153 lines
21 KiB
XML
153 lines
21 KiB
XML
<document id="19DD2FB6EB80191BDA126E88679B6057" ID-DOI="10.11646/zootaxa.1799.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5124771" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627004570370" checkinUser="felipe" docAuthor="Cline, Andrew R." docDate="2008" docId="03C587F3FFF3FFB79BA19EF47CE0FA8A" docLanguage="en" docName="zt01799p120.pdf" docOrigin="Zootaxa 1799 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1799.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Pocadius nobilis Reitter 1873" docType="treatment" docVersion="3" lastPageNumber="94" masterDocId="FFFCFF8BFFAFFFEA9B369E637E19FFB0" masterDocTitle="Revision of the sap beetle genus Pocadius Erichson, 1843 (Coleoptera: Nitidulidae: Nitidulinae" masterLastPageNumber="120" masterPageNumber="1" pageNumber="93" updateTime="1699014459094" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="02B9D9BBB07E34896EE99EC1234C050A">Revision of the sap beetle genus Pocadius Erichson, 1843 (Coleoptera: Nitidulidae: Nitidulinae</mods:title>
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<mods:namePart id="2E2F15375D3F336CA501BFCF3CA0B46D">Cline, Andrew R.</mods:namePart>
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<mods:title id="B57F41B62F619EB098E6839DC0050066">Zootaxa</mods:title>
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<mods:part id="6DB79159A72FF52B58D25194ADA1F542">
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<mods:date id="925C6DE62358C70F6ED9B2091146812E">2008</mods:date>
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<mods:number id="9828B7A865D32DE89161C48DF95194B7">2008-06-16</mods:number>
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<mods:detail id="AF2849DC170BC7903E3EC12FA9A92B59" type="volume">
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<mods:number id="51305375F275E151CD502F76BEDA4164">1799</mods:number>
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<mods:url id="113682670FCEBDED9E904E46DBC8ECDA">https://biotaxa.org/Zootaxa/article/view/zootaxa.1799.1.1</mods:url>
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</mods:location>
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<mods:classification id="FF3BA88B4A549E033B9EF8616B816720">journal article</mods:classification>
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<mods:identifier id="896A8DBA2872D5513B2B4FC36A7DFFF5" type="DOI">10.11646/zootaxa.1799.1.1</mods:identifier>
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<mods:identifier id="2FD7627C806D3840B985DB1D4841EE1E" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="FE71A367E16A00576AF433E399E48D0C" type="Zenodo-Dep">5124771</mods:identifier>
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<treatment id="03C587F3FFF3FFB79BA19EF47CE0FA8A" LSID="urn:lsid:plazi:treatment:03C587F3FFF3FFB79BA19EF47CE0FA8A" httpUri="http://treatment.plazi.org/id/03C587F3FFF3FFB79BA19EF47CE0FA8A" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">
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<subSubSection id="C376656EFFF3FFB69BA19EF47CD7FF64" pageId="92" pageNumber="93" type="nomenclature">
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<paragraph id="8BD336E5FFF3FFB69BA19EF47C1AFF01" blockId="92.[151,718,151,212]" box="[151,515,151,177]" pageId="92" pageNumber="93">
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<heading id="D09B8189FFF3FFB69BA19EF47C1AFF01" bold="true" box="[151,515,151,177]" fontSize="11" level="1" pageId="92" pageNumber="93" reason="1">
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<taxonomicName id="4C6C4D66FFF3FFB69BA19EF47C1AFF01" authority="Reitter, 1873" authorityName="Reitter" authorityYear="1873" box="[151,515,151,177]" class="Insecta" family="Nitidulidae" genus="Pocadius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="nobilis">
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<emphasis id="B918EAF7FFF3FFB69BA19EF47C1AFF01" bold="true" box="[151,515,151,177]" pageId="92" pageNumber="93">
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<emphasis id="B918EAF7FFF3FFB69BA19EF47F47FF01" bold="true" box="[151,350,151,177]" italics="true" pageId="92" pageNumber="93">Pocadius nobilis</emphasis>
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<bibRefCitation id="EFFD4B14FFF3FFB69A539EF47C1AFF01" author="Reitter, E." box="[357,515,151,177]" pageId="92" pageNumber="93" pagination="5 - 194" refId="ref85782" refString="Reitter, E. (1873) Systematische Eintheilung der Nitidularien. Verhandlungen naturgischichte Vereines in Brunn, 12, 5 - 194." type="journal article" year="1873">Reitter, 1873</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BD336E5FFF3FFB69BA19ED97CD7FF64" blockId="92.[151,718,151,212]" box="[151,718,186,212]" pageId="92" pageNumber="93">(Figs. 33, 74, 117, 160, 201, 240, 260, 272–279)</paragraph>
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</subSubSection>
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<subSubSection id="C376656EFFF3FFB69BA19F697B99FD3C" pageId="92" pageNumber="93" type="description">
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<paragraph id="8BD336E5FFF3FFB69BA19F697B99FD3C" blockId="92.[151,1437,266,2012]" pageId="92" pageNumber="93">
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<emphasis id="B918EAF7FFF3FFB69BA19F697FBDFE94" bold="true" box="[151,420,266,292]" pageId="92" pageNumber="93">Specimens examined.</emphasis>
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<typeStatus id="54D78847FFF3FFB69A989F697C4CFE94" box="[430,597,266,292]" pageId="92" pageNumber="93" type="lectotype">LECTOTYPE</typeStatus>
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(RNH), here designated,
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<specimenCount id="9D6AFD6CFFF3FFB698BA9F697DB9FEAE" box="[908,928,266,286]" pageId="92" pageNumber="93" type="male">♂</specimenCount>
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: Japan; leg. Lewis; Coll. Reitter / nobilis; Japan, Rtts. [upside down yellow label] /
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<typeStatus id="54D78847FFF3FFB699BD9F517D1CFEFC" box="[651,773,306,332]" pageId="92" pageNumber="93" type="holotype">Holotypus</typeStatus>
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1873;
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<taxonomicName id="4C6C4D66FFF3FFB6986C9F517DD1FEFC" authorityName="Erichson. Veintiuno" authorityYear="1843" box="[858,968,306,332]" class="Insecta" family="Nitidulidae" genus="Pocadius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">Pocadius</taxonomicName>
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; nobilis; Reitter [rectangular label with red trim] /
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<taxonomicName id="4C6C4D66FFF3FFB69A2E9F397F92FEC4" authorityName="Reitter" authorityYear="1873" box="[280,395,346,372]" class="Insecta" family="Nitidulidae" genus="Pocadius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="nobilis">P. nobilis</taxonomicName>
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; m.; Japan [upside down label].
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<typeStatus id="54D78847FFF3FFB698159F397A0FFEC4" box="[803,1046,346,372]" pageId="92" pageNumber="93" type="paralectotype">PARALECTOTYPE</typeStatus>
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(BMNH), here designated,
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<specimenCount id="9D6AFD6CFFF3FFB69E469F397EFFFE2C" pageId="92" pageNumber="93" type="female">two female</specimenCount>
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specimens on same paper card with the following data labels: Type; H.T. [circular label with red trim] / Japan.; G. Lewis.; 1910-320 /
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<taxonomicName id="4C6C4D66FFF3FFB6993F9FC97C6FFE73" authorityName="Erichson. Veintiuno" authorityYear="1843" box="[521,630,426,451]" class="Insecta" family="Nitidulidae" genus="Pocadius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EAF7FFF3FFB6993F9FC97C6FFE73" box="[521,630,426,451]" italics="true" pageId="92" pageNumber="93">Pocadius</emphasis>
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</taxonomicName>
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;
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<taxonomicName id="4C6C4D66FFF3FFB699B19FC97CCEFE73" authorityName="Reitter" authorityYear="1873" box="[647,727,426,451]" class="Insecta" family="Nitidulidae" genus="Pocadius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="nobilis">
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<emphasis id="B918EAF7FFF3FFB699B19FC97CCEFE73" box="[647,727,426,451]" italics="true" pageId="92" pageNumber="93">nobilis</emphasis>
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</taxonomicName>
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m. Additionally,
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<specimenCount id="9D6AFD6CFFF3FFB698909FC97A2EFE74" box="[934,1079,426,452]" pageId="92" pageNumber="93" type="generic">3 specimens</specimenCount>
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(BMNH): same data labels as
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<typeStatus id="54D78847FFF3FFB69BA19FB17EE7FE5C" box="[151,254,466,492]" pageId="92" pageNumber="93" type="holotype">holotype</typeStatus>
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but without type labels. 1 (BMNH) with same data label as
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<typeStatus id="54D78847FFF3FFB698D49FB17A50FE5C" box="[994,1097,466,492]" pageId="92" pageNumber="93" type="holotype">holotype</typeStatus>
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but with additional label of “Higo.”. 1 (BMNH) with same data label as
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<typeStatus id="54D78847FFF3FFB699849F997D01FDA4" box="[690,792,506,532]" pageId="92" pageNumber="93" type="holotype">holotype</typeStatus>
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but additional label of “Hitoyoshi.; 3.V. – 8.V.81.”. 1 (BMNH): Japan. 1 (BMNH): in puffball /
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<collectingCountry id="F37B7675FFF3FFB699B69C417CF8FD8C" box="[640,737,546,572]" name="Japan" pageId="92" pageNumber="93">JAPAN</collectingCountry>
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:; J.E.A. Lewis.; B.M. 1933-490. 1 (BMNH):
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<collectingCountry id="F37B7675FFF3FFB69FDB9C417B50FD8C" box="[1261,1353,546,572]" name="Japan" pageId="92" pageNumber="93">JAPAN</collectingCountry>
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; Kobe; Shinohara; 30-IX-’30; J.E.A. Lewis /
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<collectingCountry id="F37B7675FFF3FFB699679C297CABFDD4" box="[593,690,586,612]" name="Japan" pageId="92" pageNumber="93">JAPAN</collectingCountry>
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:; J.E.A. Lewis.; B.M. 1933-490. 1 (RMNH):
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<collectingCountry id="F37B7675FFF3FFB69FF49C297B33FDD4" box="[1218,1322,586,612]" name="South Korea" pageId="92" pageNumber="93">KOREA</collectingCountry>
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, No. 693; Kangwon Prov.; Mts. Kumgang-san; singled,
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<date id="FFD21025FFF3FFB699869C117D24FD3C" box="[688,829,626,652]" pageId="92" pageNumber="93" value="1980-09-16">16-IX-1980</date>
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; leg. Topál and Forró. 4 (ZMHB): ID labels only.
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</paragraph>
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</subSubSection>
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<subSubSection id="C376656EFFF3FFB69BF09CF97AC0FC9C" pageId="92" pageNumber="93" type="diagnosis">
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<paragraph id="8BD336E5FFF3FFB69BF09CF97AC0FC9C" blockId="92.[151,1437,266,2012]" pageId="92" pageNumber="93">
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<emphasis id="B918EAF7FFF3FFB69BF09CF97F5FFD04" bold="true" box="[198,326,666,692]" pageId="92" pageNumber="93">Diagnosis.</emphasis>
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The peculiar color pattern with dark medial longitudinal “stripe” on pronotum distinguishes
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<taxonomicName id="4C6C4D66FFF3FFB69EB09CF97EFEFD6B" authorityName="Reitter" authorityYear="1873" class="Insecta" family="Nitidulidae" genus="Pocadius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="nobilis">
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<emphasis id="B918EAF7FFF3FFB69EB09CF97EFEFD6B" italics="true" pageId="92" pageNumber="93">P. nobilis</emphasis>
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</taxonomicName>
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from other
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<collectingRegion id="49A8F807FFF3FFB69A479CA17FC8FD6C" box="[369,465,706,732]" country="Democratic Republic of the Congo" name="Orientale" pageId="92" pageNumber="93">Oriental</collectingRegion>
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and Old World
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<taxonomicName id="4C6C4D66FFF3FFB699B89CA17CE2FD6B" authorityName="Erichson. Veintiuno" authorityYear="1843" box="[654,763,706,731]" class="Insecta" family="Nitidulidae" genus="Pocadius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EAF7FFF3FFB699B89CA17CE2FD6B" box="[654,763,706,731]" italics="true" pageId="92" pageNumber="93">Pocadius</emphasis>
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</taxonomicName>
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, as well as deep punctation of pygidium; non-trapezoidal anterior pronotal margin; densely setose tegmen; sub-parallel sided median lobe with angular apical margins; basally fused ejaculatory rods; deeply cleft and separated basal piece of internal sac sclerites.
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</paragraph>
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</subSubSection>
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<subSubSection id="C376656EFFF3FFB79BF09D597D8BFB72" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93" type="description">
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<paragraph id="8BD336E5FFF3FFB69BF09D597F8BFC44" blockId="92.[151,1437,266,2012]" pageId="92" pageNumber="93">
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<emphasis id="B918EAF7FFF3FFB69BF09D597F6DFCE4" bold="true" box="[198,372,826,852]" pageId="92" pageNumber="93">Redescription</emphasis>
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. Length
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<quantity id="4C949B00FFF3FFB69AEB9D597C2BFCE4" box="[477,562,826,852]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="92" pageNumber="93" unit="mm" value="3.6">3.6mm</quantity>
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, Width 2.0mm, Depth
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<quantity id="4C949B00FFF3FFB698089D597D85FCE4" box="[830,924,826,852]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="92" pageNumber="93" unit="mm" value="0.9">0.9 mm</quantity>
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. Body convex, shining, light reddish brown with the following much darker brown to black: antennal club, temples of head, pronotal disc from apex to base, scutellum, lateral region of elytra from humeri to apices as well as apical 0.5 of elytra from suture to lateral explanation leaving. Pubescence of modest length, golden, sparsely covering dorsum. Pronotum more explanate than elytra.
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</paragraph>
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<paragraph id="8BD336E5FFF3FFB69BF09A617D08FA34" blockId="92.[151,1437,266,2012]" pageId="92" pageNumber="93">Head subequal in length and width, broadly triangular. Head surface irregularly punctate, with large and small punctures interspersed, becoming more obsolete near clypeus, interspaces smooth to alutaceous. Small punctures 3X eye facet, large ones 3–5X diameter of small punctures. Antennal club compact, broadly oval, asymmetrical with the last antennomere subequal to previous two combined. Antennomere 8 strongly flattened disc-like, 6 and 7 flattened and disc-like but not to the extent of 8, 6–8 combined length subequal to antennomere 9. Antennal scape asymmetrical, broadly convex dorsally, 2X as long as pedicel. Pedicel subcylindrical. Antennal segment 3 subequal to pedicel, gradually expanded apically. Segments 4 and 5 subquadrate, 0.5 length of segment 3. Elevated region of mentum with anterior margin broadly evenly convex, overall hemispheric, in lateral view somewhat convex. Mental surface with minute faint punctures sparsely distributed, interspaces smooth with some microreticulation.</paragraph>
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<paragraph id="8BD336E5FFF3FFB79BF09BF17AB3FE12" blockId="92.[151,1437,266,2012]" lastBlockId="93.[151,1436,152,1338]" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">Pronotum widest near middle (W:L = 2:1), evenly rounded laterally, anterior and posterior angles somewhat rounded, anterior margin broadly shallowly concave. Pronotal surface with large and small punctures interspersed. Large and small punctures similar to respective large and small punctures on head; interspaces 1–2 puncture diameters apart, smooth to alutaceous. Prosternum and epimeron irregularly punctate, punctures equal to large punctures on head, interspaces alutaceous. Prosternal process in lateral view with anterior end slightly arcuate, broad convexity medially over procoxae and posteriorly with short flat shelf; in lateral view posterior margin with medial concavity. Scutellum large, broadly rounded at apex. Scutellar surface sparsely punctate with minute punctures 0.5 diameter of small punctures on head, interspaces smooth. Eytral surface with serial rows large punctures 2X diameter of large punctures on head, each giving rise to decumbent setae. Serial small punctures 0.2 diameter of large punctures, giving rise to decumbent setae; interspaces alutaceous, large punctures separated from each other within a row by 0.5 diameter, small punctures separated by 2–3 diameters. Large and small puncture rows separated by 2 small puncture diameters. Mesoventrite extending to midway between mesocoxae with faint medial carina, deeply concave for reception of metaventrite. Mesosternal punctures aggregated near posterior margin. Metaventrite wider than long (W:L = 2.5:1). Metasternal punctures faint, equal to large punctures on head, interspaces on disc smooth to alutaceous, becoming com- pletely alutaceous laterally. Metepisternum broad, width to length 1:3.8, surface with faint large punctation similar in size to those on metaventrite, interspaces alutaceous with some microreticulation and granular areas. First abdominal sternite with broad almost truncate process 2.5X longer than second sternite. Sternites 2–4 subequal. Abdominal sternite 1 with faint punctures in all but anterior 0.33. Pygidium broadly triangular with surface densely punctate, punctures equal to large punctures on elytra, interspaces microreticulate, punctures separated by 0.25–0.2 diameter. Hypopygidium subequal to first abdominal sternite; densely punctate, punctures equal to those on sternites 2–4, interspaces rugose, separated by 0.25 diameter.</paragraph>
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<paragraph id="8BD336E5FFF2FFB79BF09FD37B9FFDDA" blockId="93.[151,1436,152,1338]" pageId="93" pageNumber="94">Protibia with apical tooth 1.5X length of tarsomere 2. Outer apical notch indistinct. Inner apical spine subequal to 1.5 length of tarsomere 1. Mesotibia modestly armed with row of slender spines along lateral edge and other stiff setae on ventral and dorsal surfaces. Outer apical tooth short and robust, 0.5 length of inner apical spine. Inner apical spine equal to tarsomeres 1–2 combined. Metatibia similarly armed as mesotibia. Outer apical process robust, 0.5 length of inner apical spine. Inner apical spine equal to tarsomeres 1–2 combined.</paragraph>
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<paragraph id="8BD336E5FFF2FFB79BF09C1B7C4EFC1A" blockId="93.[151,1436,152,1338]" pageId="93" pageNumber="94">Male genitalia moderately sclerotized. Anal sclerite with apex moderately fimbriate (Fig. 33). Spiculum gastrale with (Fig. 74) lateral region broadly rounded and widely explanate, medial border evenly concave, apical border with numerous elongate setae. Tegmen broadly rounded to truncate apically (Fig. 117), longer than wide (w:l = 1.3.1), lateral row of setae visible from median fossa to prior to apex, inner row of setae incomplete apically extending from apex of median fossa to prior to apex. Median lobe large and robust, sides subparallel laterally becoming angulate apically, 0.6 length of tegmen, apical opening well-developed (Fig. 160). Ejaculatory rods fused to each other basally but not fused to basal piece, basal piece deeply cleft apically, narrowed proximally (Fig. 201).</paragraph>
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<paragraph id="8BD336E5FFF2FFB79BF09DDB7EF0FBC2" blockId="93.[151,1436,152,1338]" pageId="93" pageNumber="94">Female genitalia overall moderately sclerotized. Gonocoxites with two baso-lateral prominences, basal ridge giving rise to two medial oblique baculi extending apicolaterally, 0.33 length of basal border. Gonocoxal apices narrowly separated with intragonocoxal invagination evenly rounded at base, gonocoxal tips evenly rounded at apex, each apex with a small lateral depression that gives rise to 4–5 primary seta (Fig. 240), tooth absent.</paragraph>
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<paragraph id="8BD336E5FFF2FFB79BF09A1C7A71FB2A" blockId="93.[151,1436,152,1338]" box="[198,1128,1151,1178]" pageId="93" pageNumber="94">
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<emphasis id="B918EAF7FFF2FFB79BF09A1C7F5AFB29" bold="true" box="[198,323,1151,1177]" pageId="93" pageNumber="94">Variation.</emphasis>
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Male specimens tend to have a broader protibial apico-lateral notch.
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</paragraph>
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<paragraph id="8BD336E5FFF2FFB79BF09AC47D8BFB72" blockId="93.[151,1436,152,1338]" box="[198,914,1191,1218]" pageId="93" pageNumber="94">
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<emphasis id="B918EAF7FFF2FFB79BF09AC47FDAFB71" bold="true" box="[198,451,1191,1217]" pageId="93" pageNumber="94">Seasonality/Habitat.</emphasis>
|
||
Known to occur from May-September.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376656EFFF2FFB79BF09AAC7CE0FA8A" pageId="93" pageNumber="94" type="distribution">
|
||
<paragraph id="8BD336E5FFF2FFB79BF09AAC7D78FAA2" blockId="93.[151,1436,152,1338]" pageId="93" pageNumber="94">
|
||
<emphasis id="B918EAF7FFF2FFB79BF09AAC7F7EFB59" bold="true" box="[198,359,1231,1257]" pageId="93" pageNumber="94">Distribution.</emphasis>
|
||
Found on the major islands of
|
||
<collectingCountry id="F37B7675FFF2FFB799EC9AB37D38FB5A" box="[730,801,1232,1258]" name="Japan" pageId="93" pageNumber="94">Japan</collectingCountry>
|
||
. The Korean specimen listed above is a new distribution record and extends the known range into mainland Asia.
|
||
</paragraph>
|
||
<paragraph id="8BD336E5FFF2FFB79BF09B7C7CE0FA8A" blockId="93.[151,1436,152,1338]" box="[198,761,1311,1338]" pageId="93" pageNumber="94">
|
||
<emphasis id="B918EAF7FFF2FFB79BF09B7C7F0AFA89" bold="true" box="[198,275,1311,1337]" pageId="93" pageNumber="94">Notes.</emphasis>
|
||
Host information available as “puffball.”
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |