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<document id="9F06AF1730F1722BCB53B41E03F07BE1" ID-DOI="10.5281/zenodo.277308" ID-GBIF-Dataset="61dd45a6-918a-4b12-8157-6090f7514613" ID-ISSN="1175-5326" ID-Zenodo-Dep="277308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460286829391" checkinUser="plazi" docAuthor="Rosa, Simone Policena" docDate="2011" docId="03C58797BE052927FF57FC7EFB05F115" docLanguage="en" docName="zt02831p022.pdf" docOrigin="Zootaxa 2831" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Triplonychus crassifemoris Rosa, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="FFFCFFEFBE00292FFFC0FF89FFB7F724" masterDocTitle="New species of Triplonychus Candèze and Globothorax Fleutiaux from Brazil (Coleoptera, Elateridae, Cardiophorinae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="6" updateTime="1698250676213" updateUser="plazi">
<mods:mods id="5BC747879B1D5F61710D9A067341A077" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C4A510F084E0E54BA330F2053F2E290A">
<mods:title id="240E5DEA9689683E6F7EB28C2D2E5843">New species of Triplonychus Candèze and Globothorax Fleutiaux from Brazil (Coleoptera, Elateridae, Cardiophorinae)</mods:title>
</mods:titleInfo>
<mods:name id="5354AF2DC9F9A39CB31F5536760513A3" type="personal">
<mods:role id="3D8A8D257719FAFBEDA5F8F30F28DA4F">
<mods:roleTerm id="85E15A124B741B7D8FEFEAC8E36F03FA">Author</mods:roleTerm>
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<mods:namePart id="21A9E32DB6189F1DC6AB7F2B1112091F">Rosa, Simone Policena</mods:namePart>
</mods:name>
<mods:typeOfResource id="F206519C665B967F3ABBD795E2B61804">text</mods:typeOfResource>
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<mods:title id="7F2EA090996EF23896548763BD7E6892">Zootaxa</mods:title>
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<mods:part id="D759594951939583B5E9BFA16EFA42CB">
<mods:date id="9E1ADC4E7EFF08481B6A1049B174B7C8">2011</mods:date>
<mods:detail id="748A91BE37BFB188D7121341FE464C3B" type="volume">
<mods:number id="2FB94F5CA99DCE2944834658B836DAFF">2831</mods:number>
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<mods:classification id="17427903844445233AA6DA9F895480FE">journal article</mods:classification>
<mods:identifier id="FAAEA9D029266A786EF27687D1C45D0A" type="DOI">10.5281/zenodo.277308</mods:identifier>
<mods:identifier id="2610DB52402E2CBD2D11DF505F71A90F" type="GBIF-Dataset">61dd45a6-918a-4b12-8157-6090f7514613</mods:identifier>
<mods:identifier id="F7E5AEB1D5F0D37B081DCBD28266525E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B8D4AF7A8B791123CBEC6226A879FE6A" type="Zenodo-Dep">277308</mods:identifier>
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<treatment id="03C58797BE052927FF57FC7EFB05F115" ID-DOI="http://doi.org/10.5281/zenodo.6188948" ID-GBIF-Taxon="119397855" ID-Zenodo-Dep="6188948" LSID="urn:lsid:plazi:treatment:03C58797BE052927FF57FC7EFB05F115" httpUri="http://treatment.plazi.org/id/03C58797BE052927FF57FC7EFB05F115" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
<subSubSection id="C376650ABE05292AFF57FC7EFDBAF316" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BD33681BE05292AFF57FC7EFDF3F335" blockId="5.[151,580,1015,1074]" box="[151,580,1015,1041]" pageId="5" pageNumber="6">
<heading id="D09B81EDBE05292AFF57FC7EFDF3F335" bold="true" box="[151,580,1015,1041]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="B918EA93BE05292AFF57FC7EFDF3F335" bold="true" box="[151,580,1015,1041]" pageId="5" pageNumber="6">
<taxonomicName id="4C6C4D02BE05292AFF57FC7EFE68F335" box="[151,479,1015,1041]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="crassifemoris" status="sp. nov.">
<emphasis id="B918EA93BE05292AFF57FC7EFE68F335" bold="true" box="[151,479,1015,1041]" italics="true" pageId="5" pageNumber="6">Triplonychus crassifemoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE05292AFE27FC7EFDF3F335" box="[487,580,1015,1041]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD33681BE05292AFF57FB93FDBAF316" blockId="5.[151,580,1015,1074]" box="[151,525,1050,1074]" pageId="5" pageNumber="6">
(
<figureCitation id="13572A04BE05292AFF5FFB93FE90F316" box="[159,295,1050,1074]" captionStart-0="FIGURES 15 – 25" captionStart-1="FIGURES 26 – 33" captionStartId-0="6.[151,264,1978,2001]" captionStartId-1="7.[151,264,1948,1971]" captionTargetBox-0="[215,1386,193,1941]" captionTargetBox-1="[205,1361,198,1893]" captionTargetId-0="figure@6.[195,1400,193,1965]" captionTargetId-1="figure@7.[202,1371,193,1911]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." captionText-1="FIGURES 26 – 33. Triplonychus crassifemoris sp. nov. 26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis." httpUri-0="https://zenodo.org/record/277311/files/figure.png" httpUri-1="https://zenodo.org/record/277312/files/figure.png" pageId="5" pageNumber="6">Figs. 15–33</figureCitation>
,
<figureCitation id="13572A04BE05292AFEF2FB93FECCF316" box="[306,379,1050,1074]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="5" pageNumber="6">80–82</figureCitation>
,
<figureCitation id="13572A04BE05292AFE46FB93FE79F316" box="[390,462,1050,1074]" captionStart="FIGURES 91 – 98" captionStartId="17.[151,264,1951,1974]" captionTargetBox="[403,1181,190,1931]" captionTargetId="figure@17.[396,1192,185,1935]" captionTargetPageId="17" captionText="FIGURES 91 – 98. Right elytron: 91, Triplonychus cruspinosus sp. nov.; 92, T. crassifemoris sp. nov.; 93, T. tibialatus sp. nov.; 94, Globothorax latidens sp. nov. Mesoscutum and scutellar shield: 95, Triplonychus cruspinosus sp. nov.; 96, T. crassifemoris sp. nov.; 97, T. tibialatus sp. nov.; 98, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277319/files/figure.png" pageId="5" pageNumber="6">92, 96</figureCitation>
,
<figureCitation id="13572A04BE05292AFE1AFB93FDB2F316" box="[474,517,1050,1074]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="5" pageNumber="6">102</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE05292AFF57FBEAFF58F3BA" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BD33681BE05292AFF57FBEAFF58F3BA" blockId="5.[151,1437,1122,2011]" pageId="5" pageNumber="6">
<emphasis id="B918EA93BE05292AFF57FBEAFEA9F35F" bold="true" box="[151,286,1123,1147]" pageId="5" pageNumber="6">Etymology.</emphasis>
From Latin,
<emphasis id="B918EA93BE05292AFE70FBEDFDB3F35F" box="[432,516,1124,1147]" italics="true" pageId="5" pageNumber="6">crassus</emphasis>
= fat,
<emphasis id="B918EA93BE05292AFD8DFBEAFD16F35F" box="[589,673,1123,1147]" italics="true" pageId="5" pageNumber="6">femoris</emphasis>
= femur, alluding to the widened and convex posterior femur of this species.
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE05292AFF05FB23FAC6F22E" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BD33681BE05292AFF05FB23FAC6F22E" blockId="5.[151,1437,1122,2011]" pageId="5" pageNumber="6">
<emphasis id="B918EA93BE05292AFF05FB23FE88F3E7" bold="true" box="[197,319,1194,1219]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Pronotum wider than long with rounded sides (
<figureCitation id="13572A04BE05292AFC92FB23FC12F3E6" box="[850,933,1194,1218]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 21</figureCitation>
); antennae surpassing the hind angles by one antennomere; mandibles large (
<figureCitation id="13572A04BE05292AFE39FB47FDFCF3C2" box="[505,587,1230,1254]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 17</figureCitation>
,
<figureCitation id="13572A04BE05292AFD98FB47FD31F3C2" box="[600,646,1230,1254]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="5" pageNumber="6">102</figureCitation>
); metacoxal plate triangular with posterior margin acute (
<figureCitation id="13572A04BE05292AFACCFB47FAD7F3C2" box="[1292,1376,1230,1254]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 25</figureCitation>
); tibiae widened apicad, metafemur and metatrochanter strongly widened dorsoventrally and convex (
<figureCitation id="13572A04BE05292AFB1BFB7BFAD2F22E" box="[1243,1381,1266,1290]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="5" pageNumber="6">Figs. 80–82</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE052927FF05FA9FFDBAF35B" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="description">
<paragraph id="8BD33681BE05292AFF05FA9FFD5DF29A" blockId="5.[151,1437,1122,2011]" pageId="5" pageNumber="6">
<emphasis id="B918EA93BE05292AFF05FA9FFEE0F20B" bold="true" box="[197,343,1302,1327]" pageId="5" pageNumber="6">Description.</emphasis>
Male (
<figureCitation id="13572A04BE05292AFE6AFA9FFDB9F20A" box="[426,526,1302,1326]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="5" pageNumber="6">Fig. 102</figureCitation>
). Body convex, densely covered with fine and short setae; setae yellow to light brown and semidecumbent to erect on dorsal surface, yellow and decumbent on ventral surface. Integument bright; evenly brown or with dorsal surface light brown to brown and ventral surface and legs lighter. Pronotal and elytral punctures not or slightly darker than the surrounding area. Total length: 6.0–7.0 mm; elytra 2.5–2.7 times longer than prothorax; elytral base 1.1 as wide as prothorax.
</paragraph>
<paragraph id="8BD33681BE05292AFF05FA43FB16F04A" blockId="5.[151,1437,1122,2011]" pageId="5" pageNumber="6">
Head (
<figureCitation id="13572A04BE05292AFECEFA43FED5F2C6" box="[270,354,1482,1506]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 20</figureCitation>
). Frons shallowly concave; anterior margin carinate, rounded and produced; frontoclypeal region steeply declivous to base of labrum and 2 times wider than long. Punctures smaller than those of pronotal disk, umbilicate 0.5–1.0 diameter apart. Antenna (
<figureCitation id="13572A04BE05292AFD54F99BFD50F10E" box="[660,743,1554,1578]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 15</figureCitation>
) surpassing the hind angles of pronotum at antennomere 10; antennomere 3 1.7 times longer than 2, subequal to the 4. Dorsal index of eye prominence 0.4–0.5. Labrum (
<figureCitation id="13572A04BE05292AFAAFF9BFFF03F156" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 16</figureCitation>
) semielliptical, strongly punctate, with long setae, anterior margin rounded or emarginated. Mandible (
<figureCitation id="13572A04BE05292AFA81F9D3FA23F156" box="[1345,1428,1626,1650]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 17</figureCitation>
) long (distal half superimpose each other) unidentate, evenly arcuate, with rows of small setae on anteromesal margin, a longitudinal short and narrow carina dorsally and a stout and longer seta than the fine setae of the lateroventral margin. Maxilla with basistipe fused to medistipe (
<figureCitation id="13572A04BE05292AFCC9F94FFCEBF1FA" box="[777,860,1734,1758]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 18</figureCitation>
), with a long stout seta and a few fine short setae; galea triangular with outer angle strongly produced and sparsely covered with long stout setae, inner angle with a short rectangular lobe densely covered with fine short setae. Labium (
<figureCitation id="13572A04BE05292AFC61F887FC45F002" box="[929,1010,1806,1830]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Fig. 19</figureCitation>
) with mentum with one seta on posterior angles; prementum notched at anteromedian margin, lateroanterior angles rounded and produced; second labial palpomere with spiniform setae apically; apical maxillary and labial palpomeres securiform.
</paragraph>
<paragraph id="8BD33681BE052927FF05F8F3FAC6F5EB" blockId="5.[151,1437,1122,2011]" lastBlockId="8.[151,1437,151,1585]" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
Thorax. Prothorax (
<figureCitation id="13572A04BE05292AFE62F8F3FD90F0B6" box="[418,551,1914,1938]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="5" pageNumber="6">Figs. 21–23</figureCitation>
) about 1.2 times wider than long, lateral margins rounded, carinate on the posterior 3/4. Pronotum convex; punctures on disk large 0.5–1.5 diameters apart, on anterior margin smaller 0.0–0.5 diameter apart, on posterior margin very small 0.5–1.0 diameter apart. Pronotal hind angles flat, short, obliquely truncate; posterior margin between hind angle and median line with a very small notch. Hypomera (
<figureCitation id="13572A04BE082927FAC7FF1EFAEDF78B" box="[1287,1370,151,175]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="8" pageNumber="9">Fig. 23</figureCitation>
) with punctation double; notosternal suture nearly straight. Prosternum (
<figureCitation id="13572A04BE082927FC55FF35FC5DF7F0" box="[917,1002,188,212]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="8" pageNumber="9">Fig. 23</figureCitation>
) 1.0–1.1 times wider than long, with punctures smaller than those of pronotal disk, umbilicate, 1–2 diameters apart. Prosternal process (
<figureCitation id="13572A04BE082927FAD2FF56FAD1F7D3" box="[1298,1382,223,247]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="8" pageNumber="9">Fig. 23</figureCitation>
) 2.5 times longer than coxal diameter with ventral surface of the posterior half flattened laterally, gradually declivous posteriorly to coxae then steeply sloping dorsad at apex (
<figureCitation id="13572A04BE082927FCE2FEAEFCCFF61B" box="[802,888,295,319]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="8" pageNumber="9">Fig. 22</figureCitation>
). Procoxal cavities closed. Scutellar shield (
<figureCitation id="13572A04BE082927FAAEFEAEFF02F640" captionStart="FIGURES 91 – 98" captionStartId="17.[151,264,1951,1974]" captionTargetBox="[403,1181,190,1931]" captionTargetId="figure@17.[396,1192,185,1935]" captionTargetPageId="17" captionText="FIGURES 91 – 98. Right elytron: 91, Triplonychus cruspinosus sp. nov.; 92, T. crassifemoris sp. nov.; 93, T. tibialatus sp. nov.; 94, Globothorax latidens sp. nov. Mesoscutum and scutellar shield: 95, Triplonychus cruspinosus sp. nov.; 96, T. crassifemoris sp. nov.; 97, T. tibialatus sp. nov.; 98, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277319/files/figure.png" pageId="8" pageNumber="9">Fig. 96</figureCitation>
) pentagonal to cordiform with a short median notch on anterior margin, abruptly elevated above the level of mesoscutum. Mesoventrite (
<figureCitation id="13572A04BE082927FE18FEFBFD9BF6AE" box="[472,556,370,394]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="8" pageNumber="9">Fig. 24</figureCitation>
) with posterior region inclined 45o above the level of anterior region; lateral lobes narrow and elongate; mesoventral cavity widened up to anterior border of mesocoxae than abruptly convergent. Mesocoxal cavity oval, closed. Mesepisternum with a weak transverse carina near the anterior inner angle; anterior inner angle rounded. Meso-metaventral suture impressed. Metaventrite (
<figureCitation id="13572A04BE082927FBFBFE56FB7EF6D3" box="[1083,1225,479,503]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="8" pageNumber="9">Figs. 24, 25</figureCitation>
) with punctuation double; metacoxal plate triangular with posterior margin acute. Elytra (
<figureCitation id="13572A04BE082927FC00FD8BFBA4F53E" box="[960,1043,514,538]" captionStart="FIGURES 91 – 98" captionStartId="17.[151,264,1951,1974]" captionTargetBox="[403,1181,190,1931]" captionTargetId="figure@17.[396,1192,185,1935]" captionTargetPageId="17" captionText="FIGURES 91 – 98. Right elytron: 91, Triplonychus cruspinosus sp. nov.; 92, T. crassifemoris sp. nov.; 93, T. tibialatus sp. nov.; 94, Globothorax latidens sp. nov. Mesoscutum and scutellar shield: 95, Triplonychus cruspinosus sp. nov.; 96, T. crassifemoris sp. nov.; 97, T. tibialatus sp. nov.; 98, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277319/files/figure.png" pageId="8" pageNumber="9">Fig. 92</figureCitation>
) slightly rounded or parallel-sided up to posterior 1/3 then tapering to apex; elytral striae with 2 rows of punctures predominantly paired; interstices 3, 5 and 7 strongly convex on apical region; convex on anterior region; interstice 9 carinate on posterior 2/3. Legs (
<figureCitation id="13572A04BE082927FF5FFDE6FE90F5A3" box="[159,295,623,647]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="8" pageNumber="9">Figs. 80–82</figureCitation>
). Pro- and mesofemur slightly widened dorsoventrally, metafemur and metatrochanter strongly widened dorsoventrally and convex. Tibiae widened apicad with rows of long spiniform setae on dorsal and apical margin, tibial spurs stout about 0.5–0.7 times as long as tarsomere 1; tarsomeres densely pilose with fine setae.
</paragraph>
<caption id="DF136609BE062929FF57F833FB81F0CB" httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="6" pageNumber="7" targetBox="[215,1386,193,1941]" targetPageId="6">
<paragraph id="8BD33681BE062929FF57F833FB81F0CB" blockId="6.[151,1436,1978,2031]" pageId="6" pageNumber="7">
<emphasis id="B918EA93BE062929FF57F833FEE1F0F4" bold="true" box="[151,342,1978,2001]" pageId="6" pageNumber="7">FIGURES 15–25.</emphasis>
<taxonomicName id="4C6C4D02BE062929FE9FF833FDC3F0F4" box="[351,628,1978,2000]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="crassifemoris" status="sp. nov.">
<emphasis id="B918EA93BE062929FE9FF833FDC3F0F4" box="[351,628,1978,2000]" italics="true" pageId="6" pageNumber="7">Triplonychus crassifemoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE062929FDBDF832FD67F0F4" box="[637,720,1979,2000]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EA93BE062929FDBDF832FD67F0F4" bold="true" box="[637,720,1979,2000]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21–23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral).
</paragraph>
</caption>
<caption id="DF136609BE072928FF57F815FC9BF0D4" httpUri="https://zenodo.org/record/277312/files/figure.png" pageId="7" pageNumber="8" targetBox="[205,1361,198,1893]" targetPageId="7">
<paragraph id="8BD33681BE072928FF57F815FC9BF0D4" blockId="7.[151,1437,1948,2032]" pageId="7" pageNumber="8">
<emphasis id="B918EA93BE072928FF57F815FEE3F095" bold="true" box="[151,340,1948,1971]" pageId="7" pageNumber="8">FIGURES 26–33.</emphasis>
<taxonomicName id="4C6C4D02BE072928FE9AF815FDDBF096" box="[346,620,1948,1970]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassifemoris" status="sp. nov.">
<emphasis id="B918EA93BE072928FE9AF815FDDBF096" box="[346,620,1948,1970]" italics="true" pageId="7" pageNumber="8">Triplonychus crassifemoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE072928FDB3F814FD73F096" box="[627,708,1949,1970]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B918EA93BE072928FDB3F814FD73F096" bold="true" box="[627,708,1949,1970]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis.
</paragraph>
</caption>
<paragraph id="8BD33681BE082927FF05FD53FD1AF4CB" blockId="8.[151,1437,151,1585]" pageId="8" pageNumber="9">
Abdomen (
<figureCitation id="13572A04BE082927FE82FD53FE24F5D6" box="[322,403,730,754]" captionStart="FIGURES 26 – 33" captionStartId="7.[151,264,1948,1971]" captionTargetBox="[205,1361,198,1893]" captionTargetId="figure@7.[202,1371,193,1911]" captionTargetPageId="7" captionText="FIGURES 26 – 33. Triplonychus crassifemoris sp. nov. 26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277312/files/figure.png" pageId="8" pageNumber="9">Fig. 27</figureCitation>
) with sides rounded, punctation double denser on lateral margin. First ventrite 3.4 times longer on lateral border than at middle, hind angles of ventrites 1–4 acute, last ventrite pentagonal. Tergite 8 (
<figureCitation id="13572A04BE082927FA82FD76FA24F433" box="[1346,1427,767,791]" captionStart="FIGURES 26 – 33" captionStartId="7.[151,264,1948,1971]" captionTargetBox="[205,1361,198,1893]" captionTargetId="figure@7.[202,1371,193,1911]" captionTargetPageId="7" captionText="FIGURES 26 – 33. Triplonychus crassifemoris sp. nov. 26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277312/files/figure.png" pageId="8" pageNumber="9">Fig. 28</figureCitation>
) evenly sclerotized, densely covered with microsetae, with a few long and stout setae on lateroposterior and posterior margins; anterior sclerotized border U-shaped emarginated. Sternite 8 (
<figureCitation id="13572A04BE082927FC21FCCEFB85F47B" box="[993,1074,839,863]" captionStart="FIGURES 26 – 33" captionStartId="7.[151,264,1948,1971]" captionTargetBox="[205,1361,198,1893]" captionTargetId="figure@7.[202,1371,193,1911]" captionTargetPageId="7" captionText="FIGURES 26 – 33. Triplonychus crassifemoris sp. nov. 26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277312/files/figure.png" pageId="8" pageNumber="9">Fig. 29</figureCitation>
) pentagonal, evenly sclerotized, with short setae lateroposteriorly, apex wide emarginate at middle. Sternite 9 (
<figureCitation id="13572A04BE082927FBE5FCE3FBD8F4A6" box="[1061,1135,874,898]" captionStart="FIGURES 26 – 33" captionStartId="7.[151,264,1948,1971]" captionTargetBox="[205,1361,198,1893]" captionTargetId="figure@7.[202,1371,193,1911]" captionTargetPageId="7" captionText="FIGURES 26 – 33. Triplonychus crassifemoris sp. nov. 26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277312/files/figure.png" pageId="8" pageNumber="9">Fig.31</figureCitation>
) with sides subparallel or divergent posteriorly, abruptly convergent at apex; lateral and posterior margins sclerotized, anterior and median region translucent, apex with short and elongate setae. Tergite 9 (
<figureCitation id="13572A04BE082927FCAFFC3BFC76F4EE" box="[879,961,946,970]" captionStart="FIGURES 26 – 33" captionStartId="7.[151,264,1948,1971]" captionTargetBox="[205,1361,198,1893]" captionTargetId="figure@7.[202,1371,193,1911]" captionTargetPageId="7" captionText="FIGURES 26 – 33. Triplonychus crassifemoris sp. nov. 26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277312/files/figure.png" pageId="8" pageNumber="9">Fig. 30</figureCitation>
) with anterior margin nearly straight, posterior lobes acute; tergite 10 (
<figureCitation id="13572A04BE082927FE20FC5EFD86F4CB" box="[480,561,983,1007]" captionStart="FIGURES 26 – 33" captionStartId="7.[151,264,1948,1971]" captionTargetBox="[205,1361,198,1893]" captionTargetId="figure@7.[202,1371,193,1911]" captionTargetPageId="7" captionText="FIGURES 26 – 33. Triplonychus crassifemoris sp. nov. 26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277312/files/figure.png" pageId="8" pageNumber="9">Fig. 30</figureCitation>
) semioval.
</paragraph>
<paragraph id="8BD33681BE082927FF05FC73FDBAF35B" blockId="8.[151,1437,151,1585]" pageId="8" pageNumber="9">
Aedeagus (
<figureCitation id="13572A04BE082927FE83FC73FE7AF336" box="[323,461,1018,1042]" captionStart="FIGURES 26 – 33" captionStartId="7.[151,264,1948,1971]" captionTargetBox="[205,1361,198,1893]" captionTargetId="figure@7.[202,1371,193,1911]" captionTargetPageId="7" captionText="FIGURES 26 – 33. Triplonychus crassifemoris sp. nov. 26, metanotum; 27, abdomen; 28, tergite 8; 29, sternite 8; 30, tergites 9 and 10; 31, sternite 9; 32, 33, aedeagus (ventral, dorsal). Abbreviations: pn, postnotum; psc, prescutum; mma, median membranous area; sc, scutum; scl, scutellum; vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277312/files/figure.png" pageId="8" pageNumber="9">Figs. 32, 33</figureCitation>
). Parameres with apex narrow, rounded and slightly convergent with one or two setae, articulated to penis by transparent membrane. Penis with a ventral sclerite elliptical about 6 times longer than wide; basal struts about 0.5 times as long as its total length, median region between basal struts without process, lateral margins tapered to rounded apex.
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE082927FF05FB02FB72F235" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BD33681BE082927FF05FB02FE32F3E3" blockId="8.[151,1437,151,1585]" pageId="8" pageNumber="9">
<emphasis id="B918EA93BE082927FF05FB02FEC1F387" bold="true" box="[197,374,1162,1187]" pageId="8" pageNumber="9">
<typeStatus id="54D78823BE082927FF05FB02FEB7F387" box="[197,256,1163,1187]" pageId="8" pageNumber="9">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54D78823BE082927FE40FB02FE50F387" box="[384,487,1163,1187]" pageId="8" pageNumber="9" type="holotype">
<emphasis id="B918EA93BE082927FE40FB02FE50F387" box="[384,487,1163,1187]" italics="true" pageId="8" pageNumber="9">Holotype</emphasis>
</typeStatus>
. [Igarapé Gurupi-Una, Aldeia Araçu, MA, 50 Km E de Canindé,
<date id="FFD21041BE082927FB28FB03FA8BF386" box="[1256,1340,1162,1186]" pageId="8" pageNumber="9" value="1966-02">II.1966</date>
, Malkin col.] male (
<collectionCode id="ED7DAE44BE082927FED6FB26FECDF3E3" box="[278,378,1199,1223]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
).
</paragraph>
<paragraph id="8BD33681BE082927FF05FB5DFCE3F3CE" blockId="8.[151,1437,151,1585]" box="[197,852,1234,1259]" pageId="8" pageNumber="9">
<typeStatus id="54D78823BE082927FF05FB5DFE80F3CF" box="[197,311,1236,1259]" pageId="8" pageNumber="9" type="paratype">
<emphasis id="B918EA93BE082927FF05FB5DFE80F3CF" box="[197,311,1236,1259]" italics="true" pageId="8" pageNumber="9">Paratypes</emphasis>
</typeStatus>
. Same label as the
<typeStatus id="54D78823BE082927FDCEFB5BFDC4F3CE" box="[526,627,1234,1258]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D6AFD08BE082927FDBEFB5BFD63F3CE" box="[638,724,1234,1258]" pageId="8" pageNumber="9" type="male">6 males</specimenCount>
(
<collectionCode id="ED7DAE44BE082927FD24FB5BFCFFF3CE" box="[740,840,1234,1258]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
).
</paragraph>
<paragraph id="8BD33681BE082927FF05FB70FB72F235" blockId="8.[151,1437,151,1585]" box="[197,1221,1270,1298]" pageId="8" pageNumber="9">
<emphasis id="B918EA93BE082927FF05FB70FEE7F236" bold="true" box="[197,336,1273,1298]" pageId="8" pageNumber="9">Occurrence</emphasis>
.
<collectingCountry id="F37B7611BE082927FE9EFB73FE70F236" box="[350,455,1274,1298]" name="Brazil" pageId="8" pageNumber="9">BRAZIL</collectingCountry>
. Maranhão state:
<geoCoordinate id="EE585046BE082927FD4CFB70FCAFF235" box="[652,792,1270,1297]" direction="west" orientation="longitude" pageId="8" pageNumber="9" precision="15" value="-46.48">46o28’48”W</geoCoordinate>
,
<geoCoordinate id="EE585046BE082927FCE3FB70FC21F235" box="[803,918,1270,1297]" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="15" value="-2.56">2o33’36”S</geoCoordinate>
(
<bibRefCitation id="EFFD4B70BE082927FC65FB70FB0AF235" author="Rocha" box="[933,1213,1273,1297]" pageId="8" pageNumber="9" refString="Rocha, M. M. &amp; Cancello, E. M. (2007) Estudo taxonomico de Cylindrotermes Holmgren (Isoptera, Termitidae, Termitinae). Papeis Avulsos de Zoologia, 47 (10), 137 - 152." type="journal article" year="2007">Rocha &amp; Cancello, 2007</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE082927FF05FA94FB05F115" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BD33681BE082927FF05FA94FB05F115" blockId="8.[151,1437,151,1585]" pageId="8" pageNumber="9">
<emphasis id="B918EA93BE082927FF05FA94FEFDF211" bold="true" box="[197,330,1309,1334]" pageId="8" pageNumber="9">Discussion.</emphasis>
<taxonomicName id="4C6C4D02BE082927FE93FA94FDC4F212" box="[339,627,1309,1334]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cruspinosus" status="sp. nov.">
<emphasis id="B918EA93BE082927FE93FA94FDC4F212" box="[339,627,1309,1334]" italics="true" pageId="8" pageNumber="9">Triplonychus cruspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE082927FDBCFA97FD6FF212" box="[636,728,1310,1334]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B918EA93BE082927FDBCFA97FD6FF212" bold="true" box="[636,728,1310,1334]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C6C4D02BE082927FCD2FA96FC70F211" box="[786,967,1309,1334]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="crassifemoris" status="sp. nov.">
<emphasis id="B918EA93BE082927FCD2FA96FC70F211" box="[786,967,1309,1334]" italics="true" pageId="8" pageNumber="9">T. crassifemoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE082927FC0FFA97FB9CF212" box="[975,1067,1310,1334]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B918EA93BE082927FC0FFA97FB9CF212" bold="true" box="[975,1067,1310,1334]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
are similar to the species of the second group listed by
<bibRefCitation id="EFFD4B70BE082927FE66FAC8FDEDF27D" author="Schwarz" box="[422,602,1345,1369]" pageId="8" pageNumber="9" refString="Schwarz, O. (1906) Coleoptera. Fam. Elateridae. In P. Wytsman (Ed.) Genera Insectorum, Fasc. 46 B. P. Wytsman, Brussels, pp. 113 - 224." type="book chapter" year="1906">Schwarz (1906)</bibRefCitation>
by their pronotum “not longer than wide” and also defined by
<bibRefCitation id="EFFD4B70BE082927FAFCFAC8FF56F259" author="Candeze" pageId="8" pageNumber="9" refString="Candeze, E. (1860) Monographie des Elaterides. Vol. 3. Memoires de la Societe Royale des Sciences de Liege, 15, 512 pp." type="book" year="1860">Candèze (1860)</bibRefCitation>
as species with prothorax as wide as long. Both species have pronotum a little wider than long.
<taxonomicName id="4C6C4D02BE082927FAD5FAEEFF0CF285" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cruspinosus" status="sp. nov.">
<emphasis id="B918EA93BE082927FAD5FAEEFF0CF285" italics="true" pageId="8" pageNumber="9">T. cruspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE082927FF02FA03FEABF286" box="[194,284,1418,1442]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B918EA93BE082927FF02FA03FEABF286" bold="true" box="[194,284,1418,1442]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C6C4D02BE082927FE2CFA03FCA3F285" authority="Candeze, 1860" authorityName="Candeze" authorityYear="1860" box="[492,788,1417,1441]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B918EA93BE082927FE2CFA03FDD4F285" box="[492,611,1418,1441]" italics="true" pageId="8" pageNumber="9">T. costatus</emphasis>
<bibRefCitation id="EFFD4B70BE082927FDAAFA00FCA3F285" author="Candeze" box="[618,788,1417,1441]" pageId="8" pageNumber="9" refString="Candeze, E. (1860) Monographie des Elaterides. Vol. 3. Memoires de la Societe Royale des Sciences de Liege, 15, 512 pp." type="book" year="1860">Candèze, 1860</bibRefCitation>
</taxonomicName>
, from Pará state, differing from it in its color pattern, longer antennae and elytral interstices 3, 5, and 7 convex, not carinate.
<taxonomicName id="4C6C4D02BE082927FC55FA27FB71F2E2" box="[917,1222,1454,1478]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="crassifemoris" status="sp. nov.">
<emphasis id="B918EA93BE082927FC55FA27FB71F2E2" box="[917,1222,1454,1478]" italics="true" pageId="8" pageNumber="9">Triplonychus crassifemoris</emphasis>
</taxonomicName>
<emphasis id="B918EA93BE082927FB0EFA27FA9FF2E2" bold="true" box="[1230,1320,1454,1478]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A22B57E8BE082927FB0EFA27FA9FF2E2" box="[1230,1320,1454,1478]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has elytra similar to the
<taxonomicName id="4C6C4D02BE082927FEF4FA5BFE61F2CD" box="[308,470,1490,1513]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cruspinosus">
<emphasis id="B918EA93BE082927FEF4FA5BFE61F2CD" box="[308,470,1490,1513]" italics="true" pageId="8" pageNumber="9">T. cruspinosus</emphasis>
</taxonomicName>
but is distinguishable from it and other species in Schwarz’s second group by its longer mandible, convex and widened metatrochanter and metafemur and widened apicad tibiae. Both species bear a ventral sclerite on the penis and scutellar shield with a short median notch on anterior margin.
</paragraph>
</subSubSection>
</treatment>
</document>
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