treatments-xml/data/03/D7/FF/03D7FF1CFFA8FF8EFF48F8E133F5FAEC.xml
2024-06-21 12:22:17 +02:00

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<document id="C94954267AF4A63DF776771225CEE41F" ID-DOI="10.5281/zenodo.3678492" ID-GBIF-Dataset="8e18452b-1d9d-462e-bac8-8c86eeb4ddc3" ID-ISSN="1942-1354" ID-Zenodo-Dep="3678492" ID-ZooBank="930EAB11-37FA-41B0-980A-1A4736527842" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582295086673" checkinUser="felipe" docAuthor="Hielkema, Auke J. &amp; Hielkema, Meindert A." docDate="2019" docId="03D7FF1CFFA8FF8EFF48F8E133F5FAEC" docLanguage="en" docName="InsectaMundi.0732.1-306.pdf.imf" docOrigin="Insecta Mundi 732 (732)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Dendropaemon (Dendropaemon) piceus" docType="treatment" docVersion="12" lastPageNumber="98" masterDocId="FFEE8764FFCBFFEDFF88FB6E317CFFF0" masterDocTitle="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas" masterLastPageNumber="306" masterPageNumber="1" pageNumber="98" updateTime="1698721431862" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="9D68020C2E962B59CE23446462CB8892">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<treatment id="03D7FF1CFFA8FF8EFF48F8E133F5FAEC" ID-DOI="http://doi.org/10.5281/zenodo.3683786" ID-GBIF-Taxon="162246642" ID-Zenodo-Dep="3683786" LSID="urn:lsid:plazi:treatment:03D7FF1CFFA8FF8EFF48F8E133F5FAEC" httpUri="http://treatment.plazi.org/id/03D7FF1CFFA8FF8EFF48F8E133F5FAEC" lastPageNumber="98" pageId="99" pageNumber="98">
<subSubSection id="C3641D81FFA8FF8EFF48F8E132D8FC57" box="[192,932,911,936]" pageId="99" pageNumber="98" type="nomenclature">
<paragraph id="8BC14E0AFFA8FF8EFF48F8E132D8FC57" blockId="99.[192,932,911,936]" box="[192,932,911,936]" pageId="99" pageNumber="98">
<heading id="D089F966FFA8FF8EFF48F8E132D8FC57" bold="true" box="[192,932,911,936]" fontSize="10" level="2" pageId="99" pageNumber="98" reason="3">
<emphasis id="B90A9218FFA8FF8EFF48F8E132D8FC57" bold="true" box="[192,932,911,936]" pageId="99" pageNumber="98">
§
<taxonomicName id="4C7E3589FFA8FF8EFF5DF8E132D8FC57" ID-CoL="7ZN25" authority="(Perty, 1830)" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[213,932,911,936]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceus" subGenus="Dendropaemon">
<emphasis id="B90A9218FFA8FF8EFF5DF8E130D9FC57" bold="true" box="[213,421,911,935]" italics="true" pageId="99" pageNumber="98">Dendropaemon</emphasis>
(
<emphasis id="B90A9218FFA8FF8EFE3FF8E133FBFC57" bold="true" box="[439,647,911,935]" italics="true" pageId="99" pageNumber="98">Dendropaemon</emphasis>
)
<emphasis id="B90A9218FFA8FF8EFD10F8FE3392FC58" bold="true" box="[664,750,912,936]" italics="true" pageId="99" pageNumber="98">piceus</emphasis>
(
<bibRefCitation id="EFEF33FBFFA8FF8EFD77F8E132E7FC57" author="Perty" box="[767,923,911,935]" firstAuthor="Perty" pageId="99" pageNumber="98" pagination="412" refId="ref219520" refString="Perty, J. A. M. 1830. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCX- VII-MDCCCXX. Frid. Fleischer; Leipzig, Germany. 224 p. [Note. Bousquet (2016: 412) discussed the year of publication and concluded that the pages 1 - 60 and the accompanying plates, which include all the referred Scarabaeoidea, were published in 1830]." type="book chapter" year="1830">Perty, 1830</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FFA8FF8EFF60F8D93265FC3C" box="[232,793,950,972]" pageId="99" pageNumber="98" type="reference_group">
<paragraph id="8BC14E0AFFA8FF8EFF60F8D93265FC3C" blockId="99.[232,793,950,972]" box="[232,793,950,972]" pageId="99" pageNumber="98">
<taxonomicName id="4C7E3589FFA8FF8EFF60F8D93342FC3B" authority="Perty 1830" authorityName="Perty" authorityYear="1830" box="[232,574,950,972]" class="Insecta" family="Scarabaeidae" genus="Eurysternus" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B90A9218FFA8FF8EFF60F8D930C7FC3C" box="[232,443,951,972]" italics="true" pageId="99" pageNumber="98">Eurysternus piceus</emphasis>
<bibRefCitation id="EFEF33FBFFA8FF8EFE49F8D93342FC3B" author="Perty" box="[449,574,950,972]" firstAuthor="Perty" pageId="99" pageNumber="98" pagination="412" refId="ref219520" refString="Perty, J. A. M. 1830. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCX- VII-MDCCCXX. Frid. Fleischer; Leipzig, Germany. 224 p. [Note. Bousquet (2016: 412) discussed the year of publication and concluded that the pages 1 - 60 and the accompanying plates, which include all the referred Scarabaeoidea, were published in 1830]." type="book chapter" year="1830">Perty 1830</bibRefCitation>
</taxonomicName>
: pl. 8, fig. 6 (
<collectingCountry id="F3690E9AFFA8FF8EFD42F8D8326DFC3C" box="[714,785,950,972]" name="Brazil" pageId="99" pageNumber="98">Brazil</collectingCountry>
)
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FFA8FF8EFF48F8B233F5FAEC" pageId="99" pageNumber="98" type="distribution">
<paragraph id="8BC14E0AFFA8FF8EFF48F8B233B7FB85" blockId="99.[192,1440,988,1141]" pageId="99" pageNumber="98">
<emphasis id="B90A9218FFA8FF8EFF48F8B2336DFC04" bold="true" box="[192,529,988,1012]" pageId="99" pageNumber="98">Distribution. “Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFFA8FF8EFD95F8B33267FC05" author="Olsoufieff" box="[541,795,989,1013]" firstAuthor="Olsoufieff" pageId="99" pageNumber="98" pagination="4 - 172" refId="ref218570" refString="Olsoufieff, G. d'. 1924. Les phanaeides (Coleoptera-Lamellicornia). Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 172." type="journal article" year="1924">Olsoufieff 1924: 122</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFFA8FF8EFCA3F8B33534FC05" author="Blackwelder, R. E." box="[811,1096,989,1013]" journalOrPublisher="United States National Museum Bulletin" pageId="99" pageNumber="98" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
(as
<taxonomicName id="4C7E3589FFA8FF8EFB0BF8B234ECFC04" box="[1155,1424,988,1012]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceum">
<emphasis id="B90A9218FFA8FF8EFB0BF8B234ECFC04" box="[1155,1424,988,1012]" italics="true" pageId="99" pageNumber="98">Dendropemon piceum</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FFA8FF8EFF48F8923060FBE4" bold="true" box="[192,284,1020,1044]" pageId="99" pageNumber="98">
<collectingCountry id="F3690E9AFFA8FF8EFF48F892306AFBE4" box="[192,278,1020,1044]" name="Brazil" pageId="99" pageNumber="98">Brazil</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFFA8FF8EFEABF89330D0FBE5" author="Perty" box="[291,428,1021,1045]" firstAuthor="Perty" pageId="99" pageNumber="98" pagination="412" refId="ref219520" refString="Perty, J. A. M. 1830. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCX- VII-MDCCCXX. Frid. Fleischer; Leipzig, Germany. 224 p. [Note. Bousquet (2016: 412) discussed the year of publication and concluded that the pages 1 - 60 and the accompanying plates, which include all the referred Scarabaeoidea, were published in 1830]." type="book chapter" year="1830">Perty 1830</bibRefCitation>
: pl. 8, fig. 6, p. 39 (drawing - as
<taxonomicName id="4C7E3589FFA8FF8EFCA7F893356AFBE4" authorityName="Perty" authorityYear="1830" box="[815,1046,1021,1044]" class="Insecta" family="Scarabaeidae" genus="Eurysternus" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B90A9218FFA8FF8EFCA7F893356AFBE4" box="[815,1046,1021,1044]" italics="true" pageId="99" pageNumber="98">Eurysternus piceus</emphasis>
</taxonomicName>
; description - as
<taxonomicName id="4C7E3589FFA8FF8EFB6FF8923070FBC4" baseAuthorityName="Perty" baseAuthorityYear="1830" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B90A9218FFA8FF8EFB6FF8923070FBC4" italics="true" pageId="99" pageNumber="98">Dendropaemon piceus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFFA8FF8EFEADFF733096FBC5" author="Gillet, J. J. E." bookContentInfo="100 p." box="[293,490,1053,1077]" editor="S. Schenkling" journalOrPublisher="Junk; Berlin" pageId="99" pageNumber="98" pagination="3 - 100" refId="ref204620" refString="Gillet, J. J. E. 1911 b. Scarabaeidae: Coprinae I. p. 3 - 100. In: S. Schenkling. Coleopterorum Catalogus Pars 38. W. Junk; Berlin. 100 p." title="Scarabaeidae: Coprinae I" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 38. W" year="1911">Gillet 1911b: 88</bibRefCitation>
(as
<taxonomicName id="4C7E3589FFA8FF8EFDA9FF723263FBC4" box="[545,799,1052,1076]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B90A9218FFA8FF8EFDA9FF723263FBC4" box="[545,799,1052,1076]" italics="true" pageId="99" pageNumber="98">Dendropemon piceus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFFA8FF8EFCB1FF73354EFBC5" author="Olsoufieff" box="[825,1074,1053,1077]" firstAuthor="Olsoufieff" pageId="99" pageNumber="98" pagination="4 - 172" refId="ref218570" refString="Olsoufieff, G. d'. 1924. Les phanaeides (Coleoptera-Lamellicornia). Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 172." type="journal article" year="1924">Olsoufieff 1924: 122</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFFA8FF8EFBC8FF733586FBC5" author="Blut, H." box="[1088,1274,1053,1077]" journalOrPublisher="Archiv fur Naturgeschichte (N. F.)" pageId="99" pageNumber="98" pagination="263 - 300" part="8" refId="ref194178" refString="Blut, H. 1939. Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte (N. F.) 8 (2): 263 - 300." title="Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon" type="journal article" year="1939">Blut 1939: 290</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFFA8FF8EFA80FF73304BFBA5" author="Blackwelder, R. E." journalOrPublisher="United States National Museum Bulletin" pageId="99" pageNumber="98" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
(as
<taxonomicName id="4C7E3589FFA8FF8EFEE5FF52330BFBA4" box="[365,631,1084,1108]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceum">
<emphasis id="B90A9218FFA8FF8EFEE5FF52330BFBA4" box="[365,631,1084,1108]" italics="true" pageId="99" pageNumber="98">Dendropemon piceum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFFA8FF8EFD18FF5332CDFBA5" author="Vaz-de-Mello" box="[656,945,1085,1109]" firstAuthor="Vaz-de-Mello" pageId="99" pageNumber="98" refId="ref228421" refString="Vaz-de-Mello, F. Z. 2000. Estado atual de conhecimento dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. p. 183 - 195. In: F. Martin-Piera, J. J. Morrone, and A. Melic (eds.). Hacia un proyecto CYTED para el inventario y estimacion de la diversidad entomologica en Iberoamerica: PrIBES 2000. Monografias Tercer Milenio; Zaragoza, Spain. 326 p." type="book" year="2000">Vaz-de-Mello 2000: 192</bibRefCitation>
(as
<emphasis id="B90A9218FFA8FF8EFC6FFF5235E5FBA4" box="[999,1177,1084,1108]" italics="true" pageId="99" pageNumber="98">Dendropaenon</emphasis>
[sic]
<taxonomicName id="4C7E3589FFA8FF8EFB55FF53302CFB84" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceum" subGenus="Dendropaenon">
(
<emphasis id="B90A9218FFA8FF8EFB6DFF5234EBFBA4" box="[1253,1431,1084,1108]" italics="true" pageId="99" pageNumber="98">Dendropaenon</emphasis>
) [sic]
<emphasis id="B90A9218FFA8FF8EFF71FF33302CFB84" box="[249,336,1117,1140]" italics="true" pageId="99" pageNumber="98">piceum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFFA8FF8EFEE0FF3333B4FB85" author="Genier, F. &amp; P. Arnaud" box="[360,712,1117,1141]" journalOrPublisher="Zootaxa" pageId="99" pageNumber="98" pagination="1 - 125" part="4099" refId="ref204208" refString="Genier, F., and P. Arnaud. 2016. Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini). Zootaxa 4099 (1): 1 - 125." title="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" type="journal article" year="2016">Génier and Arnaud 2016: 33</bibRefCitation>
.
</paragraph>
<paragraph id="8BC14E0AFFA8FF8EFF48FFEB33F5FAEC" blockId="99.[192,1440,1156,1309]" pageId="99" pageNumber="98">
<emphasis id="B90A9218FFA8FF8EFF48FFEB3074FB6D" bold="true" box="[192,264,1157,1181]" pageId="99" pageNumber="98">Note.</emphasis>
According to
<bibRefCitation id="EFEF33FBFFA8FF8EFE24FFEB326AFB6D" author="Genier, F. &amp; P. Arnaud" box="[428,790,1157,1181]" journalOrPublisher="Zootaxa" pageId="99" pageNumber="98" pagination="1 - 125" part="4099" refId="ref204208" refString="Genier, F., and P. Arnaud. 2016. Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini). Zootaxa 4099 (1): 1 - 125." title="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" type="journal article" year="2016">Génier and Arnaud (2016: 33)</bibRefCitation>
,
<taxonomicName id="4C7E3589FFA8FF8EFCA9FFEA3551FB6C" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[801,1069,1156,1180]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B90A9218FFA8FF8EFCA9FFEA3551FB6C" box="[801,1069,1156,1180]" italics="true" pageId="99" pageNumber="98">Dendropaemon piceus</emphasis>
</taxonomicName>
is only known from the Brazilian
<typeStatus id="54C5F0A8FFA8FF8EFF66FFCB3026FB4D" box="[238,346,1189,1213]" pageId="99" pageNumber="98" type="holotype">holotype</typeStatus>
. According to
<bibRefCitation id="EFEF33FBFFA8FF8EFD8AFFCB33BBFB4D" author="Perty" box="[514,711,1189,1213]" firstAuthor="Perty" pageId="99" pageNumber="98" pagination="412" refId="ref219520" refString="Perty, J. A. M. 1830. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCX- VII-MDCCCXX. Frid. Fleischer; Leipzig, Germany. 224 p. [Note. Bousquet (2016: 412) discussed the year of publication and concluded that the pages 1 - 60 and the accompanying plates, which include all the referred Scarabaeoidea, were published in 1830]." type="book chapter" year="1830">Perty (1830: 39)</bibRefCitation>
, this
<typeStatus id="54C5F0A8FFA8FF8EFC80FFCB320CFB4D" box="[776,880,1189,1213]" pageId="99" pageNumber="98" type="holotype">holotype</typeStatus>
comes from either of the states
<collectingRegion id="49BA80E8FFA8FF8EFB79FFCB3417FB4D" box="[1265,1387,1189,1213]" country="Brazil" name="Sao Paulo" pageId="99" pageNumber="98">São Paulo</collectingRegion>
and
<collectingRegion id="49BA80E8FFA8FF8EFF48FFAB3018FB2D" box="[192,356,1221,1245]" country="Brazil" name="Minas Gerais" pageId="99" pageNumber="98">Minas Gerais</collectingRegion>
. According to
<bibRefCitation id="EFEF33FBFFA8FF8EFD8EFFAB33BAFB2D" author="Blut, H." box="[518,710,1221,1245]" journalOrPublisher="Archiv fur Naturgeschichte (N. F.)" pageId="99" pageNumber="98" pagination="263 - 300" part="8" refId="ref194178" refString="Blut, H. 1939. Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte (N. F.) 8 (2): 263 - 300." title="Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon" type="journal article" year="1939">Blut (1939: 290)</bibRefCitation>
, reports of this species from the Guianas (such as in
<bibRefCitation id="EFEF33FBFFA8FF8EFAA4FFAB303BFB0D" author="Olsoufieff" firstAuthor="Olsoufieff" pageId="99" pageNumber="98" pagination="4 - 172" refId="ref218570" refString="Olsoufieff, G. d'. 1924. Les phanaeides (Coleoptera-Lamellicornia). Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 172." type="journal article" year="1924">Olsoufieff (1924: 122)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFFA8FF8EFE0BFF8B33CEFB0D" author="Blackwelder, R. E." box="[387,690,1253,1277]" journalOrPublisher="United States National Museum Bulletin" pageId="99" pageNumber="98" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder (1944: 211))</bibRefCitation>
may in fact concern
<taxonomicName id="4C7E3589FFA8FF8EFC25FF8B357CFB0C" authorityName="Laporte" authorityYear="1832" box="[941,1024,1253,1276]" class="Insecta" family="Scarabaeidae" genus="Enicotarsus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B90A9218FFA8FF8EFC25FF8B357CFB0C" box="[941,1024,1253,1276]" italics="true" pageId="99" pageNumber="98">D. ater</emphasis>
</taxonomicName>
. We regard
<taxonomicName id="4C7E3589FFA8FF8EFB1CFF8A34DCFB0C" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[1172,1440,1252,1276]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="98" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B90A9218FFA8FF8EFB1CFF8A34DCFB0C" box="[1172,1440,1252,1276]" italics="true" pageId="99" pageNumber="98">Dendropaemon piceus</emphasis>
</taxonomicName>
as not occurring in the research area.
</paragraph>
</subSubSection>
</treatment>
</document>