217 lines
26 KiB
XML
217 lines
26 KiB
XML
<document id="B46AB4BE2B297B06C74C18E3A47787B7" ID-DOI="10.5281/zenodo.181729" ID-GBIF-Dataset="4f3eeed9-6817-4a8f-8464-95f84391260e" ID-ISSN="1175-5326" ID-Zenodo-Dep="181729" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459838520192" checkinUser="plazi" docAuthor="Grave, De" docDate="2008" docId="03C5878E631B9F19FF46FC1F4946CB72" docLanguage="en" docName="zt01750p052.pdf" docOrigin="Zootaxa 1750" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Leptathanas Grave, 2008, gen. nov." docType="treatment" docVersion="9" lastPageNumber="45" masterDocId="FFFCFFF6631A9F1BFFD1FFD04D64CE48" masterDocTitle="Leptathanas powelli gen. nov., sp. nov, a new infaunal alpheid shrimp associated with upogebiid mudshrimps in Nigeria (Crustacea, Decapoda)" masterLastPageNumber="52" masterPageNumber="43" pageNumber="44" updateTime="1698224011786" updateUser="plazi">
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<mods:mods id="A691C3CE441D8FF4EAF1211D4F281BA7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4910754D3FB8936CD9D27EEA205D55EB">
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<mods:title id="BB64D0CDF7363EDC9A7AAEA21142E4EB">Leptathanas powelli gen. nov., sp. nov, a new infaunal alpheid shrimp associated with upogebiid mudshrimps in Nigeria (Crustacea, Decapoda)</mods:title>
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</mods:titleInfo>
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<mods:name id="E964B8B8378A067EADBFBCBEB068ECC1" type="personal">
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<mods:role id="DF5796A66A7FDA4F28857BB8C449AD88">
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<mods:roleTerm id="5B01224C3D98A0D159238EC69F46D223">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5C527A8A1C0AEA750AAA281CE2FDB26D">Grave, De</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7186687C0C0C96C0C583BDA39FF7721D">text</mods:typeOfResource>
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<mods:relatedItem id="5BCF0C8B635066A82223A36495F30BD6" type="host">
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<mods:titleInfo id="72E665E6D16CC1B75FC26F0ED8650633">
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<mods:title id="DF271CE2727745D5F663444641E4C6C9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5AFBB91FF73713EEC895ADC2F0A23557">
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<mods:date id="ADBEF47F738A2DC1EEB473547D27A7C2">2008</mods:date>
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<mods:detail id="7B782C420EFBA381D294F15BBBE5DD7E" type="volume">
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<mods:number id="55F609D7A6A8C23677ECD24CA1692780">1750</mods:number>
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</mods:detail>
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<mods:extent id="352E5D3F6ADDEDE95AD5AF485034B7E6" unit="page">
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<mods:start id="9DFECAFAA6235513433A08E76E8C7AF1">43</mods:start>
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<mods:end id="C79F5FF6269034E3C9203DB9A1C3C8F2">52</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="017255EB359AF83F0B01A02473C8B74B">journal article</mods:classification>
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<mods:identifier id="73EB5311A5C815AF44BA9191F70ACCB1" type="DOI">10.5281/zenodo.181729</mods:identifier>
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<mods:identifier id="18D1383B40FB327D8AC9952333F5633F" type="GBIF-Dataset">4f3eeed9-6817-4a8f-8464-95f84391260e</mods:identifier>
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<mods:identifier id="700F209FCD1EBBBFEAC0A05FCA91110B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="07C68EECF087D82997488C7D3A110658" type="Zenodo-Dep">181729</mods:identifier>
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</mods:mods>
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<treatment id="03C5878E631B9F19FF46FC1F4946CB72" ID-DOI="http://doi.org/10.5281/zenodo.6228010" ID-GBIF-Taxon="119359528" ID-Zenodo-Dep="6228010" LSID="urn:lsid:plazi:treatment:03C5878E631B9F19FF46FC1F4946CB72" httpUri="http://treatment.plazi.org/id/03C5878E631B9F19FF46FC1F4946CB72" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
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<subSubSection id="C3766513631B9F1AFF46FC1F4CC5CDA1" box="[151,417,975,1001]" pageId="1" pageNumber="44" type="nomenclature">
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<paragraph id="8BD33698631B9F1AFF46FC1F4CC5CDA1" blockId="1.[151,417,975,1001]" box="[151,417,975,1001]" pageId="1" pageNumber="44">
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<heading id="D09B81F4631B9F1AFF46FC1F4CC5CDA1" bold="true" box="[151,417,975,1001]" fontSize="11" level="1" pageId="1" pageNumber="44" reason="1">
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<emphasis id="B918EA8A631B9F1AFF46FC1F4CC5CDA1" bold="true" box="[151,417,975,1001]" pageId="1" pageNumber="44">
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<taxonomicName id="4C6C4D1B631B9F1AFF46FC1F4C48CDA1" ID-CoL="85FV4" box="[151,300,975,1001]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B918EA8A631B9F1AFF46FC1F4C48CDA1" bold="true" box="[151,300,975,1001]" italics="true" pageId="1" pageNumber="44">Leptathanas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22B57F1631B9F1AFEE5FC1F4CC5CDA1" box="[308,417,975,1001]" pageId="1" pageNumber="44" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3766513631B9F1AFF46FBCA4967C94C" pageId="1" pageNumber="44" type="diagnosis">
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<paragraph id="8BD33698631B9F1AFF46FBCA4967C94C" blockId="1.[151,1437,1050,2036]" pageId="1" pageNumber="44">
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<emphasis id="B918EA8A631B9F1AFF46FBCA4C7DCA7C" bold="true" box="[151,281,1050,1076]" pageId="1" pageNumber="44">Diagnosis:</emphasis>
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Carapace glabrous; branchiostegial margin without notch or ventral lip; cardiac notch well developed. Frontal margin with short, broadly triangular rostrum, without orbital teeth; rostral carina weak, reaching to middle of carapace. Pterygostomial angle rounded. Eyes concealed in dorsal and lateral views. Antennular peduncle robust; first segment without ventromesial tooth; stylocerite not appressed, distally acute; second segment as long as broad; lateral antennular flagellum with poorly developed accessory branch. Antenna with robust basicerite; carpocerite not over-reaching scaphocerite. Mouthparts typical for family; mandible with two-segmented palp; first maxilliped with expanded caridean lobe; second maxilliped with large epipod. Third maxilliped pediform; coxa with styliform lateral plate; tip of ultimate segment with cuspidate setae. First pereiopods (chelipeds) asymmetrical in shape, unequal in size, not sexually dimorphic, carried folded with propodus flexed against merus; basis with rudimentary exopod; merus of major cheliped mesially excavated; carpus vase-shaped; chela simple, with ventrally crenulated palm; linea impressa and adhesive discs absent; cutting edges of fingers armed with teeth, finger tips simple. Second pereiopod with five-segmented carpus, chelae simple. Third pereiopod with unarmed ischium and merus; carpus with single distoventral cuspidate seta; propodus mesially with cuspidate setae; dactylus simple. Fifth pereiopod with well developed setal brush on propodus. Second pleopod with appendix interna and appendix masculina in males, with appendix interna only in females. Telson with two pairs of dorsal and two pairs of distolateral cuspidate setae; posterior margin rounded; anal tubercles absent. Gill/exopod formula: 5 pleurobranchs (P1-5), 0 arthrobranch, 0 podobranch, 2 lobe-shaped epipods (Mxp1-2), 4 mastigobranchs (strap-like epipods) (Mxp 3, P1-3), 4 sets of setobranchs (P1-4), 4 exopods (Mxp1-3, P1[rudimentary]).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3766513631B9F1AFF17F8C249F7C964" box="[198,1171,1810,1836]" pageId="1" pageNumber="44" type="materials_examined">
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<paragraph id="8BD33698631B9F1AFF17F8C249F7C964" blockId="1.[151,1437,1050,2036]" box="[198,1171,1810,1836]" pageId="1" pageNumber="44">
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<emphasis id="B918EA8A631B9F1AFF17F8C24C0FC964" bold="true" box="[198,363,1810,1836]" pageId="1" pageNumber="44">
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<typeStatus id="54D7883A631B9F1AFF17F8C24C67C964" box="[198,259,1810,1836]" pageId="1" pageNumber="44">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C6C4D1B631B9F1AFEA3F8C34F04C964" box="[370,608,1811,1836]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="powelli" status="sp. nov.">
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<emphasis id="B918EA8A631B9F1AFEA3F8C34F04C964" box="[370,608,1811,1836]" italics="true" pageId="1" pageNumber="44">Leptathanas powelli</emphasis>
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</taxonomicName>
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<emphasis id="B918EA8A631B9F1AFDB9F8C24FA1C964" bold="true" box="[616,709,1810,1836]" pageId="1" pageNumber="44">
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<taxonomicNameLabel id="A22B57F1631B9F1AFDB9F8C24FA1C964" box="[616,709,1810,1836]" pageId="1" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, by monotypy and present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3766513631B9F19FF17F8EA4946CB72" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44" type="etymology">
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<paragraph id="8BD33698631B9F1AFF17F8EA4FF7C9EC" blockId="1.[151,1437,1050,2036]" pageId="1" pageNumber="44">
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<emphasis id="B918EA8A631B9F1AFF17F8EA4C33C91C" bold="true" box="[198,343,1850,1876]" pageId="1" pageNumber="44">Etymology:</emphasis>
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The generic name is a combination of
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<taxonomicName id="4C6C4D1B631B9F1AFCF1F8EB490FC91C" authority="Williams, 1965" authorityName="Williams" authorityYear="1965" box="[800,1131,1850,1876]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A631B9F1AFCF1F8EB4ECBC91C" box="[800,943,1851,1876]" italics="true" pageId="1" pageNumber="44">Leptalpheus</emphasis>
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Williams, 1965
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</taxonomicName>
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and
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<taxonomicName id="4C6C4D1B631B9F1AFB75F8EB48F8C91C" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[1188,1436,1850,1876]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A631B9F1AFB75F8EB4860C91C" box="[1188,1284,1851,1876]" italics="true" pageId="1" pageNumber="44">Athanas</emphasis>
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Leach,
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<quantity id="4C949B7D631B9F1AFAB0F8EA48F8C91C" box="[1377,1436,1850,1876]" metricMagnitude="1" metricUnit="m" metricValue="4.60756" pageId="1" pageNumber="44" unit="in" value="1814.0">1814</quantity>
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</taxonomicName>
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in reference to the morphological similarity of
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<taxonomicName id="4C6C4D1B631B9F1AFD09F8B34E0DC934" box="[728,873,1891,1916]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B918EA8A631B9F1AFD09F8B34E0DC934" box="[728,873,1891,1916]" italics="true" pageId="1" pageNumber="44">Leptathanas</emphasis>
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</taxonomicName>
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<emphasis id="B918EA8A631B9F1AFCA2F8B24E87C934" bold="true" box="[883,995,1890,1916]" pageId="1" pageNumber="44">
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<taxonomicNameLabel id="A22B57F1631B9F1AFCA2F8B24E87C934" box="[883,995,1890,1916]" pageId="1" pageNumber="44" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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with
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<taxonomicName id="4C6C4D1B631B9F1AFBFDF8B349DFC934" box="[1068,1211,1891,1916]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A631B9F1AFBFDF8B349DFC934" box="[1068,1211,1891,1916]" italics="true" pageId="1" pageNumber="44">Leptalpheus</emphasis>
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</taxonomicName>
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and relatedness to
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<taxonomicName id="4C6C4D1B631B9F1AFF46F85B4D93C9EC" box="[151,247,1931,1956]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A631B9F1AFF46F85B4D93C9EC" box="[151,247,1931,1956]" italics="true" pageId="1" pageNumber="44">Athanas</emphasis>
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</taxonomicName>
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(see below). Gender is masculine.
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</paragraph>
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<paragraph id="8BD33698631B9F19FF17F8624F97CC22" blockId="1.[151,1437,1050,2036]" lastBlockId="2.[151,1437,152,1338]" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
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<emphasis id="B918EA8A631B9F1AFF17F8624CBFC984" bold="true" box="[198,475,1970,1996]" pageId="1" pageNumber="44">Generic relationships:</emphasis>
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<taxonomicName id="4C6C4D1B631B9F1AFE38F8634F1EC984" box="[489,634,1971,1996]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B918EA8A631B9F1AFE38F8634F1EC984" box="[489,634,1971,1996]" italics="true" pageId="1" pageNumber="44">Leptathanas</emphasis>
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</taxonomicName>
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<emphasis id="B918EA8A631B9F1AFD50F8624F8BC984" bold="true" box="[641,751,1970,1996]" pageId="1" pageNumber="44">
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<taxonomicNameLabel id="A22B57F1631B9F1AFD50F8624F8BC984" box="[641,751,1970,1996]" pageId="1" pageNumber="44" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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is superficially similar to
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<taxonomicName id="4C6C4D1B631B9F1AFBF1F86349CBC984" box="[1056,1199,1971,1996]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A631B9F1AFBF1F86349CBC984" box="[1056,1199,1971,1996]" italics="true" pageId="1" pageNumber="44">Leptalpheus</emphasis>
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</taxonomicName>
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, e.g., by the general shape of the frontal margin, chelipeds (P1) and walking legs. However,
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<taxonomicName id="4C6C4D1B631B9F1AFC3FF80B491BC9BC" box="[1006,1151,2011,2036]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B918EA8A631B9F1AFC3FF80B491BC9BC" box="[1006,1151,2011,2036]" italics="true" pageId="1" pageNumber="44">Leptathanas</emphasis>
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</taxonomicName>
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<emphasis id="B918EA8A631B9F1AFB56F80A4991C9BC" bold="true" box="[1159,1269,2010,2036]" pageId="1" pageNumber="44">
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<taxonomicNameLabel id="A22B57F1631B9F1AFB56F80A4991C9BC" box="[1159,1269,2010,2036]" pageId="1" pageNumber="44" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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can be distinguished from
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<taxonomicName id="4C6C4D1B63189F19FEEBFF484CADCEF9" box="[314,457,152,177]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A63189F19FEEBFF484CADCEF9" box="[314,457,152,177]" italics="true" pageId="2" pageNumber="45">Leptalpheus</emphasis>
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</taxonomicName>
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by the absence of an arthrobranch at the base of Mxp3; the presence of a rudimentary exopod on the basis of P1; the absence of a mesioventral tooth on the first segment of the antennular peduncle; the absence of a deep incision and a mesial tooth on the uropodal diaeresis; the presence of cuspidate setae on the uropodal protopod; the presence of mastigobranchs on coxae of Mxp3 and P1-3 (vs. Mxp3 and P
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<quantity id="4C949B7D63189F19FF0DFEE84C42CF1A" box="[220,294,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="2" pageNumber="45" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1-4 in</quantity>
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<taxonomicName id="4C6C4D1B63189F19FEFFFEE84CD9CF19" box="[302,445,312,337]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A63189F19FEFFFEE84CD9CF19" box="[302,445,312,337]" italics="true" pageId="2" pageNumber="45">Leptalpheus</emphasis>
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</taxonomicName>
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); the minor chela with short stout fingers (vs. long and slender in
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<taxonomicName id="4C6C4D1B63189F19FB18FEE8483CCF19" box="[1225,1368,312,337]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A63189F19FB18FEE8483CCF19" box="[1225,1368,312,337]" italics="true" pageId="2" pageNumber="45">Leptalpheus</emphasis>
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</taxonomicName>
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); and the ultimate segment of the Mxp3 without distally thickened setae, but with cuspidate setae on the tip (vs. with dense, distally thickened setae and unarmed tip in
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<taxonomicName id="4C6C4D1B63189F19FD3AFE584E1ECFE9" box="[747,890,392,417]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A63189F19FD3AFE584E1ECFE9" box="[747,890,392,417]" italics="true" pageId="2" pageNumber="45">Leptalpheus</emphasis>
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</taxonomicName>
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). All species of
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<taxonomicName id="4C6C4D1B63189F19FB97FE5849B1CFE9" box="[1094,1237,392,417]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EA8A63189F19FB97FE5849B1CFE9" box="[1094,1237,392,417]" italics="true" pageId="2" pageNumber="45">Leptalpheus</emphasis>
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</taxonomicName>
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and related genera (= leptalpheoid lineage or ALF clade in Anker
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<emphasis id="B918EA8A63189F19FCD3FE604E5BCF81" box="[770,831,432,457]" italics="true" pageId="2" pageNumber="45">et al.</emphasis>
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, 2006) possess a hyper-developed arthrobranch; a deep incision and a strong mesial tooth on the uropodal diaeresis; the ultimate segment of Mxp3 with dense rows of distally thickened setae and without apical cuspidate setae; and a minor chela with long slender fingers, usually with toothed cutting edges. The absence of these features in
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<taxonomicName id="4C6C4D1B63189F19FC28FDF849EECC09" box="[1017,1162,552,577]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B918EA8A63189F19FC28FDF849EECC09" box="[1017,1162,552,577]" italics="true" pageId="2" pageNumber="45">Leptathanas</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B918EA8A63189F19FB40FDF7499BCC09" bold="true" box="[1169,1279,551,577]" pageId="2" pageNumber="45">
|
|
<taxonomicNameLabel id="A22B57F163189F19FB40FDF7499BCC09" box="[1169,1279,551,577]" pageId="2" pageNumber="45" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
</emphasis>
|
|
suggests that the new genus is not closely related to
|
|
<taxonomicName id="4C6C4D1B63189F19FD8CFD804F88CC21" box="[605,748,592,617]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B918EA8A63189F19FD8CFD804F88CC21" box="[605,748,592,617]" italics="true" pageId="2" pageNumber="45">Leptalpheus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BD3369863189F19FF14FDA84946CB72" blockId="2.[151,1437,152,1338]" pageId="2" pageNumber="45">
|
|
On the other hand, several features of
|
|
<taxonomicName id="4C6C4D1B63189F19FD5EFDA84E44CCD9" box="[655,800,632,657]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="B918EA8A63189F19FD5EFDA84E44CCD9" box="[655,800,632,657]" italics="true" pageId="2" pageNumber="45">Leptathanas</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B918EA8A63189F19FCF8FDA74EFCCCD9" bold="true" box="[809,920,631,657]" pageId="2" pageNumber="45">
|
|
<taxonomicNameLabel id="A22B57F163189F19FCF8FDA74EFCCCD9" box="[809,920,631,657]" pageId="2" pageNumber="45" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
</emphasis>
|
|
, such as the absence of an arthrobranch on Mxp3; the presence of a rudimentary exopod on P1; the presence of a cuspidate seta on the dorsal margin of the ischium of P1; and the absence of a mastigobranch on P4, are also present in members of the generic complex around
|
|
<taxonomicName id="4C6C4D1B63189F19FEF8FD204F45CD42" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[297,545,752,778]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B918EA8A63189F19FEF8FD204CEDCD41" box="[297,393,752,777]" italics="true" pageId="2" pageNumber="45">Athanas</emphasis>
|
|
Leach, 1814
|
|
</taxonomicName>
|
|
(= athanoid lineage or clade AP in Anker
|
|
<emphasis id="B918EA8A63189F19FBDEFD20492DCD41" box="[1039,1097,752,777]" italics="true" pageId="2" pageNumber="45">et al.</emphasis>
|
|
, 2006). Among these,
|
|
<taxonomicName id="4C6C4D1B63189F19FA8AFD204CFECD7A" authority="Coutiere, 1896" authorityName="Coutiere" authorityYear="1896" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B918EA8A63189F19FA8AFD204D86CD79" italics="true" pageId="2" pageNumber="45">Athanopsis</emphasis>
|
|
Coutière, 1896
|
|
</taxonomicName>
|
|
with five species all in the Indo-West Pacific (
|
|
<bibRefCitation id="EFFD4B6963189F19FC15FCC84994CD7A" author="Anker" box="[964,1264,792,818]" pageId="2" pageNumber="52" refString="Anker, A. & Ahyong, S. T. (2007 b) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007" yearSuffix="b">Anker & Ahyong, 2007b</bibRefCitation>
|
|
) appears to be morphologically closest to the new genus, especially in the general shape and size ratio of the major and minor chelipeds. For instance, the major cheliped in both genera is characterized by one or several cuspidate setae on the ischium; a distally broadened, ventrally excavated and laterally rugose merus; a tuberculate ridge on the ventral margin of the palm; and the cutting edges of the dactylus and pollex each armed with one or a few blunt teeth. However,
|
|
<taxonomicName id="4C6C4D1B63189F19FE02FC304F00CDB1" box="[467,612,992,1017]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="B918EA8A63189F19FE02FC304F00CDB1" box="[467,612,992,1017]" italics="true" pageId="2" pageNumber="45">Leptathanas</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A22B57F163189F19FDBAFC0F4FBDCDB1" box="[619,729,991,1017]" pageId="2" pageNumber="45" rank="genus">
|
|
<emphasis id="B918EA8A63189F19FDBAFC0F4FBDCDB1" bold="true" box="[619,729,991,1017]" pageId="2" pageNumber="45">gen. nov.</emphasis>
|
|
</taxonomicNameLabel>
|
|
can be easily separated from
|
|
<taxonomicName id="4C6C4D1B63189F19FBE2FC3049D1CDB1" box="[1075,1205,992,1017]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B918EA8A63189F19FBE2FC3049D1CDB1" box="[1075,1205,992,1017]" italics="true" pageId="2" pageNumber="45">Athanopsis</emphasis>
|
|
</taxonomicName>
|
|
by the reduced rostrum (vs. well developed with rounded tip in
|
|
<taxonomicName id="4C6C4D1B63189F19FD6CFBD84E5BCA69" box="[701,831,1032,1057]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B918EA8A63189F19FD6CFBD84E5BCA69" box="[701,831,1032,1057]" italics="true" pageId="2" pageNumber="45">Athanopsis</emphasis>
|
|
</taxonomicName>
|
|
); the absence of orbital teeth (present and usually well developed in
|
|
<taxonomicName id="4C6C4D1B63189F19FEA6FBE04C9DCA01" box="[375,505,1072,1097]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B918EA8A63189F19FEA6FBE04C9DCA01" box="[375,505,1072,1097]" italics="true" pageId="2" pageNumber="45">Athanopsis</emphasis>
|
|
</taxonomicName>
|
|
); the absence of a mesioventral tooth on the first segment of the antennular peduncle; the five-segmented carpus of the P2; the absence of cuspidate setae on the ischium of P3; and finally, by the presence of a row of cuspidate setae on the uropodal protopod, a feature that appears to be unique within the family (A. Anker, pers. obs.). This last feature, in combination with other characters (see above) enables to separate
|
|
<taxonomicName id="4C6C4D1B63189F19FE03FB004F07CAA1" box="[466,611,1232,1257]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="B918EA8A63189F19FE03FB004F07CAA1" box="[466,611,1232,1257]" italics="true" pageId="2" pageNumber="45">Leptathanas</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B918EA8A63189F19FDBBFB1F4FBCCAA1" bold="true" box="[618,728,1231,1257]" pageId="2" pageNumber="45">
|
|
<taxonomicNameLabel id="A22B57F163189F19FDBBFB1F4FBCCAA1" box="[618,728,1231,1257]" pageId="2" pageNumber="45" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
</emphasis>
|
|
from all other alpheid genera. The remarkable convergence between
|
|
<taxonomicName id="4C6C4D1B63189F19FED1FB284CEBCB59" box="[256,399,1272,1297]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B918EA8A63189F19FED1FB284CEBCB59" box="[256,399,1272,1297]" italics="true" pageId="2" pageNumber="45">Leptalpheus</emphasis>
|
|
</taxonomicName>
|
|
from the leptalpheoid lineage and
|
|
<taxonomicName id="4C6C4D1B63189F19FCF9FB284EDDCB59" box="[808,953,1272,1297]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="B918EA8A63189F19FCF9FB284EDDCB59" box="[808,953,1272,1297]" italics="true" pageId="2" pageNumber="45">Leptathanas</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B918EA8A63189F19FC11FB27494ACB59" bold="true" box="[960,1070,1271,1297]" pageId="2" pageNumber="45">
|
|
<taxonomicNameLabel id="A22B57F163189F19FC11FB27494ACB59" box="[960,1070,1271,1297]" pageId="2" pageNumber="45" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
</emphasis>
|
|
from the athanoid lineage may be explained by very similar infaunal life styles of these shrimps (see below).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |