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<document id="F9B02E854EE09A8765B62B0713E08A2A" ID-CLB-Dataset="8562" ID-DOI="10.5252/geodiversitas2020v42a20" ID-GBIF-Dataset="ad4dbfb6-14ce-48e2-8fe9-99fa26a9bcf4" ID-ISSN="1638-9395" ID-Zenodo-Dep="4447563" ID-ZooBank="urn:lsid:zoobank.org:pub:8FF2A078-CE45-4BF1-A681-00136F57375E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1610989940068" checkinUser="felipe" docAuthor="Ivanov, Martin, Čerňanský, Andrej, Bonilla-Salomón, Isaac &amp; Luján, Àngel Hernández" docDate="2020" docId="03C587C74309FFEBFBAEFB694906FA1F" docLanguage="en" docName="Geodiversitas.42.20.343-376.pdf" docOrigin="Geodiversitas 42 (20)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Anguinae Gray 1825" docType="treatment" docVersion="9" lastPageNumber="356" masterDocId="FFFCFFBF4304FFE4FFBCFF9C484CFFD2" masterDocTitle="Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications" masterLastPageNumber="376" masterPageNumber="343" pageNumber="354" updateTime="1698925529958" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="D57651E02EB5518722312E8F7FD6681D">Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications</mods:title>
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<mods:namePart id="7F7803AE4721A35C7A3E61DC739A7BBE">Ivanov, Martin</mods:namePart>
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<mods:namePart id="B074F0EB702C071F92360EC4EA7D48D4">Čerňanský, Andrej</mods:namePart>
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<mods:namePart id="A7AFFD2F5ABA0062944BB7AC324EA27F">Bonilla-Salomón, Isaac</mods:namePart>
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<mods:namePart id="2A39713229E3467C1A706C2B8059B59C">Luján, Àngel Hernández</mods:namePart>
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<treatment id="03C587C74309FFEBFBAEFB694906FA1F" ID-DOI="http://doi.org/10.5281/zenodo.4488221" ID-GBIF-Taxon="177200744" ID-Zenodo-Dep="4488221" LSID="urn:lsid:plazi:treatment:03C587C74309FFEBFBAEFB694906FA1F" httpUri="http://treatment.plazi.org/id/03C587C74309FFEBFBAEFB694906FA1F" lastPageId="15" lastPageNumber="356" pageId="13" pageNumber="354">
<subSubSection id="C376655A4309FFE9FBAEFB694C87FADF" box="[1042,1227,1267,1295]" pageId="13" pageNumber="354" type="nomenclature">
<paragraph id="8BD336D14309FFE9FBAEFB694C87FADF" blockId="13.[1042,1227,1267,1295]" box="[1042,1227,1267,1295]" pageId="13" pageNumber="354">
<heading id="D09B81BD4309FFE9FBAEFB694C87FADF" box="[1042,1227,1267,1295]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="354" reason="7">
<taxonomicName id="4C6C4D524309FFE9FBAEFB694CC8FADE" authorityName="Gray" authorityYear="1825" box="[1042,1156,1269,1295]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="subFamily" subFamily="Anguinae">ANGUINAE</taxonomicName>
indet.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376655A4309FFE9FB8CFA884CE7FAFC" box="[1072,1195,1299,1326]" pageId="13" pageNumber="354" type="description">
<paragraph id="8BD336D14309FFE9FB8CFA884CE7FAFC" blockId="13.[1072,1195,1299,1326]" box="[1072,1195,1299,1326]" pageId="13" pageNumber="354">
(
<figureCitation id="13572A544309FFE9FB85FA8F4C34FAFF" box="[1081,1144,1299,1325]" captionStart="FIG" captionStartId="14.[132,143,1644,1662]" captionTargetBox="[160,1427,215,1603]" captionTargetId="figure-127@14.[160,1427,214,1603]" captionTargetPageId="14" captionText="FIG. 6. — Ophisaurus and Anguinae indet. from the early Miocene (MN 4) of MWQ. Trunk vertebra Pal. 1407 (1/2001 Turtle Joint) of Ophisaurus sp. in (A) lateral, (B) dorsal, (C) ventral, (D) anterior and (E) posterior aspects. Anguinae indet.: right coronoid Pal. 1408 (1/2001 Turtle Joint) in (F) lateral, (G) medial and (H) dorsal aspects. Osteoderms Pal. 1577-1579 (2/2003 Reptile Joint) in (I-K) external aspects. Abbreviations: alp, anterolateral process; amp, anteromedial process; cd, condyle; ct, cotyle; dp, dorsal process; mr, medial ridge; muc, muscular crest; nc, neural canal; ns, neural spine; nt, notch; os, overlap surface; pof, postzygapophyseal articular facet; pp, posterior process; prf, prezygapophyseal articular facet; rd, ridge; scs, scupltured surface; syn, synapophysis. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447575" httpUri="https://zenodo.org/record/4447575/files/figure.png" pageId="13" pageNumber="354">Fig. 6</figureCitation>
F-K)
</paragraph>
</subSubSection>
<subSubSection id="C376655A4309FFE9FC91FAD14C36FA66" pageId="13" pageNumber="354" type="materials_examined">
<paragraph id="8BD336D14309FFE9FC91FAD14C36FA66" blockId="13.[811,1458,1356,1460]" pageId="13" pageNumber="354">
MATERIAL. —
<materialsCitation id="3B043C8C4309FFE9FC7BFAD14C36FA66" ID-GBIF-Occurrence="3027945305" pageId="13" pageNumber="354">
MWQ, early Miocene, Burdigalian, Orleanian,
<collectionCode id="ED7DAE144309FFE9FC93FAFB4B16FAAC" box="[815,858,1383,1406]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="13" pageNumber="354">MN</collectionCode>
4: 1/2001 Turtle Joint: one right and one left coronoid (Pal. 1408-1409), 38 osteoderms (Pal. 1410-1447); 2/2003 Reptile Joint: 374 osteoderms (Pal. 1577-1950).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376655A4309FFEBFC91FA4F4906FA1F" lastPageId="15" lastPageNumber="356" pageId="13" pageNumber="354" type="description">
<paragraph id="8BD336D14309FFE9FC91FA4F4BF1FA39" blockId="13.[811,1457,1491,2028]" box="[813,957,1491,1517]" pageId="13" pageNumber="354">DESCRIPTION</paragraph>
<paragraph id="8BD336D14309FFE9FC91FA6F4BC2F9DF" blockId="13.[811,1457,1491,2028]" box="[813,910,1523,1549]" pageId="13" pageNumber="354">
<emphasis id="B918EAC34309FFE9FC91FA6F4BC2F9DF" box="[813,910,1523,1549]" italics="true" pageId="13" pageNumber="354">Coronoid</emphasis>
</paragraph>
<paragraph id="8BD336D14309FFE9FC91F98E4DE3F83E" blockId="13.[811,1457,1491,2028]" pageId="13" pageNumber="354">Both left and right coronoids are preserved, but the right one is in better condition. The element is a chevron shaped bone, with several major processes. The dorsal process, which is located in the mid-region, is shortest. However, it is a robust structure, slightly curved posteriorly. On the lateral side, the central region bears a ridge (or keel) for muscle attachment. The posterior process is broad. On its medial side, a strong muscular crest forms the anterior border of the adductor (mandibular) fossa. The whole posterior process is bent posteromedially and, in lateral aspect, the internal side has a wrinkled appearance (it is slightly sculpted by grooves). The anteromedial and anterolateral processes are incomplete. They are separated one from another by a well-developed notch. It shows that the coronoid process of the dentary overlapped this region. A wider base is present on the anteromedial process.</paragraph>
<caption id="DF136659430AFFEAFF38F9F04DFDF90B" ID-DOI="http://doi.org/10.5281/zenodo.4447575" ID-Zenodo-Dep="4447575" httpUri="https://zenodo.org/record/4447575/files/figure.png" pageId="14" pageNumber="355" startId="14.[132,143,1644,1662]" targetBox="[160,1427,215,1603]" targetPageId="14">
<paragraph id="8BD336D1430AFFEAFF38F9F04DFDF90B" blockId="14.[132,1457,1644,1753]" pageId="14" pageNumber="355">
FIG. 6. —
<taxonomicName id="4C6C4D52430AFFEAFF68F9F0497EF9AC" authorityName="Daudin" authorityYear="1803" box="[212,306,1644,1662]" class="Reptilia" family="Anguidae" genus="Ophisaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="355" phylum="Chordata" rank="genus">
<emphasis id="B918EAC3430AFFEAFF68F9F0497EF9AC" box="[212,306,1644,1662]" italics="true" pageId="14" pageNumber="355">Ophisaurus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D52430AFFEAFEE1F9F049E7F9AC" authorityName="Gray" authorityYear="1825" box="[349,427,1644,1662]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="355" phylum="Chordata" rank="subFamily" subFamily="Anguinae">Anguinae</taxonomicName>
indet. from the early Miocene (MN 4) of MWQ. Trunk vertebra Pal. 1407 (1/2001 Turtle Joint) of
<taxonomicName id="4C6C4D52430AFFEAFB79F9F04D6FF9AC" authorityName="Daudin" authorityYear="1803" box="[1221,1315,1644,1662]" class="Reptilia" family="Anguidae" genus="Ophisaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="355" phylum="Chordata" rank="genus">
<emphasis id="B918EAC3430AFFEAFB79F9F04D6FF9AC" box="[1221,1315,1644,1662]" italics="true" pageId="14" pageNumber="355">Ophisaurus</emphasis>
</taxonomicName>
sp. in (
<emphasis id="B918EAC3430AFFEAFADDF9F04D22F9AC" bold="true" box="[1377,1390,1644,1662]" pageId="14" pageNumber="355">A</emphasis>
) lateral, (
<emphasis id="B918EAC3430AFFEAFF35F91F48DAF947" bold="true" box="[137,150,1667,1685]" pageId="14" pageNumber="355">B</emphasis>
) dorsal, (
<emphasis id="B918EAC3430AFFEAFF5EF91F48BCF947" bold="true" box="[226,240,1667,1685]" pageId="14" pageNumber="355">C</emphasis>
) ventral, (
<emphasis id="B918EAC3430AFFEAFEFDF91F4903F947" bold="true" box="[321,335,1667,1685]" pageId="14" pageNumber="355">D</emphasis>
) anterior and (
<emphasis id="B918EAC3430AFFEAFE7AF91F499EF947" bold="true" box="[454,466,1667,1685]" pageId="14" pageNumber="355">E</emphasis>
) posterior aspects.
<taxonomicName id="4C6C4D52430AFFEAFDCBF91F4A89F947" authorityName="Gray" authorityYear="1825" box="[631,709,1667,1685]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="355" phylum="Chordata" rank="subFamily" subFamily="Anguinae">Anguinae</taxonomicName>
indet.: right coronoid Pal. 1408 (1/2001 Turtle Joint) in (
<emphasis id="B918EAC3430AFFEAFB2CF91F4CD7F947" bold="true" box="[1168,1179,1667,1685]" pageId="14" pageNumber="355">F</emphasis>
) lateral, (
<emphasis id="B918EAC3430AFFEAFB5BF91F4CB9F947" bold="true" box="[1255,1269,1667,1685]" pageId="14" pageNumber="355">G</emphasis>
) medial and (
<emphasis id="B918EAC3430AFFEAFAD8F91F4D3EF947" bold="true" box="[1380,1394,1667,1685]" pageId="14" pageNumber="355">H</emphasis>
) dorsal aspects. Osteoderms Pal. 1577-1579 (2/2003 Reptile Joint) in (
<emphasis id="B918EAC3430AFFEAFD2CF9064AE7F97E" bold="true" box="[656,683,1690,1708]" pageId="14" pageNumber="355">I-K</emphasis>
) external aspects. Abbreviations:
<emphasis id="B918EAC3430AFFEAFC79F9064BACF97E" bold="true" box="[965,992,1690,1708]" pageId="14" pageNumber="355">alp</emphasis>
, anterolateral process;
<emphasis id="B918EAC3430AFFEAFB18F9064C87F97E" bold="true" box="[1188,1227,1690,1708]" pageId="14" pageNumber="355">amp</emphasis>
, anteromedial process;
<emphasis id="B918EAC3430AFFEAFA28F9064DE6F97E" bold="true" box="[1428,1450,1690,1708]" pageId="14" pageNumber="355">cd</emphasis>
, condyle;
<emphasis id="B918EAC3430AFFEAFF6CF92C48ADF910" bold="true" box="[208,225,1712,1730]" pageId="14" pageNumber="355">ct</emphasis>
, cotyle;
<emphasis id="B918EAC3430AFFEAFE94F92C4973F910" bold="true" box="[296,319,1712,1730]" pageId="14" pageNumber="355">dp</emphasis>
, dorsal process;
<emphasis id="B918EAC3430AFFEAFE72F92C49AAF910" bold="true" box="[462,486,1712,1730]" pageId="14" pageNumber="355">mr</emphasis>
, medial ridge;
<emphasis id="B918EAC3430AFFEAFDDDF92C4AC4F910" bold="true" box="[609,648,1712,1730]" pageId="14" pageNumber="355">muc</emphasis>
, muscular crest;
<emphasis id="B918EAC3430AFFEAFCABF92C4B61F910" bold="true" box="[791,813,1712,1730]" pageId="14" pageNumber="355">nc</emphasis>
, neural canal;
<emphasis id="B918EAC3430AFFEAFC1AF92C4BF7F910" bold="true" box="[934,955,1712,1730]" pageId="14" pageNumber="355">ns</emphasis>
, neural spine;
<emphasis id="B918EAC3430AFFEAFB89F92C4C0BF910" bold="true" box="[1077,1095,1712,1730]" pageId="14" pageNumber="355">nt</emphasis>
, notch;
<emphasis id="B918EAC3430AFFEAFB37F92C4CECF910" bold="true" box="[1163,1184,1712,1730]" pageId="14" pageNumber="355">os</emphasis>
, overlap surface;
<emphasis id="B918EAC3430AFFEAFA89F92C4D1EF910" bold="true" box="[1333,1362,1712,1730]" pageId="14" pageNumber="355">pof</emphasis>
, postzygapophyseal articular facet;
<emphasis id="B918EAC3430AFFEAFEE7F95B493EF90B" bold="true" box="[347,370,1735,1753]" pageId="14" pageNumber="355">pp</emphasis>
, posterior process;
<emphasis id="B918EAC3430AFFEAFDA4F95B4A7DF90B" bold="true" box="[536,561,1735,1753]" pageId="14" pageNumber="355">prf</emphasis>
, prezygapophyseal articular facet;
<emphasis id="B918EAC3430AFFEAFCE8F95B4B2AF90B" bold="true" box="[852,870,1735,1753]" pageId="14" pageNumber="355">rd</emphasis>
, ridge;
<emphasis id="B918EAC3430AFFEAFC19F95B4B88F90B" bold="true" box="[933,964,1735,1753]" pageId="14" pageNumber="355">scs</emphasis>
, scupltured surface;
<emphasis id="B918EAC3430AFFEAFBCFF95B4CDEF90B" bold="true" box="[1139,1170,1735,1753]" pageId="14" pageNumber="355">syn</emphasis>
, synapophysis. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="8BD336D1430AFFEAFF38F8AD48B5F899" blockId="14.[132,249,1841,1867]" box="[132,249,1841,1867]" pageId="14" pageNumber="355">
<heading id="D09B81BD430AFFEAFF38F8AD48B5F899" box="[132,249,1841,1867]" fontSize="11" level="2" pageId="14" pageNumber="355" reason="8">
<emphasis id="B918EAC3430AFFEAFF38F8AD48B5F899" box="[132,249,1841,1867]" italics="true" pageId="14" pageNumber="355">Osteoderms</emphasis>
</heading>
</paragraph>
<paragraph id="8BD336D1430AFFEAFF38F8CD4B45F839" blockId="14.[132,777,1873,2027]" pageId="14" pageNumber="355">
The osteoderms are of two morphotypes. The first morphotype (
<figureCitation id="13572A54430AFFEAFF7FF8ED4968F85E" box="[195,292,1905,1932]" captionStart="FIG" captionStartId="14.[132,143,1644,1662]" captionTargetBox="[160,1427,215,1603]" captionTargetId="figure-127@14.[160,1427,214,1603]" captionTargetPageId="14" captionText="FIG. 6. — Ophisaurus and Anguinae indet. from the early Miocene (MN 4) of MWQ. Trunk vertebra Pal. 1407 (1/2001 Turtle Joint) of Ophisaurus sp. in (A) lateral, (B) dorsal, (C) ventral, (D) anterior and (E) posterior aspects. Anguinae indet.: right coronoid Pal. 1408 (1/2001 Turtle Joint) in (F) lateral, (G) medial and (H) dorsal aspects. Osteoderms Pal. 1577-1579 (2/2003 Reptile Joint) in (I-K) external aspects. Abbreviations: alp, anterolateral process; amp, anteromedial process; cd, condyle; ct, cotyle; dp, dorsal process; mr, medial ridge; muc, muscular crest; nc, neural canal; ns, neural spine; nt, notch; os, overlap surface; pof, postzygapophyseal articular facet; pp, posterior process; prf, prezygapophyseal articular facet; rd, ridge; scs, scupltured surface; syn, synapophysis. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447575" httpUri="https://zenodo.org/record/4447575/files/figure.png" pageId="14" pageNumber="355">Fig. 6I, J</figureCitation>
) is represented by a slender, flat, trapezoidal osteoderm. This
<typeStatus id="54D78873430AFFEAFE80F80E4926F87E" box="[316,362,1938,1964]" pageId="14" pageNumber="355">type</typeStatus>
possesses a low medial ridge running along its entire central region, including both sculptured region and smooth overlap surface. The ornamentation is
</paragraph>
<paragraph id="8BD336D1430AFFEAFC93F8AD4CFFF839" blockId="14.[811,1457,1841,2027]" pageId="14" pageNumber="355">formed by short ridges, tubercles and pits diverging from the central region. The sculptured region forms approximately 2/3 of the entire length of the osteoderm. The rest is formed by a smooth anterior overlap surface. The lateral bevel is highest close to the overlap surface. The central part of the internal surface is pierced by three foramina.</paragraph>
<caption id="DF136659430BFFEBFF38FBAE4BE3FB5A" ID-DOI="http://doi.org/10.5281/zenodo.4447577" ID-Zenodo-Dep="4447577" httpUri="https://zenodo.org/record/4447577/files/figure.png" pageId="15" pageNumber="356" startId="15.[132,143,1074,1092]" targetBox="[132,1455,215,1033]" targetPageId="15">
<paragraph id="8BD336D1430BFFEBFF38FBAE4BE3FB5A" blockId="15.[132,1457,1074,1160]" pageId="15" pageNumber="356">
FIG. 7. —
<taxonomicName id="4C6C4D52430BFFEBFF6FFBAE4933FB96" authorityName="Szyndlar &amp; Schleich" authorityYear="1993" box="[211,383,1074,1092]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="356" phylum="Chordata" rank="species" species="hermi">
<emphasis id="B918EAC3430BFFEBFF6FFBAE497DFB96" box="[211,305,1074,1092]" italics="true" pageId="15" pageNumber="356">Bavarioboa</emphasis>
cf.
<emphasis id="B918EAC3430BFFEBFEECFBAE4933FB96" box="[336,383,1074,1092]" italics="true" pageId="15" pageNumber="356">hermi</emphasis>
</taxonomicName>
from the early Miocene (MN 4) of MWQ. Posterior trunk vertebra Pal. 1448 (1/2001 Turtle Joint) in left lateral (
<emphasis id="B918EAC3430BFFEBFABEFBAE4D43FB96" bold="true" box="[1282,1295,1074,1092]" pageId="15" pageNumber="356">A</emphasis>
), dorsal (
<emphasis id="B918EAC3430BFFEBFAE7FBAE4D24FB96" bold="true" box="[1371,1384,1074,1092]" pageId="15" pageNumber="356">B</emphasis>
), ventral (
<emphasis id="B918EAC3430BFFEBFF35FBD448DBFB88" bold="true" box="[137,151,1096,1114]" pageId="15" pageNumber="356">C</emphasis>
), cranial (
<emphasis id="B918EAC3430BFFEBFF54FBD548BAFB89" bold="true" box="[232,246,1097,1115]" pageId="15" pageNumber="356">D</emphasis>
) and caudal (
<emphasis id="B918EAC3430BFFEBFEDAFBD5493EFB89" bold="true" box="[358,370,1097,1115]" pageId="15" pageNumber="356">E</emphasis>
) views. Abbreviations:
<emphasis id="B918EAC3430BFFEBFD8FFBD54A08FB89" bold="true" box="[563,580,1097,1115]" pageId="15" pageNumber="356">ct</emphasis>
, cotyle;
<emphasis id="B918EAC3430BFFEBFD30FBD54AEEFB89" bold="true" box="[652,674,1097,1115]" pageId="15" pageNumber="356">hk</emphasis>
, haemal keel;
<emphasis id="B918EAC3430BFFEBFCA6FBD54B6AFB89" bold="true" box="[794,806,1097,1115]" pageId="15" pageNumber="356">ir</emphasis>
, interzygapophyseal ridge;
<emphasis id="B918EAC3430BFFEBFBB1FBD54C6FFB89" bold="true" box="[1037,1059,1097,1115]" pageId="15" pageNumber="356">na</emphasis>
, neural arch;
<emphasis id="B918EAC3430BFFEBFB29FBD54CE7FB89" bold="true" box="[1173,1195,1097,1115]" pageId="15" pageNumber="356">nc</emphasis>
, neural canal;
<emphasis id="B918EAC3430BFFEBFA99FBD54D76FB89" bold="true" box="[1317,1338,1097,1115]" pageId="15" pageNumber="356">ns</emphasis>
, neural spine;
<emphasis id="B918EAC3430BFFEBFF38FBC348D7FBA3" bold="true" box="[132,155,1119,1137]" pageId="15" pageNumber="356">pd</emphasis>
, paradiapophysis;
<emphasis id="B918EAC3430BFFEBFE80FBC34915FBA3" bold="true" box="[316,345,1119,1137]" pageId="15" pageNumber="356">pof</emphasis>
, postzygapophyseal articular facet;
<emphasis id="B918EAC3430BFFEBFD37FBC34AE8FBA3" bold="true" box="[651,676,1119,1137]" pageId="15" pageNumber="356">prf</emphasis>
, prezygapophyseal articular facet;
<emphasis id="B918EAC3430BFFEBFC70FBC34BA6FBA3" bold="true" box="[972,1002,1119,1137]" pageId="15" pageNumber="356">prp</emphasis>
, prezygapophyseal process;
<emphasis id="B918EAC3430BFFEBFB5DFBC34CB0FBA3" bold="true" box="[1249,1276,1119,1137]" pageId="15" pageNumber="356">scf</emphasis>
, subcentral foramen;
<emphasis id="B918EAC3430BFFEBFF38FBEA48E8FB5A" bold="true" box="[132,164,1142,1160]" pageId="15" pageNumber="356">scg</emphasis>
, subcentral groove;
<emphasis id="B918EAC3430BFFEBFEF3FBEA4927FB5A" bold="true" box="[335,363,1142,1160]" pageId="15" pageNumber="356">scr</emphasis>
, subcentral ridge;
<emphasis id="B918EAC3430BFFEBFDBBFBEA4A56FB5A" bold="true" box="[519,538,1142,1160]" pageId="15" pageNumber="356">zy</emphasis>
, zygosphene;
<emphasis id="B918EAC3430BFFEBFD28FBEA4AFFFB5A" bold="true" box="[660,691,1142,1160]" pageId="15" pageNumber="356">zyg</emphasis>
, zygantrum. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="8BD336D1430BFFEBFF27FB484906FA1F" blockId="15.[130,777,1236,1485]" pageId="15" pageNumber="356">
Osteoderms of the second morphotype (
<figureCitation id="13572A54430BFFEBFD85FB484AC9FB3C" box="[569,645,1236,1262]" captionStart="FIG" captionStartId="14.[132,143,1644,1662]" captionTargetBox="[160,1427,215,1603]" captionTargetId="figure-127@14.[160,1427,214,1603]" captionTargetPageId="14" captionText="FIG. 6. — Ophisaurus and Anguinae indet. from the early Miocene (MN 4) of MWQ. Trunk vertebra Pal. 1407 (1/2001 Turtle Joint) of Ophisaurus sp. in (A) lateral, (B) dorsal, (C) ventral, (D) anterior and (E) posterior aspects. Anguinae indet.: right coronoid Pal. 1408 (1/2001 Turtle Joint) in (F) lateral, (G) medial and (H) dorsal aspects. Osteoderms Pal. 1577-1579 (2/2003 Reptile Joint) in (I-K) external aspects. Abbreviations: alp, anterolateral process; amp, anteromedial process; cd, condyle; ct, cotyle; dp, dorsal process; mr, medial ridge; muc, muscular crest; nc, neural canal; ns, neural spine; nt, notch; os, overlap surface; pof, postzygapophyseal articular facet; pp, posterior process; prf, prezygapophyseal articular facet; rd, ridge; scs, scupltured surface; syn, synapophysis. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447575" httpUri="https://zenodo.org/record/4447575/files/figure.png" pageId="15" pageNumber="356">Fig.6K</figureCitation>
) are roughly rectangular in shape. This morphotype almost lacks the medial ridge ornamentation is arranged only in the centre to form an indistinct ridge. This osteoderm
<typeStatus id="54D78873430BFFEBFE42FAA84A60FA9C" box="[510,556,1332,1358]" pageId="15" pageNumber="356">type</typeStatus>
also differs from the first morphotype by its sculpture. It is denser because the ridges are broad, with narrow grooves between them. The smooth overlap surface is large. The internal surface is pierced by three foramina.
</paragraph>
</subSubSection>
</treatment>
</document>