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<document id="27D2ACE3BE15D2D7167A055F8F575798" ID-DOI="10.11646/zootaxa.3972.2.8" ID-GBIF-Dataset="44196629-f926-495d-87ac-d5c85ea5f8b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="242992" ID-ZooBank="8B3F8B14-97C4-4AFE-B299-2DEBC32BDE7A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461187775986" checkinUser="plazi" docAuthor="Barkalov, Anatolij V. &amp; Ståhls, Gunilla" docDate="2015" docId="03FC4D7058164D18FF1EFF0EFA2F025A" docLanguage="en" docName="zt03972p290.pdf" docOrigin="Zootaxa 3972 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheilosia Barkalov &amp; Ståhls, 2015, sp. n." docType="treatment" docVersion="7" lastPageNumber="285" masterDocId="FFC5350858134D1DFF89FF99FFB30422" masterDocTitle="Descriptions of three new species of the genus Cheilosia Meigen from China (Diptera, Syrphidae)" masterLastPageNumber="290" masterPageNumber="280" pageNumber="285" updateTime="1698599953686" updateUser="plazi">
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<mods:title id="7AF4CC4EAB41085E7FC51E98549C0B11">Descriptions of three new species of the genus Cheilosia Meigen from China (Diptera, Syrphidae)</mods:title>
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<mods:namePart id="DC79C49F92F4FE6E273BA7C3230A9442">Barkalov, Anatolij V.</mods:namePart>
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<mods:namePart id="E06E0FB32F58C8E9B2BC775889C9B67A">Ståhls, Gunilla</mods:namePart>
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<mods:date id="24E62187147043B04CF3E2F0FD829A65">2015</mods:date>
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<treatment id="03FC4D7058164D18FF1EFF0EFA2F025A" ID-DOI="http://doi.org/10.5281/zenodo.6103318" ID-GBIF-Taxon="119628794" ID-Zenodo-Dep="6103318" LSID="urn:lsid:plazi:treatment:03FC4D7058164D18FF1EFF0EFA2F025A" httpUri="http://treatment.plazi.org/id/03FC4D7058164D18FF1EFF0EFA2F025A" lastPageNumber="285" pageId="5" pageNumber="285">
<subSubSection id="C34FAFED58164D18FF1EFF0EFE9104F1" pageId="5" pageNumber="285" type="nomenclature">
<paragraph id="8BEAFC6658164D18FF1EFF0EFC870490" blockId="5.[151,820,151,211]" box="[151,820,151,178]" pageId="5" pageNumber="285">
<heading id="D0A24B0A58164D18FF1EFF0EFC870490" bold="true" box="[151,820,151,178]" fontSize="11" level="1" pageId="5" pageNumber="285" reason="1">
<emphasis id="B921207458164D18FF1EFF0EFC870490" bold="true" box="[151,820,151,178]" pageId="5" pageNumber="285">
<emphasis id="B921207458164D18FF1EFF0EFE4D0493" bold="true" box="[151,510,151,177]" italics="true" pageId="5" pageNumber="285">
<taxonomicName id="4C5587E558164D18FF1EFF0EFEBA0493" ID-CoL="3MHY" box="[151,265,151,177]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="genus">Cheilosia</taxonomicName>
(s. str.)
<taxonomicName id="4C5587E558164D18FEE3FF0EFE4D0493" authority="Barkalov &amp; Stahls" authorityName="Barkalov &amp; Stahls" box="[362,510,151,177]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="species" species="lamproptera" status="sp. nov.">lamproptera</taxonomicName>
</emphasis>
Barkalov &amp; Ståhls
<taxonomicNameLabel id="A2129D0F58164D18FD7BFF01FC870490" box="[754,820,152,178]" pageId="5" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEAFC6658164D18FF1EFF23FE9104F1" blockId="5.[151,820,151,211]" box="[151,290,186,211]" pageId="5" pageNumber="285">
(
<figureCitation id="136EE0E358164D18FF16FF23FF5304F1" box="[159,224,186,211]" captionStart="FIGURE 1. A D" captionStartId="4.[151,250,1688,1710]" captionTargetBox="[151,1429,212,1659]" captionTargetId="figure@4.[151,1435,195,1666]" captionTargetPageId="4" captionText="FIGURE 1. A D: Cheilosia (Eucartosyrphus) bullabucca sp. n. E G: Cheilosia (s. str.) lamproptera sp. n. A &amp; F: head laterally, B &amp; E: head dorsally, C: tip of wing, D &amp; G: 2 3 antennal segments laterally." httpUri="https://zenodo.org/record/242993/files/figure.png" pageId="5" pageNumber="285">Figs 1</figureCitation>
EG)
</paragraph>
</subSubSection>
<subSubSection id="C34FAFED58164D18FF1EFE9BFE130540" pageId="5" pageNumber="285" type="materials_examined">
<paragraph id="8BEAFC6658164D18FF1EFE9BFE130540" blockId="5.[151,1437,258,1441]" pageId="5" pageNumber="285">
<emphasis id="B921207458164D18FF1EFE9BFDB10539" bold="true" box="[151,514,258,283]" pageId="5" pageNumber="285">
<typeStatus id="54EE42C458164D18FF1EFE9BFF610539" box="[151,210,258,283]" pageId="5" pageNumber="285">Type</typeStatus>
material.
<typeStatus id="54EE42C458164D18FED5FE9BFE4E0539" box="[348,509,258,283]" pageId="5" pageNumber="285" type="holotype">HOLOTYPE</typeStatus>
.
</emphasis>
♀,
<collectingCountry id="F342BCF658164D18FDB5FE9BFD310539" box="[572,642,258,283]" name="China" pageId="5" pageNumber="285">China</collectingCountry>
, Yunnan Province, Gongga Shan,
<quantity id="4CAD518358164D18FBA0FE9AFBCA0539" box="[1065,1145,258,283]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="285" unit="km" value="40.0">40 km</quantity>
NW Dulong,
<geoCoordinate id="EE619AA158164D18FAB9FE9AFA2F0539" box="[1328,1436,259,283]" direction="north" orientation="latitude" pageId="5" pageNumber="285" precision="55" value="27.984">27.984°N</geoCoordinate>
<geoCoordinate id="EE619AA158164D18FF1EFEBEFEB0051D" box="[151,259,295,319]" direction="east" orientation="longitude" pageId="5" pageNumber="285" precision="55" value="98.324">98.324°E</geoCoordinate>
,
<quantity id="4CAD518358164D18FE99FEBEFEDB051C" box="[272,360,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="285" unit="m" value="1700.0">1700 m</quantity>
above sea level, 8.06.2009, leg. Blank, Liston &amp; Taeger. 0 22
<collectingCountry id="F342BCF658164D18FBBFFEBFFBCF051D" box="[1078,1148,294,319]" name="China" pageId="5" pageNumber="285">China</collectingCountry>
,
<collectionCode id="ED4464A358164D18FB00FEBEFB75051D" LSID="urn:lsid:biocol.org:col:15536" box="[1161,1222,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="5" pageNumber="285">DNA</collectionCode>
voucher specimen
<collectionCode id="ED4464A358164D18FF1EFED2FF6B0541" box="[151,216,331,355]" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="5" pageNumber="285">MZH</collectionCode>
_
<collectingCountry id="F342BCF658164D18FF6EFED2FF480541" box="[231,251,331,355]" name="Italy" pageId="5" pageNumber="285">Y</collectingCountry>
1918, in
<collectionCode id="ED4464A358164D18FED3FED2FE2F0540" box="[346,412,331,354]" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="5" pageNumber="285">MZH</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C34FAFED58164D18FF4EFEF7FCED05EC" pageId="5" pageNumber="285" type="diagnosis">
<paragraph id="8BEAFC6658164D18FF4EFEF7FCED05EC" blockId="5.[151,1437,258,1441]" pageId="5" pageNumber="285">
<emphasis id="B921207458164D18FF4EFEF7FEF205A5" bold="true" box="[199,321,366,391]" pageId="5" pageNumber="285">Diagnosis.</emphasis>
The presence of a character combination unusual for species of
<taxonomicName id="4C5587E558164D18FBE3FEF7FB6605A4" box="[1130,1237,366,390]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B921207458164D18FBE3FEF7FB6605A4" box="[1130,1237,366,390]" italics="true" pageId="5" pageNumber="285">Cheilosia</emphasis>
</taxonomicName>
, the absence of microtrichia on wing cells M and CuA, a densely haired face and mostly yellow legs, facilitates the distinction of the new species from all other members of the genus
<taxonomicName id="4C5587E558164D18FD65FE2FFCE405EC" box="[748,855,438,462]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B921207458164D18FD65FE2FFCE405EC" box="[748,855,438,462]" italics="true" pageId="5" pageNumber="285">Cheilosia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34FAFED58164D18FF4EFE43FA2200EB" pageId="5" pageNumber="285" type="description">
<paragraph id="8BEAFC6658164D18FF4EFE43FA2200EB" blockId="5.[151,1437,258,1441]" pageId="5" pageNumber="285">
<emphasis id="B921207458164D18FF4EFE43FE6D05D1" bold="true" box="[199,478,474,499]" pageId="5" pageNumber="285">Description. FEMALE.</emphasis>
Body length
<quantity id="4CAD518358164D18FDFFFE42FD7C05D0" box="[630,719,475,499]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="5" pageNumber="285" unit="mm" value="8.7">8.7 mm</quantity>
, wing length
<quantity id="4CAD518358164D18FCEEFE43FC7305D0" box="[871,960,474,499]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="285" unit="mm" value="8.5">8.5 mm</quantity>
.
<emphasis id="B921207458164D18FC42FE43FBA105D1" bold="true" box="[971,1042,474,499]" pageId="5" pageNumber="285">Head.</emphasis>
Face comparatively narrow, black, shiny, with stripes of fine grey dusting, completely covered with dense light hairs (
<figureCitation id="136EE0E358164D18FBB3FE67FBC90635" box="[1082,1146,510,535]" captionStart="FIGURE 1. A D" captionStartId="4.[151,250,1688,1710]" captionTargetBox="[151,1429,212,1659]" captionTargetId="figure@4.[151,1435,195,1666]" captionTargetPageId="4" captionText="FIGURE 1. A D: Cheilosia (Eucartosyrphus) bullabucca sp. n. E G: Cheilosia (s. str.) lamproptera sp. n. A &amp; F: head laterally, B &amp; E: head dorsally, C: tip of wing, D &amp; G: 2 3 antennal segments laterally." httpUri="https://zenodo.org/record/242993/files/figure.png" pageId="5" pageNumber="285">Fig. 1</figureCitation>
F); eye margins moderate in width, black, in lower part brownish, covered with dense grey dusting and short white hairs; cheeks comparatively low, covered with fine dusting and white hairs. Frons narrow (
<figureCitation id="136EE0E358164D18FBDAFDDFFB28067D" box="[1107,1179,582,607]" captionStart="FIGURE 1. A D" captionStartId="4.[151,250,1688,1710]" captionTargetBox="[151,1429,212,1659]" captionTargetId="figure@4.[151,1435,195,1666]" captionTargetPageId="4" captionText="FIGURE 1. A D: Cheilosia (Eucartosyrphus) bullabucca sp. n. E G: Cheilosia (s. str.) lamproptera sp. n. A &amp; F: head laterally, B &amp; E: head dorsally, C: tip of wing, D &amp; G: 2 3 antennal segments laterally." httpUri="https://zenodo.org/record/242993/files/figure.png" pageId="5" pageNumber="285">Fig. 1</figureCitation>
E), slightly narrowed posteriorly, shiny with narrow stripes of grey dusting near eyes; covered with not very long yellow hairs and mixed with some black hairs near ocellar triangle; lunula bright yellow; antennal pits broadly separated. Antenna brownish, basoflagellomere rounded, light-brown; arista long, thin, yellow, covered with distinct but short hairs (
<figureCitation id="136EE0E358164D18FF16FD4FFF6C06CD" box="[159,223,726,751]" captionStart="FIGURE 1. A D" captionStartId="4.[151,250,1688,1710]" captionTargetBox="[151,1429,212,1659]" captionTargetId="figure@4.[151,1435,195,1666]" captionTargetPageId="4" captionText="FIGURE 1. A D: Cheilosia (Eucartosyrphus) bullabucca sp. n. E G: Cheilosia (s. str.) lamproptera sp. n. A &amp; F: head laterally, B &amp; E: head dorsally, C: tip of wing, D &amp; G: 2 3 antennal segments laterally." httpUri="https://zenodo.org/record/242993/files/figure.png" pageId="5" pageNumber="285">Fig. 1</figureCitation>
G). Eyes completely covered with dense whitish hairs. Vertex flat, shiny with yellow hairs, ocellar triangle equilateral. Occiput behind ocellar triangle shiny, otherwise covered with grey dusting.
<emphasis id="B921207458164D18FB1DFD63FB470731" bold="true" box="[1172,1268,762,787]" pageId="5" pageNumber="285">Thorax.</emphasis>
Postpronotum black with brownish outer angle, covered with grey dusting and yellow hairs. Scutum and scutellum black, shiny with comparatively short semi-erect yellow hairs; postalar callus brownish, without black bristles, scutellum hind margin with long yellow hairs (some of same length or longer than scutellum length), without black bristles. Subscutellar fringe with yellow hairs, of about same length as on scutum. Pleura black with grey dusting and yellow hairs. Dorsal and ventral hair patches on katepisternum narrowly separated. Legs: coxa black with narrowly yellow tips; trochanter brown anteriorly and black posteriorly; fore femur black posteriorly and yellow on basal 1/ 5 and apical half anteriorly; mid and hind femora black with narrowly yellow base and tips, all femora covered with yellow hairs; tibia yellow with narrow brownish spot postero-medially, mid tibia with comb of strong black short hairs (ctenidium) apically; tarsi yellow with black 5th segment. Wings yellowish with yellow stigma and veins; cell M and cell CuA with large area without microtrichia; inner angle between veins M1 and R4+5 almost 90°.
<emphasis id="B921207458164D18FF1EFB15FEA00087" bold="true" box="[151,275,1164,1189]" pageId="5" pageNumber="285">Abdomen.</emphasis>
Narrower basally, the broadest part at the hind margin of tergite III, black, except grey dusted tergite I, shiny; covered with comparatively short, yellow, medially semi-erect and laterally erect hairs.
<emphasis id="B921207458164D18FB30FB29FAA700EB" bold="true" box="[1209,1300,1200,1225]" pageId="5" pageNumber="285">MALE.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34FAFED58164D18FF4EFB4DFA3000CF" box="[199,1411,1236,1261]" pageId="5" pageNumber="285" type="etymology">
<paragraph id="8BEAFC6658164D18FF4EFB4DFA3000CF" blockId="5.[151,1437,258,1441]" box="[199,1411,1236,1261]" pageId="5" pageNumber="285">
<emphasis id="B921207458164D18FF4EFB4DFEFD00CF" bold="true" box="[199,334,1236,1261]" pageId="5" pageNumber="285">Etymology.</emphasis>
The Greek word “
<taxonomicName id="4C5587E558164D18FD95FB4DFD1B00CF" box="[540,680,1236,1261]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="species" species="lamproptera">lamproptera</taxonomicName>
” means “shiny-wing” referring to the shiny wings of the species.
</paragraph>
</subSubSection>
<subSubSection id="C34FAFED58164D18FF4EFB61FA2F025A" pageId="5" pageNumber="285" type="distribution">
<paragraph id="8BEAFC6658164D18FF4EFB61FD330133" blockId="5.[151,1437,258,1441]" box="[199,640,1272,1297]" pageId="5" pageNumber="285">
<emphasis id="B921207458164D18FF4EFB61FED30133" bold="true" box="[199,352,1272,1297]" pageId="5" pageNumber="285">Distribution.</emphasis>
<collectingCountry id="F342BCF658164D18FEEEFB61FE1D0133" box="[359,430,1272,1297]" name="China" pageId="5" pageNumber="285">China</collectingCountry>
: Yunnan Province.
</paragraph>
<paragraph id="8BEAFC6658164D18FF4EFA85FDCD0183" blockId="5.[151,1437,258,1441]" pageId="5" pageNumber="285">
<emphasis id="B921207458164D18FF4EFA85FEF90117" bold="true" box="[199,330,1308,1333]" pageId="5" pageNumber="285">Taxonomy.</emphasis>
The taxon shares the following characters with taxa of nominative subgenus: antennal pits broadly separated, eyes covered with dense hairs, not very dense hairs on body and all habitus is similar with other members of
<emphasis id="B921207458164D18FEADFAFDFE61015E" box="[292,466,1380,1404]" italics="true" pageId="5" pageNumber="285">
<taxonomicName id="4C5587E558164D18FEADFAFDFE3C015E" box="[292,399,1380,1404]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="genus" sensu="stricto">Cheilosia</taxonomicName>
<taxonomicNameLabel id="A2129D0F58164D18FE1FFAFCFE61015E" box="[406,466,1381,1404]" pageId="5" pageNumber="285" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
In the key to Chinese
<taxonomicName id="4C5587E558164D18FD59FAFDFBD9015F" authority="Barkalov &amp; Cheng 2004" authorityName="Barkalov &amp; Cheng" authorityYear="2004" box="[720,1130,1380,1405]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B921207458164D18FD59FAFDFC8F015E" box="[720,828,1380,1404]" italics="true" pageId="5" pageNumber="285">Cheilosia</emphasis>
(
<bibRefCitation id="EFC4819758164D18FCC3FAFDFBD1015F" author="Barkalov" box="[842,1122,1380,1405]" pageId="5" pageNumber="285" refString="Barkalov, A. V. &amp; Cheng, X. - Y. (2004) Revision of the genus Cheilosia Meigen, 1822 (Diptera: Syrphidae) of China. Contributions on Entomology, International, 5 (3 / 4), 266 - 421." type="journal article" year="2004">Barkalov &amp; Cheng 2004</bibRefCitation>
)
</taxonomicName>
the new species is inserted at couplet 39 and will key out as following:
</paragraph>
<paragraph id="8BEAFC6658164D18FF1EFA56FA2F025A" blockId="5.[151,1436,1487,1656]" pageId="5" pageNumber="285">
39. Face strongly protruded forward........................................................
<taxonomicName id="4C5587E558164D18FB4AFA56FA2F01C7" authority="Shiraki" authorityName="Shiraki" box="[1219,1436,1487,1509]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="species" species="ochreipila">
<emphasis id="B921207458164D18FB4AFA56FAF201C7" box="[1219,1345,1487,1509]" italics="true" pageId="5" pageNumber="285">C. ochreipila</emphasis>
(Shiraki)
</taxonomicName>
- Face less protruded forward (
<figureCitation id="136EE0E358164D18FE73FA75FD830220" box="[506,560,1516,1538]" captionStart="FIGURE 1. A D" captionStartId="4.[151,250,1688,1710]" captionTargetBox="[151,1429,212,1659]" captionTargetId="figure@4.[151,1435,195,1666]" captionTargetPageId="4" captionText="FIGURE 1. A D: Cheilosia (Eucartosyrphus) bullabucca sp. n. E G: Cheilosia (s. str.) lamproptera sp. n. A &amp; F: head laterally, B &amp; E: head dorsally, C: tip of wing, D &amp; G: 2 3 antennal segments laterally." httpUri="https://zenodo.org/record/242993/files/figure.png" pageId="5" pageNumber="285">Fig. 1</figureCitation>
F).................................................................... 40 40. Smaller species: 7.0
<quantity id="4CAD518358164D18FE27F993FE4A023D" box="[430,505,1546,1568]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="285" unit="mm" value="7.3">7.3 mm</quantity>
, hind tarsi black..................................
<taxonomicName id="4C5587E558164D18FBC8F993FA280202" authority="Barkalov &amp; Cheng" authorityName="Barkalov &amp; Cheng" box="[1089,1435,1546,1568]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="species" species="intonsaformis">
<emphasis id="B921207458164D18FBC8F993FB520202" box="[1089,1249,1546,1568]" italics="true" pageId="5" pageNumber="285">C. intonsaformis</emphasis>
Barkalov &amp; Cheng
</taxonomicName>
- Bigger species: 8.512.0 mm, hind tarsi yellow except black apical segment..................................... 41 41. Cells M and CuA almost completely without microtrichia......................................
<taxonomicName id="4C5587E558164D18FB40F9DDFAEC0278" box="[1225,1375,1604,1626]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="species" species="lamproptera" status="sp. nov.">
<emphasis id="B921207458164D18FB40F9DDFAEC0278" box="[1225,1375,1604,1626]" italics="true" pageId="5" pageNumber="285">C. lamproptera</emphasis>
</taxonomicName>
<emphasis id="B921207458164D18FAEFF9DCFA2F0279" bold="true" box="[1382,1436,1605,1627]" pageId="5" pageNumber="285">
<taxonomicNameLabel id="A2129D0F58164D18FAEFF9DCFA2F0279" box="[1382,1436,1605,1627]" pageId="5" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
- Cells M and CuA completely covered with microtrichia.......................................
<taxonomicName id="4C5587E558164D18FB41F9FBFA2F025A" authority="Shiraki" authorityName="Shiraki" box="[1224,1436,1634,1656]" class="Insecta" family="Syrphidae" genus="Cheilosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="285" phylum="Arthropoda" rank="species" species="okinawae">
<emphasis id="B921207458164D18FB41F9FBFAF3025A" box="[1224,1344,1634,1656]" italics="true" pageId="5" pageNumber="285">C. okinawae</emphasis>
(Shiraki)
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>