272 lines
32 KiB
XML
272 lines
32 KiB
XML
<document id="17F1AAE59D79B6AD53CAAAAA3AD44323" ID-CLB-Dataset="8562" ID-DOI="10.5252/geodiversitas2020v42a20" ID-GBIF-Dataset="ad4dbfb6-14ce-48e2-8fe9-99fa26a9bcf4" ID-ISSN="1638-9395" ID-Zenodo-Dep="4447563" ID-ZooBank="urn:lsid:zoobank.org:pub:8FF2A078-CE45-4BF1-A681-00136F57375E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1610989940068" checkinUser="felipe" docAuthor="Ivanov, Martin, Čerňanský, Andrej, Bonilla-Salomón, Isaac & Luján, Àngel Hernández" docDate="2020" docId="03C587C7430DFFEFFC16FE2A4A12FC1D" docLanguage="en" docName="Geodiversitas.42.20.343-376.pdf" docOrigin="Geodiversitas 42 (20)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pseudopus laurillardi" docType="treatment" docVersion="10" lastPageNumber="351" masterDocId="FFFCFFBF4304FFE4FFBCFF9C484CFFD2" masterDocTitle="Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications" masterLastPageNumber="376" masterPageNumber="343" pageNumber="350" updateTime="1698925529958" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="EDE07EE20B42FE9E8F727B6274D49811">
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<mods:title id="80305D676B5073DDF4AB29D1783CCD70">Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications</mods:title>
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<mods:name id="314F88046C1C3A2B5E59536C5211DC73" type="personal">
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<mods:roleTerm id="0B0A6BA0BE8D92D76F25AB6D7492DC0B">Author</mods:roleTerm>
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<mods:namePart id="BF7EDEE7F71031B1EB2D87372EDF1E2D">Ivanov, Martin</mods:namePart>
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<mods:roleTerm id="105C02D9C7D806367915D0C67E43B6B7">Author</mods:roleTerm>
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<mods:namePart id="C6A5A402BD842A421D673A56AC26ECBA">Čerňanský, Andrej</mods:namePart>
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</mods:name>
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<mods:name id="1177B059968F4CC04E9305285AF827FB" type="personal">
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<mods:roleTerm id="85F453358C025583A0E08C99F2A6A3FA">Author</mods:roleTerm>
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<mods:namePart id="30BB63808A18FE260CFC80899088F9FB">Bonilla-Salomón, Isaac</mods:namePart>
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</mods:name>
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<mods:name id="1AB1D75C4BCFFD1A2E53A68352C17080" type="personal">
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<mods:role id="704ED9E4334167E4032240C115E9EA6E">
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<mods:roleTerm id="796A4E486D3F6134B9413903544AE0D9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="008D99003890EEFED60A980F52AEAE98">Luján, Àngel Hernández</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3CB0415168E0AF3D9D7000005464E02C">text</mods:typeOfResource>
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<mods:titleInfo id="1B9A739A66F62541E478023487060E62">
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<mods:title id="6DA4CE3A32207F8982DB1B6634336A26">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="A22C44A9CBC45E1E7E248D1390B91882">
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<mods:date id="51B649B17F34FC6C2415E44328C85A34">2020</mods:date>
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<mods:detail id="0B6D24A29E9D41F381B83891547135A5" type="pubDate">
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<mods:number id="FE54AD437F8884B37C8CB5B31B585360">2020-08-28</mods:number>
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<mods:detail id="1232CB2528D933D29D33B3D82275F15B" type="volume">
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<mods:number id="329FEDD946A970C9E8932A87E3CEC692">42</mods:number>
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</mods:detail>
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<mods:detail id="CDA321CD698E55BA178B5BD5B55FACA2" type="issue">
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<mods:number id="6EBE5BC9F02C27AE98B4B023D1C02957">20</mods:number>
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<mods:start id="E5464AACF501673249B899CA46E6B97B">343</mods:start>
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<mods:end id="F19DB07E47E5A7FDBB435F34605A5C3E">376</mods:end>
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<mods:classification id="E4E39F5CD63058FA4417D84B282C95A4">journal article</mods:classification>
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<mods:identifier id="1E5F5B3D29E02670307D02F06F38B561" type="CLB-Dataset">8562</mods:identifier>
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<mods:identifier id="2F2B151219D92D4967E9370CF25573EF" type="DOI">10.5252/geodiversitas2020v42a20</mods:identifier>
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<mods:identifier id="6329EA105458A4EA755FA67B1D1BE731" type="GBIF-Dataset">ad4dbfb6-14ce-48e2-8fe9-99fa26a9bcf4</mods:identifier>
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<mods:identifier id="F1A94AACD79134824A443F8A7F327F98" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="67D3F392A276DCED24133E6885CF35FC" type="Zenodo-Dep">4447563</mods:identifier>
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</mods:mods>
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<treatment id="03C587C7430DFFEFFC16FE2A4A12FC1D" ID-DOI="http://doi.org/10.5281/zenodo.4488219" ID-GBIF-Taxon="177200743" ID-Zenodo-Dep="4488219" LSID="urn:lsid:plazi:treatment:03C587C7430DFFEFFC16FE2A4A12FC1D" httpUri="http://treatment.plazi.org/id/03C587C7430DFFEFFC16FE2A4A12FC1D" lastPageId="11" lastPageNumber="351" pageId="9" pageNumber="350">
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||
<subSubSection id="C376655A430DFFEDFC16FE2A4D7DFE03" box="[938,1329,438,465]" pageId="9" pageNumber="350" type="nomenclature">
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||
<paragraph id="8BD336D1430DFFEDFC16FE2A4D7DFE03" blockId="9.[938,1329,438,465]" box="[938,1329,438,465]" pageId="9" pageNumber="350">
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<heading id="D09B81BD430DFFEDFC16FE2A4D7DFE03" box="[938,1329,438,465]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="350" reason="2">
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||
<taxonomicName id="4C6C4D52430DFFEDFC16FE2A4D7DFE03" authority="(Lartet, 1851)" baseAuthorityName="Lartet" baseAuthorityYear="1851" box="[938,1329,438,465]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="laurillardi">
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<emphasis id="B918EAC3430DFFEDFC16FE2A4CDFFE03" bold="true" box="[938,1171,438,465]" italics="true" pageId="9" pageNumber="350">Pseudopus laurillardi</emphasis>
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(Lartet, 1851)
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</taxonomicName>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376655A430DFFEDFB91FE4A4CE2FE23" box="[1069,1198,470,497]" pageId="9" pageNumber="350" type="description">
|
||
<paragraph id="8BD336D1430DFFEDFB91FE4A4CE2FE23" blockId="9.[1069,1198,470,497]" box="[1069,1198,470,497]" pageId="9" pageNumber="350">
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||
(
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<figureCitation id="13572A54430DFFEDFB8AFE4A4C39FE23" box="[1078,1141,470,497]" captionStart="FIG" captionStartId="11.[133,144,1917,1935]" captionTargetId="figure-20@12.[151,1436,215,1878]" captionTargetPageId="12" captionText="FIG. 5. — Pseudopus from the early Miocene (MN 4) of MWQ. Pseudopus laurillardi: parietal Pal. 1574 (2/2003 Reptile Joint) in (A) dorsal and (B) ventral aspects; parietal Pal. 1402 (1/2001 Turtle Joint) in (C) dorsal and (D) ventral aspects. Pseudopus sp.: left maxilla Pal. 1575 (2/2003 Reptile Joint) in (E) lateral and (F) medial aspects. Left quadrate Pal. 1576 (2/2003 Reptile Joint) in (G) lateral and (H) medial aspects. Right dentary Pal. 1403 (1/2001 Turtle Joint) in (I) lateral, (J) medial and (K) dorsal aspects. Trunk vertebra Pal. 1404 (1/2001 Turtle Joint) in (L) lateral, (M) dorsal, (N) ventral, (O) anterior and (P) posterior aspects. Trunk vertebra 1405 (1/2001 Turtle Joint) in (Q) anterior aspect. Abbreviations: are, arcuate edge; cc, cephalic condyle; cd, condyle; cf, facet for" figureDoi="http://doi.org/10.5281/zenodo.4487033" httpUri="https://zenodo.org/record/4487033/files/figure.png" pageId="9" pageNumber="350">Fig. 5</figureCitation>
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A-D)
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A430DFFEDFC91FD844D19FDE2" box="[813,1365,536,560]" pageId="9" pageNumber="350" type="reference_group">
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||
<paragraph id="8BD336D1430DFFEDFC91FD844D19FDE2" blockId="9.[813,1365,536,560]" box="[813,1365,536,560]" pageId="9" pageNumber="350">
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<treatmentCitationGroup id="AB7C11FF430DFFEDFC91FD844D19FDE2" box="[813,1365,536,560]" pageId="9" pageNumber="350">
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<taxonomicName id="4C6C4D52430DFFEDFC91FD844BC7FDE2" authorityName="Merrem" authorityYear="1820" box="[813,907,536,560]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="sp.">
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<emphasis id="B918EAC3430DFFEDFC91FD844BC7FDE2" box="[813,907,536,560]" italics="true" pageId="9" pageNumber="350">Pseudopus</emphasis>
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</taxonomicName>
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sp. –
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<treatmentCitation id="0ACD10C0430DFFEDFC79FD854CE4FDE2" author="IVANOV M. & MUSIL R. & BRZOBOHATY R." box="[965,1192,536,560]" page="229" pageId="9" pageNumber="350" year="2006">
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||
<bibRefCitation id="EFFD4B20430DFFEDFC79FD854CE4FDE2" author="IVANOV M. & MUSIL R. & BRZOBOHATY R." box="[965,1192,536,560]" pageId="9" pageNumber="350" pagination="223 - 239" refId="ref24955" refString="IVANOV M., MUSIL R. & BRZOBOHATY R. 2006. - Terrestrial and Marine Faunas from the Miocene Deposits of the Mokra Plateau (Drahany Upland, Czech Republic) - Impact on Palaeogeography. Beitrage zur Palaontologie 30: 223 - 239." type="journal article" year="2006">
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||
Ivanov
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<emphasis id="B918EAC3430DFFEDFBB1FD854C70FDE2" box="[1037,1084,536,560]" italics="true" pageId="9" pageNumber="350">et al.</emphasis>
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2006: 229
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</bibRefCitation>
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</treatmentCitation>
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||
, table 2 (in part).
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376655A430DFFEDFC91FDCE4C60FD4C" pageId="9" pageNumber="350" type="materials_examined">
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<paragraph id="8BD336D1430DFFEDFC91FDCE4C60FD4C" blockId="9.[813,1457,593,670]" pageId="9" pageNumber="350">
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||
MATERIAL. —
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<materialsCitation id="3B043C8C430DFFEDFC7BFDCD4C6CFD4C" ID-GBIF-Occurrence="3027945302" pageId="9" pageNumber="350">
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MWQ, early Miocene, Burdigalian, Orleanian,
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<collectionCode id="ED7DAE14430DFFEDFC93FDF04B16FD51" box="[815,858,620,643]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="9" pageNumber="350">MN</collectionCode>
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4: 1/2001 Turtle Joint: parietal (Pal. 1402). 2/2003 Reptile Joint: parietal (Pal. 1574
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</materialsCitation>
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||
).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376655A430DFFEDFC91FD4A4D4BF839" pageId="9" pageNumber="350" type="description">
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<paragraph id="8BD336D1430DFFEDFC91FD4A4D4BF839" blockId="9.[811,1463,726,2027]" pageId="9" pageNumber="350">
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||
DESCRIPTION
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<emphasis id="B918EAC3430DFFEDFC91FD694B32FCDD" box="[813,894,757,783]" italics="true" pageId="9" pageNumber="350">Parietal</emphasis>
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Two parietals are partially preserved. The specimen Pal. 1574 represents the posterior region of the parietal table, together with the base of the right supratemporal process, whereas only the central portion of the right side of the parietal is preserved in the second specimen (Pal. 1402). The anterior region of the preserved portion of the parietal table is completely covered by osteodermal shields fused to the bone. These shields bear ornamentation which consists of pits, small ridges and short grooves. The small occipital shield, located in the posterior mid-region, is trapezoidal in shape. Its anteroposteriorlength is less than that of the posteriorly located smooth area (area levis) in both specimens. However, the smooth area is large only in specimen Pal. 1574. The interparietal shield, located anterior to the occipital shield, is only hardly distinguished from the large parietal shields located laterally. There is no parietal foramen within the preserved portion of the bone. The base of the supratemporal process is broad. Here, the arcuate edge (carina arcuata;
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<emphasis id="B918EAC3430DFFEDFABCFAC84D74FABF" box="[1280,1336,1364,1389]" italics="true" pageId="9" pageNumber="350">sensu</emphasis>
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<bibRefCitation id="EFFD4B20430DFFEDFAFCFACF4BF6FA5F" author="KLEMBARA J. & DOBIAsOVA K. & HAIN M. & YARYHIN O." pageId="9" pageNumber="350" pagination="460 - 502" refId="ref25649" refString="KLEMBARA J., DOBIAsOVA K., HAIN M. & YARYHIN O. 2017 a. - Skull anatomy and ontogeny of legless lizard Pseudopus apodus (Pallas, 1775): heterochronic influences on form. The Anatomical Record 300 (3): 460 - 502. https: // doi. org / 10.1002 / ar. 23532" type="journal article" year="2017">
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Klembara
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<emphasis id="B918EAC3430DFFEDFC91FAE84B2BFA5F" box="[813,871,1395,1421]" italics="true" pageId="9" pageNumber="350">et al.</emphasis>
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2017a
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</bibRefCitation>
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) is developed. The supratemporal process gradually widens distally (this can be observed especially in specimen Pal. 1402). On the internal side, the process bears a robust ventrolateral ridge. In the larger specimen, Pal. 1574, a free ventrolateral surface is present lateral to the ridge. This is absent in the smaller Pal. 1402. In this specimen, a longitudinal facet for the supratemporal can be observed on the posterolateral side of the ridge. This facet reaches approximately to the level of the posterior margin of the parietal table. On the internal side of specimen Pal. 1574, a large oval parietal fossa is located in the posterior mid-region. It is bordered ventrally by the parietal lamina. Posterior to the parietal fossa, a postfoveal crest (crista postfovealis) is located. Between the postfoveal crests and the posterior margin of the parietal table, a shal - low parietal notch is present. The cranial parietal ridges (cristae cranii parietalis) are partly preserved, the right one can be observed in the specimen Pal. 1402. Lateral to the cranial ridge, muscular facies are well developed, forming the lateral border of the parietal.
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</paragraph>
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</subSubSection>
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<caption id="DF136659430EFFEEFF38F90A4986F93E" ID-DOI="http://doi.org/10.5281/zenodo.4447573" ID-Zenodo-Dep="4447573" httpUri="https://zenodo.org/record/4447573/files/figure.png" pageId="10" pageNumber="351" startId="10.[132,143,1686,1704]" targetBox="[132,1455,215,1644]" targetPageId="10">
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<paragraph id="8BD336D1430EFFEEFF38F90A4986F93E" blockId="10.[132,1457,1686,1772]" pageId="10" pageNumber="351">
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FIG. 4. —
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<taxonomicName id="4C6C4D52430EFFEEFF6FF90A497AF97A" authorityName="Estes, de Queiroz & Gauthier" authorityYear="1988" box="[211,310,1686,1704]" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="346" rank="superFamily" superFamily="Lacertoidea">Lacertoidea</taxonomicName>
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indet. (
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<emphasis id="B918EAC3430EFFEEFECDF90A4932F97A" bold="true" box="[369,382,1686,1704]" pageId="10" pageNumber="351">A</emphasis>
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,
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<emphasis id="B918EAC3430EFFEEFE34F90A49D9F97A" bold="true" box="[392,405,1686,1704]" pageId="10" pageNumber="351">B</emphasis>
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) and?
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<taxonomicName id="4C6C4D52430EFFEEFE6DF90A4A62F97A" authorityName="Oppel" authorityYear="1811" box="[465,558,1686,1704]" kingdom="Animalia" pageId="10" pageNumber="350" rank="superFamily" superFamily="Scincoidea">Scincoidea</taxonomicName>
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indet. (
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<emphasis id="B918EAC3430EFFEEFDD6F90A4A34F97A" bold="true" box="[618,632,1686,1704]" pageId="10" pageNumber="351">C</emphasis>
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||
,
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||
<emphasis id="B918EAC3430EFFEEFD3DF90A4AC3F97A" bold="true" box="[641,655,1686,1704]" pageId="10" pageNumber="351">D</emphasis>
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) from the early Miocene (MN 4) of MWQ. Left dentary Pal. 1571 (2/2003 Reptile Joint) in (
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<emphasis id="B918EAC3430EFFEEFADBF90A4D38F97A" bold="true" box="[1383,1396,1686,1704]" pageId="10" pageNumber="351">A</emphasis>
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) lateral and (
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<emphasis id="B918EAC3430EFFEEFF12F93148F7F96D" bold="true" box="[174,187,1709,1727]" pageId="10" pageNumber="351">B</emphasis>
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) medial aspects with detail of teeth. Left maxilla Pal. 1573 (2/2003 Reptile Joint) in (
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<emphasis id="B918EAC3430EFFEEFCC7F9314BC5F96D" bold="true" box="[891,905,1709,1727]" pageId="10" pageNumber="351">C</emphasis>
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) lateral and (
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<emphasis id="B918EAC3430EFFEEFC49F9314C4FF96D" bold="true" box="[1013,1027,1709,1727]" pageId="10" pageNumber="351">D</emphasis>
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) medial aspects with detail of teeth. Abbreviations:
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<emphasis id="B918EAC3430EFFEEFF38F95F48D6F907" bold="true" box="[132,154,1731,1749]" pageId="10" pageNumber="351">dc</emphasis>
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, dental crest;
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<emphasis id="B918EAC3430EFFEEFEA8F95F497CF907" bold="true" box="[276,304,1731,1749]" pageId="10" pageNumber="351">cm</emphasis>
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, carina maxillaris;
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<emphasis id="B918EAC3430EFFEEFE72F95F4995F907" bold="true" box="[462,473,1731,1749]" pageId="10" pageNumber="351">lf</emphasis>
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, labial foramina;
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<emphasis id="B918EAC3430EFFEEFDD6F95F4ACAF907" bold="true" box="[618,646,1731,1749]" pageId="10" pageNumber="351">Mg</emphasis>
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||
, Meckel`s groove;
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<emphasis id="B918EAC3430EFFEEFC9BF95F4B71F907" bold="true" box="[807,829,1731,1749]" pageId="10" pageNumber="351">np</emphasis>
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, nasal process;
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<emphasis id="B918EAC3430EFFEEFC75F95F4BA4F907" bold="true" box="[969,1000,1731,1749]" pageId="10" pageNumber="351">sbs</emphasis>
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, subdental shelf;
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<emphasis id="B918EAC3430EFFEEFBC3F95F4CD6F907" bold="true" box="[1151,1178,1731,1749]" pageId="10" pageNumber="351">srs</emphasis>
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, supradental shelf;
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<emphasis id="B918EAC3430EFFEEFAFEF95F4D1FF907" bold="true" box="[1346,1363,1731,1749]" pageId="10" pageNumber="351">st</emphasis>
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, striations. Scale bars: A-D, 1 mm; B’, D’, 100 µm.
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</paragraph>
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</caption>
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<subSubSection id="C376655A430EFFEFFF38F8AE4A12FC1D" lastPageId="11" lastPageNumber="352" pageId="10" pageNumber="351" type="discussion">
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<paragraph id="8BD336D1430EFFEEFF38F8AE48A4F898" blockId="10.[131,775,1842,2030]" box="[132,232,1842,1868]" pageId="10" pageNumber="351">REMARKS</paragraph>
|
||
<paragraph id="8BD336D1430EFFEFFF38F8CD4A12FC1D" blockId="10.[131,775,1842,2030]" lastBlockId="11.[130,777,215,975]" lastPageId="11" lastPageNumber="352" pageId="10" pageNumber="351">
|
||
The presence of muscular facies (see
|
||
<bibRefCitation id="EFFD4B20430EFFEEFDACF8CD4AB2F8BE" author="KLEMBARA J. & DOBIAsOVA K. & HAIN M. & YARYHIN O." box="[528,766,1873,1900]" pageId="10" pageNumber="351" pagination="460 - 502" refId="ref25649" refString="KLEMBARA J., DOBIAsOVA K., HAIN M. & YARYHIN O. 2017 a. - Skull anatomy and ontogeny of legless lizard Pseudopus apodus (Pallas, 1775): heterochronic influences on form. The Anatomical Record 300 (3): 460 - 502. https: // doi. org / 10.1002 / ar. 23532" type="journal article" year="2017">
|
||
Klembara
|
||
<emphasis id="B918EAC3430EFFEEFD3CF8CE4AF8F8B9" box="[640,692,1873,1899]" italics="true" pageId="10" pageNumber="351">et al.</emphasis>
|
||
2017a
|
||
</bibRefCitation>
|
||
) on both parietals and the presence of the postfoveal crest allow the allocation of this material to
|
||
<taxonomicName id="4C6C4D52430EFFEEFDA6F80D4ACDF879" authorityName="Merrem" authorityYear="1820" box="[538,641,1937,1963]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="351" phylum="Chordata" rank="genus">
|
||
<emphasis id="B918EAC3430EFFEEFDA6F80D4ACDF879" box="[538,641,1937,1963]" italics="true" pageId="10" pageNumber="351">Pseudopus</emphasis>
|
||
</taxonomicName>
|
||
(although it should be noted that short postfoveal crests are also present in
|
||
<taxonomicName id="4C6C4D52430EFFEEFF1FF84D4910F839" box="[163,348,2001,2027]" class="Reptilia" family="Anguidae" genus="Ophisaurus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="holeci">
|
||
<emphasis id="B918EAC3430EFFEEFF1FF84D4910F839" box="[163,348,2001,2027]" italics="true" pageId="10" pageNumber="351">Ophisaurus holeci</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<bibRefCitation id="EFFD4B20430EFFEEFED2F84D4A54F839" author="KLEMBARA J." box="[366,536,2001,2027]" pageId="10" pageNumber="351" pagination="854 - 866" refId="ref25357" refString="KLEMBARA J. 2012. - A new species of Pseudopus (Squamata, Anguidae) from the early Miocene of North-West Bohemia (Czech Republic). Journal of Vertebrate Paleontology 32: 854 - 866. https: // doi. org / 10.1080 / 02724634.2012.670177" type="journal article" year="2012">Klembara 2012</bibRefCitation>
|
||
; Čerňanský & Klem- bara 2017] and in some cases, narrow muscular facies can be observed in very large adult individuals of this taxon as well [see
|
||
<bibRefCitation id="EFFD4B20430EFFEEFCE0F8ED4C02F859" author="KLEMBARA J. & HAIN M. & CERNANSKY A." box="[860,1102,1905,1931]" pageId="10" pageNumber="351" pagination="1 - 12" refId="ref25712" refString="KLEMBARA J., HAIN M. & CERNANSKY A. 2017 b. - The first record of anguine lizards (anguimorpha, anguidae) from the early Miocene locality Ulm - Westtangente in Germany. Historical Biology: 1 - 12. https: // doi. org / 10.1080 / 08912963.2017.1416469" type="book chapter" year="2017">
|
||
Klembara
|
||
<emphasis id="B918EAC3430EFFEEFC71F8EE4C4DF859" box="[973,1025,1905,1931]" italics="true" pageId="10" pageNumber="351">et al.</emphasis>
|
||
2017b
|
||
</bibRefCitation>
|
||
]). Three species of
|
||
<taxonomicName id="4C6C4D52430EFFEEFA9DF8ED4DC4F859" authorityName="Merrem" authorityYear="1820" box="[1313,1416,1905,1931]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="351" phylum="Chordata" rank="genus">
|
||
<emphasis id="B918EAC3430EFFEEFA9DF8ED4DC4F859" box="[1313,1416,1905,1931]" italics="true" pageId="10" pageNumber="351">Pseudopus</emphasis>
|
||
</taxonomicName>
|
||
are documented from early Miocene deposits of Europe:
|
||
<taxonomicName id="4C6C4D52430EFFEEFAD1F80E4B3AF819" baseAuthorityName="Lartet" baseAuthorityYear="1851" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="laurillardi">
|
||
<emphasis id="B918EAC3430EFFEEFAD1F80E4B3AF819" italics="true" pageId="10" pageNumber="351">P. laurillardi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D52430EFFEEFC39F82E4C6AF819" authorityName="Klembara" authorityYear="2012" box="[901,1062,1969,1995]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="ahnikoviensis">
|
||
<emphasis id="B918EAC3430EFFEEFC39F82E4C6AF819" box="[901,1062,1969,1995]" italics="true" pageId="10" pageNumber="351">P. ahnikoviensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C4D52430EFFEEFBDCF82E4C9FF819" box="[1120,1235,1969,1995]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="confertus">
|
||
<emphasis id="B918EAC3430EFFEEFBDCF82E4C9FF819" box="[1120,1235,1969,1995]" italics="true" pageId="10" pageNumber="351">P. confertus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFFD4B20430EFFEEFAB0F82D4B29F839" author="KLEMBARA J. & BOHME M. & RUMMEL M." pageId="10" pageNumber="351" pagination="159 - 196" refId="ref25523" refString="KLEMBARA J., BOHME M. & RUMMEL M. 2010. - Revision of the anguine lizard Pseudopus laurillardi (Squamata, Anguidae) from the Miocene of Europe, with comments on paleoecology. Journal of Paleontology 84 (2): 159 - 196. https: // doi. org / 10.1666 / 09 - 033 R 1.1" type="journal article" year="2010">
|
||
Klembara
|
||
<emphasis id="B918EAC3430EFFEEFAC7F82E4DFCF819" box="[1403,1456,1969,1995]" italics="true" pageId="10" pageNumber="351">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFD4B20430EFFEEFCCEF84D4C55F839" author="KLEMBARA J." box="[882,1049,2001,2027]" pageId="10" pageNumber="351" pagination="854 - 866" refId="ref25357" refString="KLEMBARA J. 2012. - A new species of Pseudopus (Squamata, Anguidae) from the early Miocene of North-West Bohemia (Czech Republic). Journal of Vertebrate Paleontology 32: 854 - 866. https: // doi. org / 10.1080 / 02724634.2012.670177" type="journal article" year="2012">Klembara 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFD4B20430EFFEEFB9BF84D4C82F83E" author="KLEMBARA J." box="[1063,1230,2001,2028]" pageId="10" pageNumber="351" pagination="171 - 195" refId="ref25405" refString="KLEMBARA J. 2015. - New finds of anguines (Squamata, Anguidae) from the Early Miocene of North-West Bohemia (Czech Republic). Palaontologische Zeitschrift 89: 171 - 195. https: // doi. org / 10.1007 / s 12542 - 014 - 0226 - 4" type="journal article" year="2015">Klembara 2015</bibRefCitation>
|
||
–
|
||
<taxonomicName id="4C6C4D52430EFFEEFB56F84E4D00F839" box="[1258,1356,2002,2027]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="rugosus">
|
||
<emphasis id="B918EAC3430EFFEEFB56F84E4D00F839" box="[1258,1356,2002,2027]" italics="true" pageId="10" pageNumber="351">P. rugosus</emphasis>
|
||
</taxonomicName>
|
||
was later renamed to
|
||
<taxonomicName id="4C6C4D52430FFFEFFEB9FF44493BFF23" box="[261,375,215,241]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="confertus">
|
||
<emphasis id="B918EAC3430FFFEFFEB9FF44493BFF23" box="[261,375,215,241]" italics="true" pageId="11" pageNumber="352">P. confertus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFFD4B20430FFFEFFE22FF4B4A8BFF23" author="KLEMBARA J. & RUMMEL M." box="[414,711,215,241]" pageId="11" pageNumber="352" pagination="20 - 44" refId="ref25457" refString="KLEMBARA J. & RUMMEL M. 2018. - New material of Ophisaurus, Anguis and Pseudopus (Squamata, Anguidae, Anguinae) from the Miocene of the Czech Republic and Germany and systematic revision and palaeobiogeography of the Cenozoic Anguinae. Geological Magazine 155 (1): 20 - 44. https: // doi. org / 10.1017 / S 0016756816000753" type="journal article" year="2018">Klembara & Rummel 2018</bibRefCitation>
|
||
). The parietal Pal. 1574 clearly exhibits features present in
|
||
<taxonomicName id="4C6C4D52430FFFEFFD12FF6448F9FEE3" baseAuthorityName="Lartet" baseAuthorityYear="1851" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="laurillardi">
|
||
<emphasis id="B918EAC3430FFFEFFD12FF6448F9FEE3" italics="true" pageId="11" pageNumber="352">P. laurillardi</emphasis>
|
||
</taxonomicName>
|
||
: 1) the presence of free ventrolateral surface (facies ventrolateralis
|
||
<emphasis id="B918EAC3430FFFEFFF40FEA4497CFE83" box="[252,304,312,337]" italics="true" pageId="11" pageNumber="352">sensu</emphasis>
|
||
<bibRefCitation id="EFFD4B20430FFFEFFE8AFEAB4A5BFE83" author="KLEMBARA J. & BOHME M. & RUMMEL M." box="[310,535,311,337]" pageId="11" pageNumber="352" pagination="159 - 196" refId="ref25523" refString="KLEMBARA J., BOHME M. & RUMMEL M. 2010. - Revision of the anguine lizard Pseudopus laurillardi (Squamata, Anguidae) from the Miocene of Europe, with comments on paleoecology. Journal of Paleontology 84 (2): 159 - 196. https: // doi. org / 10.1666 / 09 - 033 R 1.1" type="journal article" year="2010">
|
||
Klembara
|
||
<emphasis id="B918EAC3430FFFEFFE1AFEA44995FE83" box="[422,473,311,337]" italics="true" pageId="11" pageNumber="352">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
) of the supratemporal process, located lateral to the ventrolateral ridge (this feature is absent in
|
||
<taxonomicName id="4C6C4D52430FFFEFFEB8FEEB49E9FE43" authorityName="Klembara" authorityYear="2012" box="[260,421,375,401]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="ahnikoviensis">
|
||
<emphasis id="B918EAC3430FFFEFFEB8FEEB49E9FE43" box="[260,421,375,401]" italics="true" pageId="11" pageNumber="352">P. ahnikoviensis</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="EFFD4B20430FFFEFFE65FEEB4ACDFE43" author="KLEMBARA J." box="[473,641,375,401]" pageId="11" pageNumber="352" pagination="854 - 866" refId="ref25357" refString="KLEMBARA J. 2012. - A new species of Pseudopus (Squamata, Anguidae) from the early Miocene of North-West Bohemia (Czech Republic). Journal of Vertebrate Paleontology 32: 854 - 866. https: // doi. org / 10.1080 / 02724634.2012.670177" type="journal article" year="2012">Klembara 2012</bibRefCitation>
|
||
); and 2) the
|
||
<typeStatus id="54D78873430FFFEFFF38FE0B48FEFE63" box="[132,178,407,433]" pageId="11" pageNumber="352">type</typeStatus>
|
||
of sculpture present on osteodermal shields fused to the bone, consisting of pits, small ridges and grooves rather than being vermicular as it is in
|
||
<taxonomicName id="4C6C4D52430FFFEFFE31FE4B4A6BFE22" authorityName="Klembara" authorityYear="2012" box="[397,551,470,496]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="ahnikoviensis">
|
||
<emphasis id="B918EAC3430FFFEFFE31FE4B4A6BFE22" box="[397,551,470,496]" italics="true" pageId="11" pageNumber="352">P. ahnikoviensis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFFD4B20430FFFEFFDE9FE4A4AB9FE22" author="KLEMBARA J." box="[597,757,470,496]" pageId="11" pageNumber="352" pagination="854 - 866" refId="ref25357" refString="KLEMBARA J. 2012. - A new species of Pseudopus (Squamata, Anguidae) from the early Miocene of North-West Bohemia (Czech Republic). Journal of Vertebrate Paleontology 32: 854 - 866. https: // doi. org / 10.1080 / 02724634.2012.670177" type="journal article" year="2012">Klembara 2012</bibRefCitation>
|
||
). Another feature is the absence of the parietal foramen in the preserved portion of the parietal – in
|
||
<taxonomicName id="4C6C4D52430FFFEFFDB8FD8B4A38FDE2" box="[516,628,534,560]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="confertus">
|
||
<emphasis id="B918EAC3430FFFEFFDB8FD8B4A38FDE2" box="[516,628,534,560]" italics="true" pageId="11" pageNumber="352">P.confertus</emphasis>
|
||
</taxonomicName>
|
||
, this foramen is located closer to the posterior margin of the floor of the parietal fossa (seeKlembara 2015). We can therefore estimate that the parietal foramen in Pal. 1574 was located more anteriorly than it is in
|
||
<taxonomicName id="4C6C4D52430FFFEFFEFAFD0A49F4FD62" box="[326,440,662,688]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="confertus">
|
||
<emphasis id="B918EAC3430FFFEFFEFAFD0A49F4FD62" box="[326,440,662,688]" italics="true" pageId="11" pageNumber="352">P. confertus</emphasis>
|
||
</taxonomicName>
|
||
. The second parietal, specimen Pal. 1402, is problematic. The free ventrolateral surface of the supratemporal process appears to be absent. However, if size is considered, this specimen represents a small individual. In the small parietal described by
|
||
<bibRefCitation id="EFFD4B20430FFFEFFE74FC894AE3FCFD" author="KLEMBARA J. & BOHME M. & RUMMEL M." box="[456,687,789,815]" pageId="11" pageNumber="352" pagination="159 - 196" refId="ref25523" refString="KLEMBARA J., BOHME M. & RUMMEL M. 2010. - Revision of the anguine lizard Pseudopus laurillardi (Squamata, Anguidae) from the Miocene of Europe, with comments on paleoecology. Journal of Paleontology 84 (2): 159 - 196. https: // doi. org / 10.1666 / 09 - 033 R 1.1" type="journal article" year="2010">
|
||
Klembara
|
||
<emphasis id="B918EAC3430FFFEFFD8BFC8A4A25FCFD" box="[567,617,789,815]" italics="true" pageId="11" pageNumber="352">et al.</emphasis>
|
||
(2010
|
||
</bibRefCitation>
|
||
: p. 166, fig. 5B), this feature is only incipiently developed and thus can be influenced by ontogeny. If our allocation is correct, then the parietal Pal. 1402 represents a juvenile individual of
|
||
<taxonomicName id="4C6C4D52430FFFEFFF3FFC0A494DFC7D" baseAuthorityName="Lartet" baseAuthorityYear="1851" box="[131,257,917,943]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="laurillardi">
|
||
<emphasis id="B918EAC3430FFFEFFF3FFC0A494DFC7D" box="[131,257,917,943]" italics="true" pageId="11" pageNumber="352">P.laurillardi</emphasis>
|
||
</taxonomicName>
|
||
(this can be supported by the presence of a similar
|
||
<typeStatus id="54D78873430FFFEFFF38FC2948FEFC1D" box="[132,178,949,975]" pageId="11" pageNumber="352">type</typeStatus>
|
||
of ornamentation to that in Pal. 1574).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |