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<document id="150E2C342D2111A638C3CD00AD6274D4" ID-CLB-Dataset="58033" ID-DOI="10.5852/ejt.2023.882.2177" ID-GBIF-Dataset="57d1d191-2bf5-4229-a0f5-9ab472a114a6" ID-ISSN="2118-9773" ID-Zenodo-Dep="8177581" ID-ZooBank="67C909E4-C700-4F8D-B8CE-5FD9B2C5D549" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1690201644379" checkinUser="felipe" docAuthor="Korn, Dieter &amp; Weyer, Dieter" docDate="2023" docId="03EA5C14CAB885FCFDAFF939FC4486C9" docLanguage="en" docName="ejt-882_1-230.pdf" docOrigin="European Journal of Taxonomy 882" docSource="http://dx.doi.org/10.5852/ejt.2023.882.2177" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Paprothites kullmanni Korn &amp; Weyer 2023, sp. nov." docType="treatment" docUuid="E347E068-0C6F-4263-BE30-7540C182E55C" docUuidSource="ZooBank" docVersion="15" lastPageNumber="180" masterDocId="FFD3246CCA09854FFFCEFF90FFB2846D" masterDocTitle="The ammonoids from the Gattendorfia Limestone of Oberrödinghausen (Early Carboniferous; Rhenish Mountains, Germany)" masterLastPageNumber="230" masterPageNumber="1" pageNumber="178" updateTime="1699545181523" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="8399039EF794DCDADE538E92E4F8ECE4">
<mods:title id="99925F90AF05EABF966A05BE6E02A7A6">The ammonoids from the Gattendorfia Limestone of Oberrödinghausen (Early Carboniferous; Rhenish Mountains, Germany)</mods:title>
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<mods:name id="3ECD1BE8AE6856966BF90426257B9C52" type="personal">
<mods:role id="2152F91FC09F2C3C3B636403A842AA52">
<mods:roleTerm id="DF23AA53E3657BA0FF6AA17AAD0D6477">Author</mods:roleTerm>
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<mods:namePart id="6C3FA10881B229D46BAF34AC74583112">Korn, Dieter</mods:namePart>
<mods:nameIdentifier id="140EB5DC81A35150668B6FB7A6F8AC36" type="LSID">286CA4F3-7EBC-4AEF-A66A-B2508D001367</mods:nameIdentifier>
<mods:affiliation id="44BD47714D3B9B45B6F47596ADCA2AC9">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="E13F7A2ED5858F346D48C41DA921C422" type="email">dieter.korn@mfn.berlin</mods:nameIdentifier>
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<mods:role id="02939659D5FE2A9D98592BB6C309D7D1">
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<mods:namePart id="67A8A462881D29244DDFF927874C216B">Weyer, Dieter</mods:namePart>
<mods:nameIdentifier id="46B28B5990AC91F4876A83B6F21DBF6A" type="LSID">A09A1738-C70E-4F22-A069-8B7DB4A1753D</mods:nameIdentifier>
<mods:affiliation id="F81216AE95D65F28ACF02191B8AA5462">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="27F8080484FA78BCCE9F879B174F71F6" type="email">dieter.weyer@t-online.de</mods:nameIdentifier>
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<mods:typeOfResource id="113478347113FF3594FF5588838C7469">text</mods:typeOfResource>
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<mods:title id="B84D5D739EA681DB2F442F7797219C02">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="6ED70CD8EFD899CF03752DF760BA02CD">
<mods:date id="6E0D2DA9C44AF8BB03F61DDB423FD2C5">2023</mods:date>
<mods:detail id="E2D07E67FA55EC98BFD44A8E8541672B" type="pubDate">
<mods:number id="7641DFDD9F9D2B0B1C25319D661B6A6B">2023-07-19</mods:number>
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<mods:detail id="DDA1E04CCC3DFF685F318969C27717ED" type="volume">
<mods:number id="45FF32E6B740A8063BCFA41963B1842D">882</mods:number>
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<mods:start id="FDE4A9675C9BB1EAF6596FBF838805C9">1</mods:start>
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<mods:url id="5DD5D07C351A9E87BA1100C0B2ADC658">http://dx.doi.org/10.5852/ejt.2023.882.2177</mods:url>
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<mods:classification id="64940B84CC928FE4C87CE1301F5004B1">journal article</mods:classification>
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<mods:identifier id="76DBD751B9144042A58E911F6367B8E2" type="DOI">10.5852/ejt.2023.882.2177</mods:identifier>
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<treatment id="03EA5C14CAB885FCFDAFF939FC4486C9" ID-DOI="http://doi.org/10.5281/zenodo.8184524" ID-GBIF-Taxon="212420449" ID-Zenodo-Dep="8184524" LSID="urn:lsid:zoobank.org:act:E347E068-0C6F-4263-BE30-7540C182E55C" httpUri="http://treatment.plazi.org/id/03EA5C14CAB885FCFDAFF939FC4486C9" lastPageId="179" lastPageNumber="180" pageId="177" pageNumber="178">
<subSubSection id="C359BE89CAB885FEFDAFF939FB098285" pageId="177" pageNumber="178" type="nomenclature">
<paragraph id="8BFCED02CAB885FEFDAFF939FC6182A9" blockId="177.[376,1211,1705,1803]" box="[609,979,1705,1732]" pageId="177" pageNumber="178">
<heading id="D0B45A6ECAB885FEFDAFF939FC6182A9" box="[609,979,1705,1732]" centered="true" fontSize="11" level="2" pageId="177" pageNumber="178" reason="2">
<taxonomicName id="4C439681CAB885FEFDAFF939FCC382A9" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[609,881,1705,1732]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="177" pageNumber="178" phylum="Mollusca" rank="species" species="kullmanni" status="sp. nov.">
<emphasis id="B9373110CAB885FEFDAFF939FCC382A9" bold="true" box="[609,881,1705,1732]" italics="true" pageId="177" pageNumber="178">Paprothites kullmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCAB885FEFCB6F93AFC6182A9" box="[888,979,1706,1732]" pageId="177" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BFCED02CAB885FEFEB6F95DFB098285" blockId="177.[376,1211,1705,1803]" box="[376,1211,1741,1768]" pageId="177" pageNumber="178">
<uri id="FFD2E100CAB885FEFEB6F95DFB098285" box="[376,1211,1741,1768]" pageId="177" pageNumber="178">
urn:lsid:zoobank.org:act:
<uuid id="FFE5D7D7CAB885FEFD59F95DFB098285" box="[663,1211,1741,1768]" pageId="177" pageNumber="178">E347E068-0C6F-4263-BE30-7540C182E55C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CAB885FEFD69F960FC3E8366" box="[679,908,1776,1803]" pageId="177" pageNumber="178" type="description">
<paragraph id="8BFCED02CAB885FEFD69F960FC3E8366" blockId="177.[376,1211,1705,1803]" box="[679,908,1776,1803]" pageId="177" pageNumber="178">
<figureCitation id="1378F187CAB885FEFD69F960FCBE8367" box="[679,780,1776,1802]" captionStart="Fig" captionStartId="178.[189,232,1928,1954]" captionTargetBox="[210,1365,1486,1882]" captionTargetId="figure-325@178.[209,1378,1478,1893]" captionTargetPageId="178" captionText="Fig. 109. Paprothites kullmanni sp. nov. A. Holotype MB.C.5270 (Kullmann 1968 Coll.) from Oese. B. Paratype MB.C.31211 (Weyer 1993–1994 Coll.) from the Oberrödinghausen railway cutting, bed 3e. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8177809" httpUri="https://zenodo.org/record/8177809/files/figure.png" pageId="177" pageNumber="178">Fig. 109</figureCitation>
;
<tableCitation id="C6C1D8B9CAB885FEFCD6F960FC3E8366" box="[792,908,1776,1803]" captionStart="Table 105" captionStartId="179.[189,258,265,291]" captionTargetPageId="179" captionText="Table 105. Conch measurements, ratios and rates of the holotype of Paprothites kullmanni sp. nov." pageId="177" pageNumber="178">Table 105</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CAB885FEFF73F8A1FCDC8389" pageId="177" pageNumber="178" type="diagnosis">
<paragraph id="8BFCED02CAB885FEFF73F8A1FE848326" blockId="177.[189,310,1841,1867]" box="[189,310,1841,1867]" pageId="177" pageNumber="178">
<heading id="D0B45A6ECAB885FEFF73F8A1FE848326" bold="true" box="[189,310,1841,1867]" fontSize="11" level="3" pageId="177" pageNumber="178" reason="3">
<emphasis id="B9373110CAB885FEFF73F8A1FE848326" bold="true" box="[189,310,1841,1867]" pageId="177" pageNumber="178">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CAB885FEFF73F8F0FCDC8389" blockId="177.[188,1399,1888,2020]" pageId="177" pageNumber="178">
Species of
<taxonomicName id="4C439681CAB885FEFEF4F8F0FE708317" authorityName="Bartzsch &amp; Weyer" authorityYear="1987" box="[314,450,1888,1914]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="177" pageNumber="178" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CAB885FEFEF4F8F0FE708317" box="[314,450,1888,1914]" italics="true" pageId="177" pageNumber="178">Paprothites</emphasis>
</taxonomicName>
with a conch reaching
<quantity id="4CBB40E7CAB885FEFD01F8F0FC928316" box="[719,800,1888,1915]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="177" pageNumber="178" unit="mm" value="30.0">30 mm</quantity>
diameter. Conch at
<quantity id="4CBB40E7CAB885FEFBC9F8F0FBEA8316" box="[1031,1112,1888,1915]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="177" pageNumber="178" unit="mm" value="25.0">25 mm</quantity>
dm extremely discoidal, evolute (ww/dm ~0.33; uw/dm ~0.50). Whorl profile at
<quantity id="4CBB40E7CAB885FEFC9BF813FC1883F3" box="[853,938,1923,1950]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="177" pageNumber="178" unit="mm" value="25.0">25 mm</quantity>
dm weakly depressed (ww/wh ~1.25); coiling rate low (WER ~1.60). Flanks and venter flattened, umbilical margin rounded. Sculpture on the flank with 40 sharp ribs with concave course on the flanks.
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CABB85FDFF73FE99FAC5853C" box="[189,1399,265,338]" pageId="178" pageNumber="179" type="etymology">
<paragraph id="8BFCED02CABB85FDFF73FE99FEF6854E" blockId="178.[189,324,265,291]" box="[189,324,265,291]" pageId="178" pageNumber="179">
<heading id="D0B45A6ECABB85FDFF73FE99FEF6854E" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="178" pageNumber="179" reason="3">
<emphasis id="B9373110CABB85FDFF73FE99FEF6854E" bold="true" box="[189,324,265,291]" pageId="178" pageNumber="179">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CABB85FDFF73FEA7FAC5853C" blockId="178.[189,1399,311,338]" box="[189,1399,311,338]" pageId="178" pageNumber="179">Named after Jürgen Kullmann (1931–2018) in appreciation of his studies on Carboniferous ammonoids.</paragraph>
</subSubSection>
<subSubSection id="C359BE89CABB85FDFF73FEE8FEEF86BB" pageId="178" pageNumber="179" type="materials_examined">
<paragraph id="8BFCED02CABB85FDFF73FEE8FE1885FF" blockId="178.[189,426,376,402]" box="[189,426,376,402]" pageId="178" pageNumber="179">
<heading id="D0B45A6ECABB85FDFF73FEE8FE1885FF" bold="true" box="[189,426,376,402]" fontSize="11" level="3" pageId="178" pageNumber="179" reason="3">
<emphasis id="B9373110CABB85FDFF73FEE8FE1885FF" bold="true" box="[189,426,376,402]" pageId="178" pageNumber="179">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CABB85FDFF24FE37FEEE85AC" blockId="178.[189,1399,423,520]" box="[234,348,423,449]" pageId="178" pageNumber="179">
<emphasis id="B9373110CABB85FDFF24FE37FEEE85AC" bold="true" box="[234,348,423,449]" pageId="178" pageNumber="179">
<typeStatus id="54F853A0CABB85FDFF24FE37FEEE85AC" box="[234,348,423,449]" pageId="178" pageNumber="179" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFCED02CABB85FDFF73FE5AFDC28665" blockId="178.[189,1399,423,520]" pageId="178" pageNumber="179">
<materialsCitation id="3B2BE75FCABB85FDFF73FE5AFDC28665" ID-GBIF-Occurrence="4165071313" country="Germany" location="Rhenish Mountains" pageId="178" pageNumber="179" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F354AD92CABB85FDFF73FE5AFEE38589" box="[189,337,458,484]" name="Germany" pageId="178" pageNumber="179">GERMANY</collectingCountry>
•
<location id="8E9CBBD9CABB85FDFEBEFE5AFDEE8589" LSID="urn:lsid:plazi:treatment:03EA5C14CAB885FCFDAFF939FC4486C9:8E9CBBD9CABB85FDFEBEFE5AFDEE8589" box="[368,604,458,484]" country="Germany" name="Rhenish Mountains" pageId="178" pageNumber="179">Rhenish Mountains</location>
, Oese, old quarry; Hangenberg Limestone; Kullmann 1968 Coll.; illustrated in
<figureCitation id="1378F187CABB85FDFE96FE7DFE7D8665" box="[344,463,493,520]" captionStart="Fig" captionStartId="178.[189,232,1928,1954]" captionTargetBox="[210,1365,1486,1882]" captionTargetId="figure-325@178.[209,1378,1478,1893]" captionTargetPageId="178" captionText="Fig. 109. Paprothites kullmanni sp. nov. A. Holotype MB.C.5270 (Kullmann 1968 Coll.) from Oese. B. Paratype MB.C.31211 (Weyer 1993–1994 Coll.) from the Oberrödinghausen railway cutting, bed 3e. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8177809" httpUri="https://zenodo.org/record/8177809/files/figure.png" pageId="178" pageNumber="179">Fig. 109A</figureCitation>
; MB.C.5270.
</materialsCitation>
</paragraph>
<paragraph id="8BFCED02CABB85FDFF24FDBFFED58624" blockId="178.[189,1399,559,726]" box="[234,359,559,585]" pageId="178" pageNumber="179">
<emphasis id="B9373110CABB85FDFF24FDBFFED58624" bold="true" box="[234,359,559,585]" pageId="178" pageNumber="179">
<typeStatus id="54F853A0CABB85FDFF24FDBFFED58624" box="[234,359,559,585]" pageId="178" pageNumber="179" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFCED02CABB85FDFF73FDC2FEEF86BB" blockId="178.[189,1399,559,726]" pageId="178" pageNumber="179">
<materialsCitation id="3B2BE75FCABB85FDFF73FDC2FC7786E2" ID-GBIF-Occurrence="4165071560" country="Germany" county="Weyer" location="Rhenish Mountains" municipality="Hangenberg Limestone" pageId="178" pageNumber="179" specimenCount="2" typeStatus="paratype">
<collectingCountry id="F354AD92CABB85FDFF73FDC2FEE38601" box="[189,337,594,620]" name="Germany" pageId="178" pageNumber="179">GERMANY</collectingCountry>
•
<specimenCount id="9D45268BCABB85FDFEB6FDC2FDA68601" box="[376,532,594,620]" pageId="178" pageNumber="179" type="generic">2 specimens</specimenCount>
;
<location id="8E9CBBD9CABB85FDFDE9FDC2FCA58601" LSID="urn:lsid:plazi:treatment:03EA5C14CAB885FCFDAFF939FC4486C9:8E9CBBD9CABB85FDFDE9FDC2FCA58601" box="[551,791,594,620]" country="Germany" county="Weyer" municipality="Hangenberg Limestone" name="Rhenish Mountains" pageId="178" pageNumber="179">Rhenish Mountains</location>
,
<location id="8E9CBBD9CABB85FDFCE4FDC2FBB48601" LSID="urn:lsid:plazi:treatment:03EA5C14CAB885FCFDAFF939FC4486C9:8E9CBBD9CABB85FDFCE4FDC2FBB48601" box="[810,1030,594,620]" country="Germany" county="Weyer" municipality="Hangenberg Limestone" name="Oberrodinghausen" pageId="178" pageNumber="179">Oberrödinghausen</location>
, railway cutting;
<collectingMunicipality id="6B987778CABB85FDFB26FDC2FE8E86E2" pageId="178" pageNumber="179">Hangenberg Limestone</collectingMunicipality>
, bed 3d2;
<collectingCounty id="629D958ECABB85FDFE0CFDE6FDBC86FD" box="[450,526,630,656]" pageId="178" pageNumber="179">Weyer</collectingCounty>
1993–1994 Coll.; MB.C.31210.1–2
</materialsCitation>
•
<materialsCitation id="3B2BE75FCABB85FDFC26FDE5FEEB86BB" ID-GBIF-Occurrence="4165071654" country="Germany" county="Weyer" location="Rhenish Mountains" municipality="Hangenberg Limestone" pageId="178" pageNumber="179" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D45268BCABB85FDFC26FDE5FBC786E2" box="[1000,1141,629,655]" pageId="178" pageNumber="179" type="generic">1 specimen</specimenCount>
;
<location id="8E9CBBD9CABB85FDFB4BFDE5FAC186E2" LSID="urn:lsid:plazi:treatment:03EA5C14CAB885FCFDAFF939FC4486C9:8E9CBBD9CABB85FDFB4BFDE5FAC186E2" box="[1157,1395,629,655]" country="Germany" county="Weyer" municipality="Hangenberg Limestone" name="Rhenish Mountains" pageId="178" pageNumber="179">Rhenish Mountains</location>
,
<location id="8E9CBBD9CABB85FDFF73FD08FE2B86DF" LSID="urn:lsid:plazi:treatment:03EA5C14CAB885FCFDAFF939FC4486C9:8E9CBBD9CABB85FDFF73FD08FE2B86DF" box="[189,409,664,690]" country="Germany" county="Weyer" municipality="Hangenberg Limestone" name="Oberrodinghausen" pageId="178" pageNumber="179">Oberrödinghausen</location>
, railway cutting;
<collectingMunicipality id="6B987778CABB85FDFD45FD08FC1F86DF" box="[651,941,664,690]" pageId="178" pageNumber="179">Hangenberg Limestone</collectingMunicipality>
, bed 3e;
<collectingCounty id="629D958ECABB85FDFBF2FD09FB3A86DE" box="[1084,1160,665,691]" pageId="178" pageNumber="179">Weyer</collectingCounty>
1993–1994 Coll.; MB.C.31211
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CABB85FDFF73FD6DFC8D8121" pageId="178" pageNumber="179" type="description">
<paragraph id="8BFCED02CABB85FDFF73FD6DFEFC877A" blockId="178.[189,334,765,791]" box="[189,334,765,791]" pageId="178" pageNumber="179">
<heading id="D0B45A6ECABB85FDFF73FD6DFEFC877A" bold="true" box="[189,334,765,791]" fontSize="11" level="3" pageId="178" pageNumber="179" reason="3">
<emphasis id="B9373110CABB85FDFF73FD6DFEFC877A" bold="true" box="[189,334,765,791]" pageId="178" pageNumber="179">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CABB85FDFF73FCBCFDD98074" blockId="178.[188,1400,811,1049]" pageId="178" pageNumber="179">
<typeStatus id="54F853A0CABB85FDFF73FCBCFE98872B" box="[189,298,812,838]" pageId="178" pageNumber="179" type="holotype">Holotype</typeStatus>
MB.C.5270 (
<figureCitation id="1378F187CABB85FDFE01FCBCFDF9872B" box="[463,587,812,838]" captionStart="Fig" captionStartId="178.[189,232,1928,1954]" captionTargetBox="[210,1365,1486,1882]" captionTargetId="figure-325@178.[209,1378,1478,1893]" captionTargetPageId="178" captionText="Fig. 109. Paprothites kullmanni sp. nov. A. Holotype MB.C.5270 (Kullmann 1968 Coll.) from Oese. B. Paratype MB.C.31211 (Weyer 1993–1994 Coll.) from the Oberrödinghausen railway cutting, bed 3e. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8177809" httpUri="https://zenodo.org/record/8177809/files/figure.png" pageId="178" pageNumber="179">Fig. 109A</figureCitation>
) is a specimen with
<quantity id="4CBB40E7CABB85FDFC8DFCBBFC25872B" box="[835,919,811,838]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="178" pageNumber="179" unit="mm" value="26.0">26 mm</quantity>
conch diameter, which allows the study of the conch geometry, the ornament and parts of the suture line. The conch is extremely discoidal with a wide umbilicus (ww/dm = 0.33; uw/dm = 0.52) and a low coiling rate (WER = 1.60). The whorl profile is weakly depressed (ww/wh = 1.23) and shows flattened, weakly convergent flanks, which are separated from the also flattened venter by a tightly rounded ventrolateral shoulder. On the penultimate half whorl, the venter shows a shallow longitudinal groove that is not located on the centre of the venter, but asymmetrically on the right side.
</paragraph>
<paragraph id="8BFCED02CABB85FDFF73FBD0FC2780A9" blockId="178.[188,1399,1088,1220]" pageId="178" pageNumber="179">The sculpture consists of sharp ribs confined to the flanks, which are highest on the middle of the flank and emerge as small nodes in the ventrolateral area. There are 24 of these ribs on the last half whorl; the penultimate half whorl has 16 ribs. They run across the flank with a concave arc directed forward. The suture line shows a lanceolate, narrow and deep external lobe.</paragraph>
<paragraph id="8BFCED02CABB85FDFF73FB7CFC8D8121" blockId="178.[189,1400,1259,1356]" pageId="178" pageNumber="179">
<typeStatus id="54F853A0CABB85FDFF73FB7CFE96816B" box="[189,292,1260,1286]" pageId="178" pageNumber="179" type="paratype">Paratype</typeStatus>
MB.C.31211 (109B) is a fragment of a specimen of about
<quantity id="4CBB40E7CABB85FDFC2EFB7BFB86816B" box="[992,1076,1259,1286]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="178" pageNumber="179" unit="mm" value="25.0">25 mm</quantity>
conch diameter. It shows a modification of the sculpture in that the ribs extend beyond the venter and form chevron-shaped sinuses. In the other aspects it largely agrees with the
<typeStatus id="54F853A0CABB85FDFD1EFAA2FC898121" box="[720,827,1330,1356]" pageId="178" pageNumber="179" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF3CBD8ACABB85FDFF73F818FE7F8384" ID-DOI="http://doi.org/10.5281/zenodo.8177809" ID-Zenodo-Dep="8177809" httpUri="https://zenodo.org/record/8177809/files/figure.png" pageId="178" pageNumber="179" startId="178.[189,232,1928,1954]" targetBox="[210,1365,1486,1882]" targetPageId="178">
<paragraph id="8BFCED02CABB85FDFF73F818FE7F8384" blockId="178.[188,1399,1928,2025]" pageId="178" pageNumber="179">
<emphasis id="B9373110CABB85FDFF73F818FE9883CF" bold="true" box="[189,298,1928,1954]" pageId="178" pageNumber="179">Fig. 109.</emphasis>
<taxonomicName id="4C439681CABB85FDFEFDF818FD8883CF" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[307,570,1928,1954]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="178" pageNumber="179" phylum="Mollusca" rank="species" species="kullmanni" status="sp. nov.">
<emphasis id="B9373110CABB85FDFEFDF818FD8883CF" box="[307,570,1928,1954]" italics="true" pageId="178" pageNumber="179">Paprothites kullmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCABB85FDFD8DF819FD1283CE" box="[579,672,1929,1955]" pageId="178" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9373110CABB85FDFD66F818FD0F83CF" bold="true" box="[680,701,1928,1954]" pageId="178" pageNumber="179">A</emphasis>
. Holotype MB.C.5270 (Kullmann 1968 Coll.) from Oese.
<emphasis id="B9373110CABB85FDFF71F83BFF6283A8" bold="true" box="[191,208,1963,1989]" pageId="178" pageNumber="179">B</emphasis>
. Paratype MB.C.31211 (Weyer 1993–1994 Coll.) from the Oberrödinghausen railway cutting, bed 3e. Scale bar units = 1 mm.
</paragraph>
</caption>
<caption id="DF3CBD8ACABA85FCFF73FE99FAF48549" box="[189,1350,265,292]" pageId="179" pageNumber="180" startId="179.[189,258,265,291]" targetBox="[197,1374,344,445]" targetIsTable="true" targetPageId="179">
<paragraph id="8BFCED02CABA85FCFF73FE99FAF48549" blockId="179.[189,1350,265,292]" box="[189,1350,265,292]" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFF73FE99FE8F854E" bold="true" box="[189,317,265,291]" pageId="179" pageNumber="180">Table 105.</emphasis>
Conch measurements, ratios and rates of the holotype of
<taxonomicName id="4C439681CABA85FCFC2EFE99FB56854E" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[992,1252,265,291]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="kullmanni" status="sp. nov.">
<emphasis id="B9373110CABA85FCFC2EFE99FB56854E" box="[992,1252,265,291]" italics="true" pageId="179" pageNumber="180">Paprothites kullmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCABA85FCFB22FE9AFAF48549" box="[1260,1350,266,292]" pageId="179" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BFCED02CABA85FCFF28FEC8FAFF85D0" pageId="179" pageNumber="180">
<table id="F9431FA2CABA7AB0FF0BFEC8FAEC85D0" box="[197,1374,344,445]" colsContinueFrom="177.[197,1376,344,445]" colsContinueIn="187.[197,1372,379,481]" gridcols="11" gridrows="3" pageId="179" pageNumber="180">
<tr id="3573EF40CABA7AB0FF0BFEC8FAEC851D" box="[197,1374,344,368]" gridrow="0" pageId="179" pageNumber="180">
<th id="76A2863CCABA7AB0FF0BFEC8FEFD851D" box="[197,335,344,368]" gridcol="0" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFF28FEC8FEFD851D" bold="true" box="[230,335,344,368]" pageId="179" pageNumber="180">specimen</emphasis>
</th>
<th id="76A2863CCABA7AB0FE42FEC8FE7A851D" box="[396,456,344,368]" gridcol="1" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFE59FEC8FE0E851D" bold="true" box="[407,444,344,368]" pageId="179" pageNumber="180">dm</emphasis>
</th>
<th id="76A2863CCABA7AB0FE3DFEC8FD90851D" box="[499,546,344,368]" gridcol="2" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFE36FEC8FDAD851D" bold="true" box="[504,543,344,368]" pageId="179" pageNumber="180">ww</emphasis>
</th>
<th id="76A2863CCABA7AB0FD9FFEC8FD32851D" box="[593,640,344,368]" gridcol="3" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFD99FEC8FDCB851D" bold="true" box="[599,633,344,368]" pageId="179" pageNumber="180">wh</emphasis>
</th>
<th id="76A2863CCABA7AB0FD66FEC8FD56851D" box="[680,740,344,368]" gridcol="4" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFD7BFEC8FD65851D" bold="true" box="[693,727,344,368]" pageId="179" pageNumber="180">uw</emphasis>
</th>
<th id="76A2863CCABA7AB0FCC3FEC8FC8E851D" box="[781,828,344,368]" gridcol="5" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFCD8FEC8FC80851D" bold="true" box="[790,818,344,368]" pageId="179" pageNumber="180">ah</emphasis>
</th>
<th id="76A2863CCABA7AB0FCABFEC8FC0A851D" box="[869,952,344,368]" gridcol="6" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFCABFEC8FC0A851D" bold="true" box="[869,952,344,368]" pageId="179" pageNumber="180">ww/dm</emphasis>
</th>
<th id="76A2863CCABA7AB0FC13FEC8FB9F851D" box="[989,1069,344,368]" gridcol="7" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFC13FEC8FB9F851D" bold="true" box="[989,1069,344,368]" pageId="179" pageNumber="180">ww/wh</emphasis>
</th>
<th id="76A2863CCABA7AB0FB9CFEC8FB12851D" box="[1106,1184,344,368]" gridcol="8" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFB9CFEC8FB12851D" bold="true" box="[1106,1184,344,368]" pageId="179" pageNumber="180">uw/dm</emphasis>
</th>
<th id="76A2863CCABA7AB0FB08FEC8FAB4851D" box="[1222,1286,344,368]" gridcol="9" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFB08FEC8FAB4851D" bold="true" box="[1222,1286,344,368]" pageId="179" pageNumber="180">WER</emphasis>
</th>
<th id="76A2863CCABA7AB0FAE1FEC8FAEC851D" box="[1327,1374,344,368]" gridcol="10" gridrow="0" pageId="179" pageNumber="180">
<emphasis id="B9373110CABA85FCFAE1FEC8FAEC851D" bold="true" box="[1327,1374,344,368]" pageId="179" pageNumber="180">IZR</emphasis>
</th>
</tr>
<tr id="3573EF40CABA7AB0FF0BFE13FAEC85F1" box="[197,1374,387,412]" gridrow="1" pageId="179" pageNumber="180">
<th id="76A2863CCABA7AB0FF0BFE13FEFD85F1" box="[197,335,387,412]" gridcol="0" gridrow="1" pageId="179" pageNumber="180">MB.C.5270</th>
<td id="76A2863CCABA7AB0FE42FE13FE7A85F1" box="[396,456,387,412]" gridcol="1" gridrow="1" pageId="179" pageNumber="180">26.12</td>
<td id="76A2863CCABA7AB0FE3DFE13FD9085F1" box="[499,546,387,412]" gridcol="2" gridrow="1" pageId="179" pageNumber="180">8.52</td>
<td id="76A2863CCABA7AB0FD9FFE13FD3285F1" box="[593,640,387,412]" gridcol="3" gridrow="1" pageId="179" pageNumber="180">6.92</td>
<td id="76A2863CCABA7AB0FD66FE13FD5685F1" box="[680,740,387,412]" gridcol="4" gridrow="1" pageId="179" pageNumber="180">13.53</td>
<td id="76A2863CCABA7AB0FCC3FE13FC8E85F1" box="[781,828,387,412]" gridcol="5" gridrow="1" pageId="179" pageNumber="180">5.49</td>
<td id="76A2863CCABA7AB0FCABFE13FC0A85F1" box="[869,952,387,412]" gridcol="6" gridrow="1" pageId="179" pageNumber="180">0.33</td>
<td id="76A2863CCABA7AB0FC13FE13FB9F85F1" box="[989,1069,387,412]" gridcol="7" gridrow="1" pageId="179" pageNumber="180">1.23</td>
<td id="76A2863CCABA7AB0FB9CFE13FB1285F1" box="[1106,1184,387,412]" gridcol="8" gridrow="1" pageId="179" pageNumber="180">0.52</td>
<td id="76A2863CCABA7AB0FB08FE13FAB485F1" box="[1222,1286,387,412]" gridcol="9" gridrow="1" pageId="179" pageNumber="180">1.60</td>
<td id="76A2863CCABA7AB0FAE1FE13FAEC85F1" box="[1327,1374,387,412]" gridcol="10" gridrow="1" pageId="179" pageNumber="180">1.00</td>
</tr>
<tr id="3573EF40CABA7AB0FF0BFE35FAEC85D0" box="[197,1374,421,445]" gridrow="2" pageId="179" pageNumber="180">
<th id="76A2863CCABA7AB0FF0BFE35FEFD85D0" box="[197,335,421,445]" gridcol="0" gridrow="2" pageId="179" pageNumber="180">MB.C.5270</th>
<td id="76A2863CCABA7AB0FE42FE35FE7A85D0" box="[396,456,421,445]" gridcol="1" gridrow="2" pageId="179" pageNumber="180">21.85</td>
<td id="76A2863CCABA7AB0FE3DFE35FD9085D0" box="[499,546,421,445]" gridcol="2" gridrow="2" pageId="179" pageNumber="180">7.42</td>
<td id="76A2863CCABA7AB0FD9FFE35FD3285D0" box="[593,640,421,445]" gridcol="3" gridrow="2" pageId="179" pageNumber="180">6.04</td>
<td id="76A2863CCABA7AB0FD66FE35FD5685D0" box="[680,740,421,445]" gridcol="4" gridrow="2" pageId="179" pageNumber="180">11.61</td>
<td id="76A2863CCABA7AB0FCC3FE35FC8E85D0" box="[781,828,421,445]" gridcol="5" gridrow="2" pageId="179" pageNumber="180">–</td>
<td id="76A2863CCABA7AB0FCABFE35FC0A85D0" box="[869,952,421,445]" gridcol="6" gridrow="2" pageId="179" pageNumber="180">0.34</td>
<td id="76A2863CCABA7AB0FC13FE35FB9F85D0" box="[989,1069,421,445]" gridcol="7" gridrow="2" pageId="179" pageNumber="180">1.23</td>
<td id="76A2863CCABA7AB0FB9CFE35FB1285D0" box="[1106,1184,421,445]" gridcol="8" gridrow="2" pageId="179" pageNumber="180">0.53</td>
<td id="76A2863CCABA7AB0FB08FE35FAB485D0" box="[1222,1286,421,445]" gridcol="9" gridrow="2" pageId="179" pageNumber="180">–</td>
<td id="76A2863CCABA7AB0FAE1FE35FAEC85D0" box="[1327,1374,421,445]" gridcol="10" gridrow="2" pageId="179" pageNumber="180">–</td>
</tr>
</table>
</paragraph>
<subSubSection id="C359BE89CABA85FCFF73FE7FFC4486C9" pageId="179" pageNumber="180" type="discussion">
<paragraph id="8BFCED02CABA85FCFF73FE7FFE9D8664" blockId="179.[189,303,495,521]" box="[189,303,495,521]" pageId="179" pageNumber="180">
<heading id="D0B45A6ECABA85FCFF73FE7FFE9D8664" bold="true" box="[189,303,495,521]" fontSize="11" level="3" pageId="179" pageNumber="180" reason="3">
<emphasis id="B9373110CABA85FCFF73FE7FFE9D8664" bold="true" box="[189,303,495,521]" pageId="179" pageNumber="180">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CABA85FCFF73FD8EFC4486C9" blockId="179.[188,1399,542,676]" pageId="179" pageNumber="180">
<taxonomicName id="4C439681CABA85FCFF73FD8EFE728655" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[189,448,542,568]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="kullmanni" status="sp. nov.">
<emphasis id="B9373110CABA85FCFF73FD8EFE728655" box="[189,448,542,568]" italics="true" pageId="179" pageNumber="180">Paprothites kullmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCABA85FCFE09FD8FFD938654" box="[455,545,543,569]" pageId="179" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C439681CABA85FCFD77FD8FFCD48655" box="[697,870,542,568]" class="Cephalopoda" family="Pseudarietitidae" genus="Pseudarietites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="dorsoplanus">
<emphasis id="B9373110CABA85FCFD77FD8FFCD48655" box="[697,870,542,568]" italics="true" pageId="179" pageNumber="180">P. dorsoplanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C439681CABA85FCFCBAFD8FFBAC8655" baseAuthorityName="Vohringer" baseAuthorityYear="1960" box="[884,1054,543,568]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="raricostatus">
<emphasis id="B9373110CABA85FCFCBAFD8FFBAC8655" box="[884,1054,543,568]" italics="true" pageId="179" pageNumber="180">P. raricostatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C439681CABA85FCFB9BFD8FFB758655" authority="Korn &amp; Weyer, 2023" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[1109,1223,542,568]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="B9373110CABA85FCFB9BFD8FFB758655" box="[1109,1223,542,568]" italics="true" pageId="179" pageNumber="180">P. beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2048C6BCABA85FCFB01FD8FFA9B8654" box="[1231,1321,543,569]" pageId="179" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
by the very sharp ribs.
<taxonomicName id="4C439681CABA85FCFE4FFDD2FD228631" authorityName="Korn &amp; Weyer" authorityYear="2003" box="[385,656,578,604]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="ruzhencevi">
<emphasis id="B9373110CABA85FCFE4FFDD2FD228631" box="[385,656,578,604]" italics="true" pageId="179" pageNumber="180">Paprothites ruzhencevi</emphasis>
</taxonomicName>
has sharp ribs, but shows a rounded whorl profile without the distinct ventrolateral shoulder present in
<taxonomicName id="4C439681CABA85FCFD53FDF7FC9D86ED" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[669,815,614,640]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="kullmanni">
<emphasis id="B9373110CABA85FCFD53FDF7FC9D86ED" box="[669,815,614,640]" italics="true" pageId="179" pageNumber="180">P. kullmanni</emphasis>
</taxonomicName>
; furthermore,
<taxonomicName id="4C439681CABA85FCFC17FDF7FBC786ED" authorityName="Korn &amp; Weyer" authorityYear="2003" box="[985,1141,614,640]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="ruzhencevi">
<emphasis id="B9373110CABA85FCFC17FDF7FBC786ED" box="[985,1141,614,640]" italics="true" pageId="179" pageNumber="180">P. ruzhencevi</emphasis>
</taxonomicName>
has a more depressed whorl profile (ww/wh ~1.50 or more) than
<taxonomicName id="4C439681CABA85FCFD78FD1AFCF586C9" authorityName="Korn &amp; Weyer" authorityYear="2023" box="[694,839,650,676]" class="Cephalopoda" family="Pseudarietitidae" genus="Paprothites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="179" pageNumber="180" phylum="Mollusca" rank="species" species="kullmanni">
<emphasis id="B9373110CABA85FCFD78FD1AFCF586C9" box="[694,839,650,676]" italics="true" pageId="179" pageNumber="180">P. kullmanni</emphasis>
</taxonomicName>
(w/wh ~1.25).
</paragraph>
</subSubSection>
</treatment>
</document>
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