" docVersion="8" lastPageNumber="1037" masterDocId="FFFCFF8E8417FFA2FFB13A12FFEBFFAA" masterDocTitle="On the micropterous Quedius (Raphirus) species with a punctate scutellum of Turkey (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="1039" masterPageNumber="1029" pageNumber="1036" updateTime="1698926418021" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9351B88F6A5DF72AD04533A79F433A42">On the micropterous Quedius (Raphirus) species with a punctate scutellum of Turkey (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="C09D2F6C948FD1B6F071D1F5E51CEF79">Assing, Volker</mods:namePart>
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<mods:title id="AAF2249484F4F2FA613FEEFB6154C2A3">Linzer biologische Beiträge</mods:title>
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<mods:date id="B2D0237B67A46C79DC0A9AC27587A3C5">2017</mods:date>
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<mods:number id="BC304B94ACDB5D998D1089DEBB65D7BF">2017-12-11</mods:number>
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<treatment id="03C587F68410FFAAFF123905FDCAFA74" ID-DOI="http://doi.org/10.5281/zenodo.4527172" ID-GBIF-Taxon="177313204" ID-Zenodo-Dep="4527172" LSID="urn:lsid:plazi:treatment:03C587F68410FFAAFF123905FDCAFA74" httpUri="http://treatment.plazi.org/id/03C587F68410FFAAFF123905FDCAFA74" lastPageId="8" lastPageNumber="1037" pageId="7" pageNumber="1036">
<subSubSection id="C376656B8410FFA5FF123905FD7DFC87" box="[163,662,791,813]" pageId="7" pageNumber="1036" type="nomenclature">
<paragraph id="8BD336E08410FFA5FF123905FD7DFC87" blockId="7.[163,662,791,813]" box="[163,662,791,813]" pageId="7" pageNumber="1036">
<heading id="D09B818C8410FFA5FF123905FD7DFC87" box="[163,662,791,813]" fontSize="8" level="2" pageId="7" pageNumber="1036" reason="2">
<taxonomicName id="4C6C4D638410FFA5FF123905FD7DFC87" ID-CoL="4R32M" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[163,662,791,813]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops" subGenus="Raphirus">
<emphasis id="B918EAF28410FFA5FF123905FD7DFC87" bold="true" box="[163,662,791,813]" pageId="7" pageNumber="1036">
<emphasis id="B918EAF28410FFA5FF123905FF11FC87" bold="true" box="[163,250,791,813]" italics="true" pageId="7" pageNumber="1036">Quedius</emphasis>
(
<emphasis id="B918EAF28410FFA5FEB83905FE81FC87" bold="true" box="[265,362,791,813]" italics="true" pageId="7" pageNumber="1036">Raphirus</emphasis>
)
<emphasis id="B918EAF28410FFA5FEC83905FE5DFC87" bold="true" box="[377,438,791,813]" italics="true" pageId="7" pageNumber="1036">boops</emphasis>
(
<bibRefCitation id="EFFD4B118410FFA5FE723905FD64FC87" author="GRAVENHORST J" box="[451,655,791,813]" pageId="7" pageNumber="1036" refId="ref5293" refString="GRAVENHORST J. L. C. (1802): Coleoptera microptera Brunsvicensia. - Brunsuigae: 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376656B8410FFA5FF123951FC63FB8B" pageId="7" pageNumber="1036" type="reference_group">
<paragraph id="8BD336E08410FFA5FF123951FDA1FCF3" blockId="7.[163,1071,835,1216]" box="[163,586,835,857]" pageId="7" pageNumber="1036">
<treatmentCitationGroup id="AB7C11CE8410FFA5FF123951FDA1FCF3" box="[163,586,835,857]" pageId="7" pageNumber="1036">
<treatmentCitation id="0ACD10F18410FFA5FF123951FDDCFCF2" author="GRAVENHORST J" box="[163,567,835,856]" page="21" pageId="7" pageNumber="1036" year="1802">
<taxonomicName id="4C6C4D638410FFA5FF123951FDDCFCF2" authority="GRAVENHORST, 1802: 21" authorityName="Gravenhorst, V. Gusarov" authorityPageNumber="21" authorityYear="1802" box="[163,567,835,856]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FF123951FEA2FCF2" box="[163,329,835,856]" italics="true" pageId="7" pageNumber="1036">Staphylinus boops</emphasis>
<bibRefCitation id="EFFD4B118410FFA5FEFF3951FDDCFCF2" author="GRAVENHORST J" box="[334,567,835,856]" pageId="7" pageNumber="1036" refId="ref5293" refString="GRAVENHORST J. L. C. (1802): Coleoptera microptera Brunsvicensia. - Brunsuigae: 206 pp." type="book" year="1802">
GRAVENHORST, 1802:
<specimenCount id="9D6AFD698410FFA5FD913951FDDCFCF2" box="[544,567,835,856]" pageId="7" pageNumber="1036" type="generic">21</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD336E08410FFA5FF12394FFD56FCD8" blockId="7.[163,1071,835,1216]" box="[163,701,861,883]" pageId="7" pageNumber="1036">
<treatmentCitationGroup id="AB7C11CE8410FFA5FF12394FFD56FCD8" box="[163,701,861,883]" pageId="7" pageNumber="1036">
<treatmentCitation id="0ACD10F18410FFA5FF12394FFD8FFCD8" author="NORDMANN A" box="[163,612,861,883]" page="78" pageId="7" pageNumber="1036" year="1837">
<taxonomicName id="4C6C4D638410FFA5FF12394FFD8FFCD8" authority="NORDMANN, 1837" authorityName="NORDMANN" authorityYear="1837" box="[163,612,861,883]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1038" phylum="Arthropoda" rank="subSpecies" species="boops" subSpecies="tauricus">
<emphasis id="B918EAF28410FFA5FF12394FFE7AFCD9" box="[163,401,861,883]" italics="true" pageId="7" pageNumber="1036">Philonthus boops tauricus</emphasis>
<bibRefCitation id="EFFD4B118410FFA5FE26394CFD8FFCD8" author="NORDMANN A" box="[407,612,861,883]" pageId="7" pageNumber="1036" refId="ref5661" refString="NORDMANN A. VON (1837): Symbolae ad monographiam staphylinorum. Ex Academiae Caesareae Scientiarum. - Petropoli: Academiae Caesareae Scientiarum: 167 pp." type="book" year="1837">NORDMANN, 1837: 78</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<emphasis id="B918EAF28410FFA5FDDF394FFD56FCD8" bold="true" box="[622,701,861,882]" pageId="7" pageNumber="1036">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD336E08410FFA5FF123965FDB5FC26" blockId="7.[163,1071,835,1216]" box="[163,606,887,909]" pageId="7" pageNumber="1036">
<treatmentCitationGroup id="AB7C11CE8410FFA5FF123965FDB5FC26" box="[163,606,887,909]" pageId="7" pageNumber="1036">
<treatmentCitation id="0ACD10F18410FFA5FF123965FDEDFC27" author="TOTTENHAM C" box="[163,518,887,909]" page="258" pageId="7" pageNumber="1036" year="1948">
<taxonomicName id="4C6C4D638410FFA5FF123965FDEDFC27" authority="TOTTENHAM, 1948: 258" authorityName="TOTTENHAM" authorityPageNumber="258" authorityYear="1948" box="[163,518,887,909]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="crius">
<emphasis id="B918EAF28410FFA5FF123965FECAFC27" box="[163,289,887,909]" italics="true" pageId="7" pageNumber="1036">Quedius crius</emphasis>
<bibRefCitation id="EFFD4B118410FFA5FE96396AFDEDFC27" author="TOTTENHAM C" box="[295,518,887,909]" pageId="7" pageNumber="1036" pagination="241 - 258" refId="ref6020" refString="TOTTENHAM C. E. (1948): A revision of the British species of Arphirus TOTTENHAM (subgenus of Quedius STEPHENS) (Col., Staphylinidae). - The Entomologist's Monthly Magazine 84: 241 - 258." type="journal article" year="1948">TOTTENHAM, 1948: 258</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<emphasis id="B918EAF28410FFA5FDBE3965FDB5FC26" bold="true" box="[527,606,887,908]" pageId="7" pageNumber="1036">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD336E08410FFA5FF123983FD4FFC0C" blockId="7.[163,1071,835,1216]" box="[163,676,913,935]" pageId="7" pageNumber="1036">
<treatmentCitationGroup id="AB7C11CE8410FFA5FF123983FD4FFC0C" box="[163,676,913,935]" pageId="7" pageNumber="1036">
<treatmentCitation id="0ACD10F18410FFA5FF123983FDDDFC0C" author="FAGEL G" box="[163,566,913,935]" page="113" pageId="7" pageNumber="1036" year="1960">
<taxonomicName id="4C6C4D638410FFA5FF123983FDDDFC0C" ID-CoL="4R3PT" authority="FAGEL, 1960" authorityName="FAGEL" authorityYear="1960" box="[163,566,913,935]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1038" phylum="Arthropoda" rank="subSpecies" species="boops" subSpecies="islandicus">
<emphasis id="B918EAF28410FFA5FF123983FE67FC0C" box="[163,396,913,934]" italics="true" pageId="7" pageNumber="1036">Quedius boops islandicus</emphasis>
<bibRefCitation id="EFFD4B118410FFA5FE233980FDDDFC0C" author="FAGEL G" box="[402,566,913,935]" pageId="7" pageNumber="1036" pagination="108 - 123" refId="ref5221" refString="FAGEL G. (1960): Contribution a la connaissance des Staphylinidae. LXIV. Qu'est le Quedius boops GRAVENHORST? - Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 96: 108 - 123." type="journal article" year="1960">
FAGEL, 1960:
<specimenCount id="9D6AFD698410FFA5FDA53983FDDDFC0C" box="[532,566,913,934]" pageId="7" pageNumber="1036" type="generic">113</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.;
<emphasis id="B918EAF28410FFA5FDE53983FD4FFC0C" bold="true" box="[596,676,913,934]" pageId="7" pageNumber="1036">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD336E08410FFA5FF1239A3FC63FB8B" blockId="7.[163,1071,835,1216]" pageId="7" pageNumber="1036">
<materialsCitation id="3B043CBD8410FFA5FF1239A3FC63FB8B" ID-GBIF-Occurrence="3033390308" collectionCode="T, V, MNB" determinerName="I. Gusarov" location="Mus" pageId="7" pageNumber="1036" specimenCount="3" stateProvince="Mus" typeStatus="lectotype">
<collectionCode id="ED7DAE258410FFA5FF1239A3FF5AFC6C" box="[163,177,945,966]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="1036">T</collectionCode>
y p e m a t e r i a l e x a m i n e d:
<taxonomicName id="4C6C4D638410FFA5FDE339A2FD0AFC6F" ID-CoL="4R32M" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[594,737,944,965]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FDE339A2FD0AFC6F" box="[594,737,944,965]" italics="true" pageId="7" pageNumber="1036">Quedius boops</emphasis>
</taxonomicName>
:
<typeStatus id="54D788428410FFA5FD4539A3FCA4FC6C" box="[756,847,945,966]" pageId="7" pageNumber="1036" type="lectotype">Lectotype</typeStatus>
3 [dissected prior to present study], present designation: &quot;6255 / Braunschweig, Hellwig / Grav Type
<emphasis id="B918EAF28410FFA5FC1239D5FC32FC76" box="[931,985,967,988]" italics="true" pageId="7" pageNumber="1036">boops</emphasis>
! /
<taxonomicName id="4C6C4D638410FFA5FC4939D5FF28FC59" ID-CoL="4R32M" authority="Gr." authorityName="Gr." class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FC4939D5FBC5FC76" box="[1016,1070,967,988]" italics="true" pageId="7" pageNumber="1036">boops</emphasis>
Gr.
</taxonomicName>
* Gyll. / Zool.
<collectingRegion id="49A8F8028410FFA5FEF839CCFE9FFC59" box="[329,372,990,1011]" country="Turkey" name="Mus" pageId="7" pageNumber="1036">Mus</collectingRegion>
.
<collectingRegion id="49A8F8028410FFA5FECC39CCFE5DFC5E" box="[381,438,990,1012]" country="Germany" name="Berlin" pageId="7" pageNumber="1036">Berlin</collectingRegion>
/
<typeStatus id="54D788428410FFA5FE7939CCFDC6FC59" box="[456,557,990,1011]" pageId="7" pageNumber="1036" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="4C6C4D638410FFA5FD8239CCFC2FFC59" authority="Gravenhorst, V. Gusarov" authorityName="Gravenhorst, V. Gusarov" authorityYear="1802" box="[563,964,990,1012]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FD8239CCFD32FC59" box="[563,729,990,1011]" italics="true" pageId="7" pageNumber="1036">Staphylinus boops</emphasis>
Gravenhorst, V. Gusarov
</taxonomicName>
des. 1993 /
<taxonomicName id="4C6C4D638410FFA5FF1239E6FE9DFBA0" ID-CoL="4R32M" authority="(Grav.)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[163,374,1012,1034]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FF1239E6FEC7FBA3" box="[163,300,1012,1033]" italics="true" pageId="7" pageNumber="1036">Quedius boops</emphasis>
(Grav.)
</taxonomicName>
3,
<collectionCode id="ED7DAE258410FFA5FE2D39E7FE5BFBA0" box="[412,432,1013,1034]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="1036" type="Museum">V</collectionCode>
.
<determinerName id="7E0FDA248410FFA5FE0239E7FDF8FBA3" box="[435,531,1012,1034]" pageId="7" pageNumber="1036">I. Gusarov</determinerName>
det. 1993 /
<typeStatus id="54D788428410FFA5FD3539E7FD02FBA0" box="[644,745,1013,1034]" pageId="7" pageNumber="1036" type="lectotype">Lectotypus</typeStatus>
3
<taxonomicName id="4C6C4D638410FFA5FCBB39E6FBC1FBA0" authority="Gravenhorst" authorityName="Gravenhorst, V. Gusarov" authorityYear="1802" box="[778,1066,1012,1034]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FCBB39E6FC5AFBA3" box="[778,945,1012,1033]" italics="true" pageId="7" pageNumber="1036">Staphylinus boops</emphasis>
Gravenhorst
</taxonomicName>
, desig.
<collectionCode id="ED7DAE258410FFA5FF6F3E19FF1AFB8A" box="[222,241,1035,1056]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="1036" type="Museum">V</collectionCode>
.
<bibRefCitation id="EFFD4B118410FFA5FF4B3E19FE80FB8A" author="ASSING V" box="[250,363,1035,1057]" pageId="7" pageNumber="1036" pagination="207 - 228" refId="ref5116" refString="ASSING V. (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 49 (1): 207 - 228." type="journal article" year="2017">Assing 2017</bibRefCitation>
/
<taxonomicName id="4C6C4D638410FFA5FECC3E19FD62FB8B" ID-CoL="4R32M" authority="(Gravenhorst)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[381,649,1035,1057]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FECC3E19FDEEFB8A" box="[381,517,1035,1056]" italics="true" pageId="7" pageNumber="1036">Quedius boops</emphasis>
(Gravenhorst)
</taxonomicName>
3, det.
<collectionCode id="ED7DAE258410FFA5FD653E19FD0CFB8A" box="[724,743,1035,1056]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="1036" type="Museum">V</collectionCode>
. Assing&quot; (
<collectionCode id="ED7DAE258410FFA5FCF43E19FC96FB8B" box="[837,893,1035,1057]" pageId="7" pageNumber="1036">MNB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376656B8410FFAAFF123E30FDCAFA74" lastPageId="8" lastPageNumber="1037" pageId="7" pageNumber="1036" type="description">
<paragraph id="8BD336E08410FFA5FF123E30FC02FBCF" blockId="7.[163,1071,835,1216]" pageId="7" pageNumber="1036">
<taxonomicName id="4C6C4D638410FFA5FF123E30FECDFB9D" authorityName="TOTTENHAM" authorityYear="1948" box="[163,294,1058,1079]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="crius">
<emphasis id="B918EAF28410FFA5FF123E30FECDFB9D" box="[163,294,1058,1079]" italics="true" pageId="7" pageNumber="1036">Quedius crius</emphasis>
</taxonomicName>
:
<materialsCitation id="3B043CBD8410FFA5FE873E30FEDEFBCF" ID-GBIF-Occurrence="3033390310" collectingDate="1637-05-01" collectionCode="TYPE, T, V, BMNH" collectorName="C. E. Tottenham" pageId="7" pageNumber="1036" specimenCount="2" typeStatus="holotype">
<typeStatus id="54D788428410FFA5FE873E30FE61FB92" box="[310,394,1058,1080]" pageId="7" pageNumber="1036" type="holotype">Holotype</typeStatus>
3: &quot;
<typeStatus id="54D788428410FFA5FE0E3E30FDFEFB92" box="[447,533,1058,1080]" pageId="7" pageNumber="1036" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C6C4D638410FFA5FD813E30FD01FB9D" authority="Tott." authorityName="TOTTENHAM" authorityYear="1948" box="[560,746,1058,1079]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="crius">
<emphasis id="B918EAF28410FFA5FD813E30FD59FB9D" box="[560,690,1058,1079]" italics="true" pageId="7" pageNumber="1036">Quedius crius</emphasis>
Tott.
</taxonomicName>
<collectionCode id="ED7DAE258410FFA5FD433E30FCC0FB9D" box="[754,811,1058,1079]" pageId="7" pageNumber="1036">TYPE</collectionCode>
(Askwith,
<date id="FFD210208410FFA5FC123E33FC1DFB9D" box="[931,1014,1057,1079]" pageId="7" pageNumber="1036" value="1637-05-01">
<collectingDate id="EF96E9C88410FFA5FC123E33FC1DFB9D" box="[931,1014,1057,1079]" pageId="7" pageNumber="1036" value="1637-05-01">1-v-1637</collectingDate>
</date>
[sic], C.E.
<collectionCode id="ED7DAE258410FFA5FF7A3E2AFF30FBE7" box="[203,219,1080,1101]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="1036">T</collectionCode>
. /
<collectorName id="269953368410FFA5FF433E2AFE69FBE4" box="[242,386,1080,1102]" pageId="7" pageNumber="1036">C.E. Tottenham</collectorName>
collection. B.M. 1974-587. /
<taxonomicName id="4C6C4D638410FFA5FD233E2AFC49FBE4" authority="(Gravenhorst)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[658,930,1080,1102]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FD233E2AFCF1FBE7" box="[658,794,1080,1101]" italics="true" pageId="7" pageNumber="1036">Quedius boops</emphasis>
(Gravenhorst)
</taxonomicName>
, det.
<collectionCode id="ED7DAE258410FFA5FC623E2BFC0DFBE4" box="[979,998,1081,1102]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="1036" type="Museum">V</collectionCode>
.
<bibRefCitation id="EFFD4B118410FFA5FC5E3E2AFF39FBCE" author="ASSING V" pageId="7" pageNumber="1036" pagination="207 - 228" refId="ref5116" refString="ASSING V. (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 49 (1): 207 - 228." type="journal article" year="2017">Assing 2017</bibRefCitation>
&quot; (
<collectionCode id="ED7DAE258410FFA5FF573E5DFEC4FBCF" box="[230,303,1103,1125]" pageId="7" pageNumber="1036">BMNH</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54D788428410FFA5FE8C3E5DFE72FBCE" box="[317,409,1103,1124]" pageId="7" pageNumber="1036" type="paratype">Paratypes</typeStatus>
: 233: same data as
<typeStatus id="54D788428410FFA5FDFE3E5DFD4AFBCF" box="[591,673,1103,1125]" pageId="7" pageNumber="1036" type="holotype">holotype</typeStatus>
, but labelled as
<typeStatus id="54D788428410FFA5FC823E5DFC61FBCE" box="[819,906,1103,1124]" pageId="7" pageNumber="1036" type="paratype">paratypes</typeStatus>
(BMNH).
</paragraph>
<paragraph id="8BD336E08410FFA5FF123E74FDD5FB6A" blockId="7.[163,1071,835,1216]" pageId="7" pageNumber="1036">
<emphasis id="B918EAF28410FFA5FF123E74FE7CFBD1" box="[163,407,1126,1147]" italics="true" pageId="7" pageNumber="1036">
<taxonomicName id="4C6C4D638410FFA5FF123E74FEDBFBD1" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[163,304,1126,1147]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">Quedius boops</taxonomicName>
islandicus
</emphasis>
:
<materialsCitation id="3B043CBD8410FFA5FE193E74FCB9FB03" ID-GBIF-Occurrence="3033390305" collectionCode="V, MNB" collectorName="Dr. Gruner S. G." country="Iceland" determinerName="G. Fagel" location="Island" pageId="7" pageNumber="1036" specimenCount="2" typeStatus="holotype">
<typeStatus id="54D788428410FFA5FE193E74FE17FBD6" box="[424,508,1126,1148]" pageId="7" pageNumber="1036" type="holotype">Holotype</typeStatus>
3 [dissected prior to present study]: &quot;Island, Reynivellir 22.8.07.
<collectorName id="269953368410FFA5FF453E6FFE62FB3B" box="[244,393,1148,1170]" pageId="7" pageNumber="1036">Dr. Gruner S.G.</collectorName>
/
<typeStatus id="54D788428410FFA5FE2C3E6EFE10FB38" box="[413,507,1148,1170]" pageId="7" pageNumber="1036" type="holotype">Holotypus</typeStatus>
/
<determinerName id="7E0FDA248410FFA5FDA13E6EFDB4FB38" box="[528,607,1148,1170]" pageId="7" pageNumber="1036">G. Fagel</determinerName>
det., 1959,
<emphasis id="B918EAF28410FFA5FD7F3E6EFC51FB3B" box="[718,954,1148,1169]" italics="true" pageId="7" pageNumber="1036">
<taxonomicName id="4C6C4D638410FFA5FD7F3E6EFCBCFB3B" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[718,855,1148,1169]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">Quedius boops</taxonomicName>
islandicus
</emphasis>
nov. / Zool. Mus. Berlin /
<taxonomicName id="4C6C4D638410FFA5FE973E81FDDCFB03" authority="(Gravenhorst)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[294,567,1171,1193]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FE973E81FE44FB02" box="[294,431,1171,1192]" italics="true" pageId="7" pageNumber="1036">Quedius boops</emphasis>
(Gravenhorst)
</taxonomicName>
, det.
<collectionCode id="ED7DAE258410FFA5FDDA3E81FD95FB02" box="[619,638,1171,1192]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="1036" type="Museum">V</collectionCode>
.
<bibRefCitation id="EFFD4B118410FFA5FD393E81FD16FB02" author="ASSING V" box="[648,765,1171,1193]" pageId="7" pageNumber="1036" pagination="207 - 228" refId="ref5116" refString="ASSING V. (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 49 (1): 207 - 228." type="journal article" year="2017">Assing 2017</bibRefCitation>
&quot; (
<collectionCode id="ED7DAE258410FFA5FCA23E81FCA0FB03" box="[787,843,1171,1193]" pageId="7" pageNumber="1036">MNB</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54D788428410FFA5FCEA3E81FC41FB02" box="[859,938,1171,1192]" pageId="7" pageNumber="1036" type="paratype">Paratype</typeStatus>
<specimenCount id="9D6AFD698410FFA5FC033E8AFC55FB07" box="[946,958,1176,1197]" pageId="7" pageNumber="1036" type="female">♀</specimenCount>
: same data as
<typeStatus id="54D788428410FFA5FF0C3EB8FEE4FB6A" box="[189,271,1194,1216]" pageId="7" pageNumber="1036" type="holotype">holotype</typeStatus>
, but labelled as
<typeStatus id="54D788428410FFA5FE103EB8FE04FB15" box="[417,495,1194,1215]" pageId="7" pageNumber="1036" type="paratype">paratype</typeStatus>
(MNB).
</paragraph>
<paragraph id="8BD336E08410FFA5FF123EDCFDB8FAF3" blockId="7.[163,1071,1230,1369]" pageId="7" pageNumber="1036">
C o m m e n t: The original description of
<taxonomicName id="4C6C4D638410FFA5FDDD3EDCFD2CFB49" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[620,711,1230,1251]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FDDD3EDCFD2CFB49" box="[620,711,1230,1251]" italics="true" pageId="7" pageNumber="1036">Q. boops</emphasis>
</taxonomicName>
is based on an unspecified number of
<typeStatus id="54D788428410FFA5FF0E3EF9FEF3FAAA" box="[191,280,1259,1280]" pageId="7" pageNumber="1036" type="syntype">syntypes</typeStatus>
without specified locality from &quot;
<collectingRegion id="49A8F8028410FFA5FDDD3EF9FD76FAAA" box="[620,669,1259,1280]" country="Turkey" name="Mus" pageId="7" pageNumber="1036">Mus</collectingRegion>
. Prof. Hellwig&quot; (
<bibRefCitation id="EFFD4B118410FFA5FCFA3EF9FBC8FAAB" author="GRAVENHORST J" box="[843,1059,1259,1281]" pageId="7" pageNumber="1036" refId="ref5293" refString="GRAVENHORST J. L. C. (1802): Coleoptera microptera Brunsvicensia. - Brunsuigae: 206 pp." type="book" year="1802">GRAVENHORST 1802</bibRefCitation>
). The sole
<typeStatus id="54D788428410FFA5FEB63F1BFEBDFAB4" box="[263,342,1289,1310]" pageId="7" pageNumber="1036" type="syntype">syntype</typeStatus>
in the historical collection of the MNB was studied and labelled as
<typeStatus id="54D788428410FFA5FF123F34FEEBFA96" box="[163,256,1318,1340]" pageId="7" pageNumber="1036" type="lectotype">lectotype</typeStatus>
by Vladimir Gusarov, but this designation was never published. The specimen, a male, is here designated as the
<typeStatus id="54D788428410FFA5FE5E3F51FDA4FAF3" box="[495,591,1347,1369]" pageId="7" pageNumber="1036" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BD336E08410FFAAFF123F7AFCC8FDD7" blockId="7.[163,1071,1383,1582]" lastBlockId="8.[163,1071,204,637]" lastPageId="8" lastPageNumber="1037" pageId="7" pageNumber="1036">
According to SCHÜLKE &amp; SMETANA (2015), the distribution of
<emphasis id="B918EAF28410FFA5FC9B3F7BFC0AFAD4" box="[810,993,1384,1406]" italics="true" pageId="7" pageNumber="1036">
<taxonomicName id="4C6C4D638410FFA5FC9B3F7BFC6DFAD7" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[810,902,1384,1406]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">Q. boops</taxonomicName>
tauricus
</emphasis>
is confined to
<collectingCountry id="F37B76708410FFA5FF4A3F94FEA3FA36" box="[251,328,1414,1436]" name="Turkey" pageId="7" pageNumber="1036">Turkey</collectingCountry>
. The original description of
<taxonomicName id="4C6C4D638410FFA5FDDE3F94FCC9FA31" box="[623,802,1414,1435]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF28410FFA5FDDE3F94FCC9FA31" box="[623,802,1414,1435]" italics="true" pageId="7" pageNumber="1036">Philonthus boops</emphasis>
</taxonomicName>
&quot;
<taxonomicName id="4C6C4D638410FFA5FC823F94FC47FA31" box="[819,940,1414,1435]" pageId="7" pageNumber="1036" rank="variety" variety="taurica">
var.
<emphasis id="B918EAF28410FFA5FCD23F94FC47FA31" box="[867,940,1414,1435]" italics="true" pageId="7" pageNumber="1036">taurica</emphasis>
</taxonomicName>
&quot; [sic], however, is based on an unspecified number of specimens collected &quot;in Tauro-Chersoneso. (Coll. D. Stevenii)&quot; (
<bibRefCitation id="EFFD4B118410FFA5FE373FD2FDA9FA7C" author="NORDMANN A" box="[390,578,1472,1494]" pageId="7" pageNumber="1036" refId="ref5661" refString="NORDMANN A. VON (1837): Symbolae ad monographiam staphylinorum. Ex Academiae Caesareae Scientiarum. - Petropoli: Academiae Caesareae Scientiarum: 167 pp." type="book" year="1837">NORDMANN 1837</bibRefCitation>
). Taurus Chersonesus is an old name for the Crimean Peninsula (
<collectingCountry id="F37B76708410FFA5FEC33FCCFE2CFA5E" box="[370,455,1502,1524]" name="Ukraine" pageId="7" pageNumber="1036">Ukraine</collectingCountry>
) and not situated in the Taurus range,
<collectingCountry id="F37B76708410FFA5FCE33FCCFC74FA5E" box="[850,927,1502,1524]" name="Turkey" pageId="7" pageNumber="1036">Turkey</collectingCountry>
. According to
<bibRefCitation id="EFFD4B118410FFA5FF123FE9FEB0F9BB" author="HERMAN L" box="[163,347,1531,1553]" pageId="7" pageNumber="1036" refId="ref5393" refString="HERMAN L. H. (2001 b): Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Volumes I-VII. - Bulletin of the American Museum of Natural History 265: 4218 pp." type="book" year="2001">HERMAN (2001b)</bibRefCitation>
, there is only one subsequent record of
<emphasis id="B918EAF28410FFA5FCA43FE9FC3FF9BB" box="[789,980,1531,1553]" italics="true" pageId="7" pageNumber="1036">
<taxonomicName id="4C6C4D638410FFA5FCA43FE9FC9EF9BA" box="[789,885,1531,1552]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1036" phylum="Arthropoda" rank="species" species="boops">Q. boops</taxonomicName>
tauricus
</emphasis>
.
<bibRefCitation id="EFFD4B118410FFA5FC543FE9FF0DF984" author="KORGE H" pageId="7" pageNumber="1036" pagination="105 - 126" refId="ref5487" refString="KORGE H. (1964): Carabiden- und Staphylinidenfunde in den Pontischen Gebirgen Kleinasiens und in Mazedonien (Coleoptera). - Reichenbachia 4 (14): 105 - 126." type="journal article" year="1964">KORGE (1964)</bibRefCitation>
tentatively recorded it from Soğanlı Geçidi,
<collectingRegion id="49A8F8028410FFA5FD033C0AFCEDF984" box="[690,774,1560,1582]" country="Turkey" name="Bayburt" pageId="7" pageNumber="1036">Bayburt</collectingRegion>
, based on a single specimen. In the beginning of the passage on
<emphasis id="B918EAF2841FFFAAFDAB3ADEFD3FFF4B" box="[538,724,204,225]" italics="true" pageId="8" pageNumber="1037">
<taxonomicName id="4C6C4D63841FFFAAFDAB3ADEFD93FF4B" box="[538,632,204,225]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">Q. boops</taxonomicName>
tauricus
</emphasis>
(pp.
<specimenCount id="9D6AFD69841FFFAAFCBE3ADEFCA5FF48" box="[783,846,204,226]" pageId="8" pageNumber="1037" type="generic">124 f.</specimenCount>
) he states that it is a male, but it can be inferred from the last sentence (&quot;Zur endgültigen taxonomischen Wertung dieser Form muss der Fund eines 3 abgewartet werden.&quot;; p. 125) that the specimen is in fact a female. According to
<bibRefCitation id="EFFD4B11841FFFAAFDE93B36FCF1FE90" author="HORN W &amp; KAHLE I &amp; FRIESE G" box="[600,794,292,314]" pageId="8" pageNumber="1037" refId="ref5439" refString="HORN W., KAHLE I., FRIESE G. &amp; R. GAEDIKE (1990): Collectiones entomologicae. Ein Kompendium uber den Verbleib entomologischer Sammlungen der Welt bis 1960. - Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin: 573 pp." type="book" year="1990">HORN et al. (1990)</bibRefCitation>
, most of the Steven collection is deposited in the Zoological Museum of Moscow State University. A request for the loan of the
<typeStatus id="54D78842841FFFAAFE8A3B4DFE8CFEDE" box="[315,359,351,372]" pageId="8" pageNumber="1037">type</typeStatus>
material remained unanswered. There are three specimens, two males and one female, of
<taxonomicName id="4C6C4D63841FFFAAFEDA3B6EFE2CFE3B" box="[363,455,380,401]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFEDA3B6EFE2CFE3B" box="[363,455,380,401]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
from the Crimean Peninsula (&quot;N-slope of Ai Petri Mt.,
<quantity id="4C949B05841FFFAAFBB93B6EFF52FE04" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="1037" unit="m" value="900.0">900 m</quantity>
, beech forest,
<date id="FFD21020841FFFAAFEE13B8BFE1BFE04" box="[336,496,409,430]" pageId="8" pageNumber="1037" value="2001-07-20" valueMax="2001-07-24" valueMin="2001-07-20">20-24.VII.2001</date>
, leg. Koval&quot;) in my collection. They are identical to
<taxonomicName id="4C6C4D63841FFFAAFBA43B8BFF34FE61" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFBA43B8BFF34FE61" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
in every respect, both in external and the male sexual characters. It can be concluded, therefore, that the
<typeStatus id="54D78842841FFFAAFE003BC6FE36FE43" box="[433,477,468,489]" pageId="8" pageNumber="1037">type</typeStatus>
material of
<emphasis id="B918EAF2841FFFAAFDEB3BC6FCFAFE43" box="[602,785,468,489]" italics="true" pageId="8" pageNumber="1037">
<taxonomicName id="4C6C4D63841FFFAAFDEB3BC6FD5DFE43" box="[602,694,468,489]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">Q. boops</taxonomicName>
tauricus
</emphasis>
either refers to
<taxonomicName id="4C6C4D63841FFFAAFC043BC6FBFAFE43" box="[949,1041,468,489]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFC043BC6FBFAFE43" box="[949,1041,468,489]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
or to a different species. In order to dispose of a doubtful name, the former option is preferred and
<emphasis id="B918EAF2841FFFAAFEA3381DFE23FD8E" box="[274,456,526,548]" italics="true" pageId="8" pageNumber="1037">
<taxonomicName id="4C6C4D63841FFFAAFEA3381DFE85FD89" box="[274,366,526,548]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">Q. boops</taxonomicName>
tauricus
</emphasis>
is synonymized with
<taxonomicName id="4C6C4D63841FFFAAFD18381DFCEFFD89" box="[681,772,526,548]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFD18381DFCEFFD89" box="[681,772,526,548]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
. There is little doubt that the record by
<bibRefCitation id="EFFD4B11841FFFAAFEBF383EFE49FDE8" author="KORGE H" box="[270,418,556,578]" pageId="8" pageNumber="1037" pagination="105 - 126" refId="ref5487" refString="KORGE H. (1964): Carabiden- und Staphylinidenfunde in den Pontischen Gebirgen Kleinasiens und in Mazedonien (Coleoptera). - Reichenbachia 4 (14): 105 - 126." type="journal article" year="1964">KORGE (1964)</bibRefCitation>
refers to
<taxonomicName id="4C6C4D63841FFFAAFDB9383EFD8DFDEB" box="[520,614,556,577]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFDB9383EFD8DFDEB" box="[520,614,556,577]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
, a highly variable species. I have examined material of
<taxonomicName id="4C6C4D63841FFFAAFEAC385BFE91FDF4" box="[285,378,585,606]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFEAC385BFE91FDF4" box="[285,378,585,606]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
from various provinces in North
<collectingCountry id="F37B7670841FFFAAFD6C385BFCDFFDF5" box="[733,820,585,607]" name="Turkey" pageId="8" pageNumber="1037">Anatolia</collectingCountry>
(
<collectingRegion id="49A8F802841FFFAAFCF5385BFC63FDF4" box="[836,904,585,606]" country="Turkey" name="Duzce" pageId="8" pageNumber="1037">Düzce</collectingRegion>
,
<collectingRegion id="49A8F802841FFFAAFC27385BFC01FDF5" box="[918,1002,585,607]" country="Turkey" name="Sakarya" pageId="8" pageNumber="1037">Sakarya</collectingRegion>
,
<collectingRegion id="49A8F802841FFFAAFC46385BFBC0FDF5" box="[1015,1067,585,607]" country="Turkey" name="Bolu" pageId="8" pageNumber="1037">Bolu</collectingRegion>
,
<collectingRegion id="49A8F802841FFFAAFF123875FEF2FDD6" box="[163,281,615,636]" country="Turkey" name="Kastamonu" pageId="8" pageNumber="1037">Kastamonu</collectingRegion>
,
<collectingRegion id="49A8F802841FFFAAFE923874FE9DFDD1" box="[291,374,614,635]" country="Turkey" name="Samsun" pageId="8" pageNumber="1037">Samsun</collectingRegion>
,
<collectingRegion id="49A8F802841FFFAAFE313874FE5CFDD6" box="[384,439,614,636]" country="Turkey" name="Ordu" pageId="8" pageNumber="1037">Ordu</collectingRegion>
,
<collectingRegion id="49A8F802841FFFAAFE703874FDD7FDD6" box="[449,572,614,636]" country="Turkey" name="Gumushane" pageId="8" pageNumber="1037">Gümüşhane</collectingRegion>
,
<collectingRegion id="49A8F802841FFFAAFDF73874FD75FDD6" box="[582,670,614,636]" country="Turkey" name="Trabzon" pageId="8" pageNumber="1037">Trabzon</collectingRegion>
; all in cAss).
</paragraph>
<paragraph id="8BD336E0841FFFAAFF12389EFE4CFB3E" blockId="8.[163,1070,652,1172]" pageId="8" pageNumber="1037">
The original description of
<taxonomicName id="4C6C4D63841FFFAAFE74389EFDBDFD0B" authorityName="TOTTENHAM" authorityYear="1948" box="[453,598,652,673]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="crius">
<emphasis id="B918EAF2841FFFAAFE74389EFDBDFD0B" box="[453,598,652,673]" italics="true" pageId="8" pageNumber="1037">Quedius crius</emphasis>
</taxonomicName>
is based on a male
<typeStatus id="54D78842841FFFAAFC81389EFC63FD08" box="[816,904,652,674]" pageId="8" pageNumber="1037" type="holotype">holotype</typeStatus>
and three male
<typeStatus id="54D78842841FFFAAFF1238BBFEEFFD14" box="[163,260,681,702]" pageId="8" pageNumber="1037" type="paratype">paratypes</typeStatus>
from &quot;Askwith (MY)&quot;, and an unspecified number of &quot;specimens in the Sharp collection at the British Museum from Rannoch (PM), Arran (B), Dumfries and Morton Mains (DF), Polmont (SG), Garlochhead (DN), and Inverness-shire&quot; (
<bibRefCitation id="EFFD4B11841FFFAAFC1C38F6FF32FCBD" author="TOTTENHAM C" pageId="8" pageNumber="1037" pagination="241 - 258" refId="ref6020" refString="TOTTENHAM C. E. (1948): A revision of the British species of Arphirus TOTTENHAM (subgenus of Quedius STEPHENS) (Col., Staphylinidae). - The Entomologist's Monthly Magazine 84: 241 - 258." type="journal article" year="1948">TOTTENHAM 1948</bibRefCitation>
). The name was subsequently regarded as a synonym (
<bibRefCitation id="EFFD4B11841FFFAAFC9F3913FC3AFCBD" author="SMETANA A" box="[814,977,769,791]" pageId="8" pageNumber="1037" pagination="133 - 155" refId="ref5810" refString="SMETANA A. (1962): Bestimmungstabelle der mitteleuroaischen Arten der Gattung Quedius STEPH. (Col., Staphylinidae). - Entomologische Blatter 58 (3): 133 - 155." type="journal article" year="1962">SMETANA 1962</bibRefCitation>
) or as a subspecies of
<emphasis id="B918EAF2841FFFAAFE8C390DFE54FC9E" box="[317,447,799,820]" italics="true" pageId="8" pageNumber="1037">Q. asturicus</emphasis>
BERNHAUER, 1918 (
<bibRefCitation id="EFFD4B11841FFFAAFD13390CFCC2FC9E" author="LOHSE G" box="[674,809,798,820]" pageId="8" pageNumber="1037" pagination="1 - 264" refId="ref5545" refString="LOHSE G. A. (1964): Staphylinidae I. (Micropeplinae bis Tachyporinae). - In: FREUDE H., HARDE K. &amp; G. A. LOHSE (eds), Die Kafer Mitteuropas 4. - Krefeld: Goecke &amp; Evers: 1 - 264." type="book chapter" year="1964">LOHSE 1964</bibRefCitation>
,
<bibRefCitation id="EFFD4B11841FFFAAFC88390CFC33FC99" author="COIFFAIT H" box="[825,984,798,819]" pageId="8" pageNumber="1037" refId="ref5172" refString="COIFFAIT H. (1978): Coleopteres staphylinides de la region paleartique [sic] occidentale. III. Sous famille Staphylininae, Tribu Quediini. Sous famille Paederinae, Tribu Pinophilini. - Supplement a la Nouvelle Revue d'Entomologie 8 (4): 364 pp." type="book" year="1978">COIFFAIT 1978</bibRefCitation>
,
<bibRefCitation id="EFFD4B11841FFFAAFC59390DFF32FCF8" author="LUCHT W" pageId="8" pageNumber="1037" pagination="1 - 342" refId="ref5635" refString="LUCHT W. H. (1987): Die Kafer Mitteuropas. Katalog. - Krefeld: Goecke &amp; Evers: 1 - 342." type="book chapter" year="1987">LUCHT 1987</bibRefCitation>
), and finally as a synonym of
<taxonomicName id="4C6C4D63841FFFAAFDA8392EFD6BFCFB" authorityName="GRIDELLI" authorityYear="1925" box="[537,640,828,849]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="B918EAF2841FFFAAFDA8392EFD6BFCFB" box="[537,640,828,849]" italics="true" pageId="8" pageNumber="1037">Q. reitteri</emphasis>
</taxonomicName>
(LOHSE &amp; LUCHT 1989,
<bibRefCitation id="EFFD4B11841FFFAAFC34392EFBC1FCF8" author="HERMAN L" box="[901,1066,828,850]" pageId="8" pageNumber="1037" refId="ref5393" refString="HERMAN L. H. (2001 b): Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Volumes I-VII. - Bulletin of the American Museum of Natural History 265: 4218 pp." type="book" year="2001">HERMAN 2001b</bibRefCitation>
,
<bibRefCitation id="EFFD4B11841FFFAAFF12394BFEAFFCC4" author="SMETANA A" box="[163,324,857,878]" pageId="8" pageNumber="1037" pagination="653 - 671" refId="ref5930" refString="SMETANA A. (2004): Staphylinidae, subtribe Quediina KRAATZ, 1857. - In: LOBL I. &amp; A. SMETANA (eds), Catalogue of Palaearctic Coleoptera. II. Hydrophiloidea-Histeroidea- Staphylinoidea. - Apollo Books, Stenstrup, pp. 653 - 671." type="book chapter" year="2004">SMETANA 2004</bibRefCitation>
, SCHÜLKE &amp; SMETANA 2015), whereas
<bibRefCitation id="EFFD4B11841FFFAAFD6E394BFCB5FCC5" author="POPE R" box="[735,862,857,879]" pageId="8" pageNumber="1037" pagination="1 - 105" refId="ref5689" refString="POPE R. D. (1977): Coleoptera; pp. 1 - 90. - In: KLOET G. S. &amp; W. D. HINCKS (eds), A checklist of British Insects. Second edition. Part 3. - London: Royal Entomological Society: I- XIV + 1 - 105." type="book chapter" year="1977">POPE (1977)</bibRefCitation>
suspected it to be a synonym of
<taxonomicName id="4C6C4D63841FFFAAFE953965FE6AFC21" box="[292,385,886,908]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFE953965FE6AFC21" box="[292,385,886,908]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
. An examination of the
<typeStatus id="54D78842841FFFAAFDCE3964FD33FC26" box="[639,728,886,908]" pageId="8" pageNumber="1037" type="holotype">holotype</typeStatus>
and two
<typeStatus id="54D78842841FFFAAFC8B3965FC70FC26" box="[826,923,887,908]" pageId="8" pageNumber="1037" type="paratype">paratypes</typeStatus>
from the type locality revealed that the aedeagus is identical to that of
<taxonomicName id="4C6C4D63841FFFAAFD473986FCBFFC03" box="[758,852,916,937]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFD473986FCBFFC03" box="[758,852,916,937]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
, a species subject to intraspecific variation of body colour (ranging from reddish-brown to black) and also in the length of the elytra and of the hind wings (ranging from submacropterous to macropterous). According to an unpublished study of thousands of adult specimens of
<taxonomicName id="4C6C4D63841FFFAAFBA439FEFF34FBB4" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFBA439FEFF34FBB4" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
in North
<collectingCountry id="F37B7670841FFFAAFEFA3E1BFE47FBB4" box="[331,428,1033,1054]" name="Germany" pageId="8" pageNumber="1037">Germany</collectingCountry>
, only a small proportion of the material had fully developed wings and flight muscles. The relative elytral length (EL/PL) of the examined type material of
<taxonomicName id="4C6C4D63841FFFAAFF5B3E56FED6FBF3" authorityName="TOTTENHAM" authorityYear="1948" box="[234,317,1092,1113]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="crius">
<emphasis id="B918EAF2841FFFAAFF5B3E56FED6FBF3" box="[234,317,1092,1113]" italics="true" pageId="8" pageNumber="1037">Q. crius</emphasis>
</taxonomicName>
is 0.59-0.62 and thus well within the range of
<taxonomicName id="4C6C4D63841FFFAAFC943E56FC69FBF3" box="[805,898,1092,1113]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFC943E56FC69FBF3" box="[805,898,1092,1113]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
, whereas that of
<taxonomicName id="4C6C4D63841FFFAAFF123E73FEE2FBDD" authorityName="GRIDELLI" authorityYear="1925" box="[163,265,1121,1143]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="B918EAF2841FFFAAFF123E73FEE2FBDD" box="[163,265,1121,1143]" italics="true" pageId="8" pageNumber="1037">Q. reitteri</emphasis>
</taxonomicName>
ranges from approximately 0.45 to 0.52. In consequence,
<taxonomicName id="4C6C4D63841FFFAAFCEE3E73FC59FBDD" authorityName="TOTTENHAM" authorityYear="1948" box="[863,946,1121,1143]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="crius">
<emphasis id="B918EAF2841FFFAAFCEE3E73FC59FBDD" box="[863,946,1121,1143]" italics="true" pageId="8" pageNumber="1037">Q. crius</emphasis>
</taxonomicName>
is placed in synonymy with
<taxonomicName id="4C6C4D63841FFFAAFEF73E6DFE4AFB39" box="[326,417,1150,1172]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFEF73E6DFE4AFB39" box="[326,417,1150,1172]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD336E0841FFFAAFF123EB6FDCAFA74" blockId="8.[163,1071,1188,1503]" pageId="8" pageNumber="1037">
In the original description of
<emphasis id="B918EAF2841FFFAAFE7C3EB6FD7CFB13" box="[461,663,1188,1209]" italics="true" pageId="8" pageNumber="1037">
<taxonomicName id="4C6C4D63841FFFAAFE7C3EB6FDC3FB13" box="[461,552,1188,1209]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">Q. boops</taxonomicName>
islandicus
</emphasis>
, which is based on a male
<typeStatus id="54D78842841FFFAAFC1B3EB6FBE8FB10" box="[938,1027,1188,1210]" pageId="8" pageNumber="1037" type="holotype">holotype</typeStatus>
and three
<typeStatus id="54D78842841FFFAAFF503ED3FEA9FB7C" box="[225,322,1217,1238]" pageId="8" pageNumber="1037" type="paratype">paratypes</typeStatus>
from &quot;
<collectingCountry id="F37B7670841FFFAAFE223ED3FE3DFB7D" box="[403,470,1217,1239]" name="Iceland" pageId="8" pageNumber="1037">Island</collectingCountry>
: Reynivellir&quot;,
<bibRefCitation id="EFFD4B11841FFFAAFDC53ED3FCEDFB7D" author="FAGEL G" box="[628,774,1217,1239]" pageId="8" pageNumber="1037" pagination="108 - 123" refId="ref5221" refString="FAGEL G. (1960): Contribution a la connaissance des Staphylinidae. LXIV. Qu'est le Quedius boops GRAVENHORST? - Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 96: 108 - 123." type="journal article" year="1960">FAGEL (1960)</bibRefCitation>
gives some minor external characters (coloration, microsculpture, head width, shapes of antennomeres, coarser punctation of pronotum, relatively short elytra, elytral punctation coarser and less dense, abdominal punctation) distinguishing this taxon from the nominal subspecies and states that the aedeagus is identical. As mentioned before,
<taxonomicName id="4C6C4D63841FFFAAFD043F25FCFBFAE1" box="[693,784,1334,1356]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFD043F25FCFBFAE1" box="[693,784,1334,1356]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
is highly variable regarding external characters (coloration, length of elytra, punctation, etc.). Moreover, the presence of a distinct subspecies in
<collectingCountry id="F37B7670841FFFAAFE093F63FDE8FA2D" box="[440,515,1393,1415]" name="Iceland" pageId="8" pageNumber="1037">Iceland</collectingCountry>
would be implausible from a zoogeographic point of view. An examination of the
<typeStatus id="54D78842841FFFAAFE6D3F9CFDDEFA0E" box="[476,565,1422,1444]" pageId="8" pageNumber="1037" type="holotype">holotype</typeStatus>
and one female
<typeStatus id="54D78842841FFFAAFD583F9DFCABFA0E" box="[745,832,1423,1444]" pageId="8" pageNumber="1037" type="paratype">paratype</typeStatus>
revealed that they are well within the range of intraspecific variation of
<taxonomicName id="4C6C4D63841FFFAAFD193FBEFCEFFA6B" box="[680,772,1452,1473]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFD193FBEFCEFFA6B" box="[680,772,1452,1473]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
. Hence,
<emphasis id="B918EAF2841FFFAAFCD33FBEFBC5FA6B" box="[866,1070,1452,1473]" italics="true" pageId="8" pageNumber="1037">
<taxonomicName id="4C6C4D63841FFFAAFCD33FBEFC2BFA6B" box="[866,960,1452,1473]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">Q. boops</taxonomicName>
islandicus
</emphasis>
is placed in synonymy with
<taxonomicName id="4C6C4D63841FFFAAFE713FDBFDF0FA74" box="[448,539,1481,1502]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1037" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="B918EAF2841FFFAAFE713FDBFDF0FA74" box="[448,539,1481,1502]" italics="true" pageId="8" pageNumber="1037">Q. boops</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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