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<mods:title id="CF248E9D701394346A99ECD36CFCCC09">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<heading id="D089F966FF6CFF4AFF48FA2733EAFE91" bold="true" box="[192,662,329,353]" fontSize="10" level="2" pageId="167" pageNumber="166" reason="3">
<taxonomicName id="4C7E3589FF6CFF4AFF48FA2733EAFE91" authority="Ratcliffe, 2008" authorityName="Ratcliffe" authorityYear="2008" box="[192,662,329,353]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="B90A9218FF6CFF4AFF48FA2733EAFE91" bold="true" box="[192,662,329,353]" pageId="167" pageNumber="166">
<emphasis id="B90A9218FF6CFF4AFF48FA2730B0FE91" bold="true" box="[192,460,329,353]" italics="true" pageId="167" pageNumber="166">Cyclocephala chera</emphasis>
<bibRefCitation id="EFEF33FBFF6CFF4AFE5CFA2733EAFE91" author="Ratcliffe" box="[468,662,329,353]" firstAuthor="Ratcliffe" pageId="167" pageNumber="166" pagination="221 - 241" refId="ref221746" refString="Ratcliffe, B. C. 2008. More new species of Cyclocephala Dejean, 1821 from South America (Scarabaeidae: Dynastidae: Cyclocephalini). The Coleopterists Bulletin 62 (2): 221 - 241." type="journal article" year="2008">Ratcliffe, 2008</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8BC14E0AFF6CFF4AFF60FA1E327FFE76" blockId="167.[232,771,368,390]" box="[232,771,368,390]" pageId="167" pageNumber="166">
<treatmentCitationGroup id="AB6E6924FF6CFF4AFF60FA1E327FFE76" box="[232,771,368,390]" pageId="167" pageNumber="166">
<taxonomicName id="4C7E3589FF6CFF4AFF60FA1E327FFE76" authority="Ratcliffe 2008: 226" authorityName="Ratcliffe" authorityPageNumber="226" authorityYear="2008" box="[232,771,368,390]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="chera">
<treatmentCitation id="0ADF681BFF6CFF4AFF60FA1E33EAFE76" author="Ratcliffe" box="[232,662,368,390]" page="226" pageId="167" pageNumber="166" year="2008">
<emphasis id="B90A9218FF6CFF4AFF60FA1E30C0FE75" box="[232,444,368,389]" italics="true" pageId="167" pageNumber="166">Cyclocephala chera</emphasis>
<bibRefCitation id="EFEF33FBFF6CFF4AFE4BFA1E33EAFE76" author="Ratcliffe" box="[451,662,368,390]" firstAuthor="Ratcliffe" pageId="167" pageNumber="166" pagination="221 - 241" refId="ref221746" refString="Ratcliffe, B. C. 2008. More new species of Cyclocephala Dejean, 1821 from South America (Scarabaeidae: Dynastidae: Cyclocephalini). The Coleopterists Bulletin 62 (2): 221 - 241." type="journal article" year="2008">Ratcliffe 2008: 226</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF6CFF4AFD2CFA1E3380FE76" box="[676,764,368,390]" name="Guyana" pageId="167" pageNumber="166">Guyana</collectingCountry>
)
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF6CFF4AFF48FAF8328AFD06" pageId="167" pageNumber="166" type="distribution">
<paragraph id="8BC14E0AFF6CFF4AFF48FAF830E6FE1E" blockId="167.[192,1440,406,494]" pageId="167" pageNumber="166">
<emphasis id="B90A9218FF6CFF4AFF48FAF83095FE5E" bold="true" box="[192,489,406,430]" pageId="167" pageNumber="166">
Distribution.
<collectingCountry id="F3690E9AFF6CFF4AFEFEFAF8309EFE5E" box="[374,482,406,430]" name="Guyana" pageId="167" pageNumber="166">Guyana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF6CFF4AFE66FAF833ACFE5E" author="Ratcliffe" box="[494,720,406,430]" firstAuthor="Ratcliffe" pageId="167" pageNumber="166" pagination="221 - 241" refId="ref221746" refString="Ratcliffe, B. C. 2008. More new species of Cyclocephala Dejean, 1821 from South America (Scarabaeidae: Dynastidae: Cyclocephalini). The Coleopterists Bulletin 62 (2): 221 - 241." type="journal article" year="2008">Ratcliffe 2008: 226</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF6CFF4AFD51FAF9328CFE5E" author="Moore" box="[729,1008,406,430]" etAl="et al." firstAuthor="Moore" pageId="167" pageNumber="166" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 160</bibRefCitation>
.
<emphasis id="B90A9218FF6CFF4AFC71FAF835FBFE5E" bold="true" box="[1017,1159,406,430]" pageId="167" pageNumber="166">
<collectingCountry id="F3690E9AFF6CFF4AFC71FAF835FCFE5E" box="[1017,1152,406,430]" name="Suriname" pageId="167" pageNumber="166">Suriname</collectingCountry>
:
</emphasis>
MHCN -
<emphasis id="B90A9218FF6CFF4AFB7FFAF830F8FE3E" bold="true" pageId="167" pageNumber="166">NEW COUN- TRY RECORD</emphasis>
(Suriname,
<collectingRegion id="49BA80E8FF6CFF4AFD95FAD93326FE3E" box="[541,602,439,462]" country="Suriname" name="Para" pageId="167" pageNumber="166">Para</collectingRegion>
, Colakreek, (
<geoCoordinate id="EE4A28CDFF6CFF4AFD74FADB3205FE3E" box="[764,889,437,462]" degrees="5" direction="north" minutes="27" orientation="latitude" pageId="167" pageNumber="166" precision="15" seconds="45" value="5.4624996">5°2745″N</geoCoordinate>
,
<geoCoordinate id="EE4A28CDFF6CFF4AFC0DFADB356AFE3E" box="[901,1046,437,462]" degrees="55" direction="west" minutes="13" orientation="longitude" pageId="167" pageNumber="166" precision="15" seconds="45" value="-55.229168">55°1345″W</geoCoordinate>
). 6/
<date id="FFC068CAFF6CFF4AFBC9FAD835D5FE3E" box="[1089,1193,438,462]" pageId="167" pageNumber="166" value="2013-01-09">9-I-2013</date>
. A. J. Hielkema. On UV/HPL.); NZCS.
</paragraph>
<paragraph id="8BC14E0AFF6CFF4AFF48FA90328AFD06" blockId="167.[192,1440,510,758]" pageId="167" pageNumber="166">
<emphasis id="B90A9218FF6CFF4AFF48FA903074FDE6" bold="true" box="[192,264,510,534]" pageId="167" pageNumber="166">Note.</emphasis>
<bibRefCitation id="EFEF33FBFF6CFF4AFE86FA90337DFDE6" author="Ratcliffe" box="[270,513,510,534]" firstAuthor="Ratcliffe" pageId="167" pageNumber="166" pagination="221 - 241" refId="ref221746" refString="Ratcliffe, B. C. 2008. More new species of Cyclocephala Dejean, 1821 from South America (Scarabaeidae: Dynastidae: Cyclocephalini). The Coleopterists Bulletin 62 (2): 221 - 241." type="journal article" year="2008">Ratcliffe (2008: 226)</bibRefCitation>
describes
<taxonomicName id="4C7E3589FF6CFF4AFDF5FA90321AFDE6" authorityName="Ratcliffe" authorityYear="2008" box="[637,870,510,534]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="B90A9218FF6CFF4AFDF5FA90321AFDE6" box="[637,870,510,534]" italics="true" pageId="167" pageNumber="166">Cyclocephala chera</emphasis>
</taxonomicName>
based on
<specimenCount id="9D788583FF6CFF4AFC52FA903533FDE6" box="[986,1103,510,534]" pageId="167" pageNumber="166" type="female">10 female</specimenCount>
specimens from
<collectingCountry id="F3690E9AFF6CFF4AFA9FFA903407FDE6" box="[1303,1403,510,534]" name="Guyana" pageId="167" pageNumber="166">Guyana</collectingCountry>
. In this description, Ratcliffe compares
<taxonomicName id="4C7E3589FF6CFF4AFDFBF97033ABFDC6" authorityName="Ratcliffe" authorityYear="2008" box="[627,727,542,566]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="B90A9218FF6CFF4AFDFBF97033ABFDC6" box="[627,727,542,566]" italics="true" pageId="167" pageNumber="166">C. chera</emphasis>
</taxonomicName>
with
<taxonomicName id="4C7E3589FF6CFF4AFC93F97035DBFDC6" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[795,1191,542,566]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="complanata">
<emphasis id="B90A9218FF6CFF4AFC93F97032B0FDC6" box="[795,972,542,566]" italics="true" pageId="167" pageNumber="166">C. complanata</emphasis>
<bibRefCitation id="EFEF33FBFF6CFF4AFC5AF97035DBFDC6" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[978,1191,542,566]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="167" pageNumber="166" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister, 1847</bibRefCitation>
</taxonomicName>
(p. 48),
<taxonomicName id="4C7E3589FF6CFF4AFA8FF9703033FDA6" authority="Sharp, 1877" authorityName="Sharp" authorityYear="1877" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="conspicua">
<emphasis id="B90A9218FF6CFF4AFA8FF97034DCFDC6" box="[1287,1440,542,566]" italics="true" pageId="167" pageNumber="166">C. conspicua</emphasis>
<bibRefCitation id="EFEF33FBFF6CFF4AFF48F9503033FDA6" author="Sharp" box="[192,335,574,598]" firstAuthor="Sharp" pageId="167" pageNumber="166" pagination="129 - 138" refId="ref224536" refString="Sharp, D. 1877. Description of some new species of beetles (Scarabaeidae) from Central America. Journal of the Linnean Society, Zoology 13: 129 - 138." type="journal article" year="1877">Sharp, 1877</bibRefCitation>
</taxonomicName>
(p. 135) and
<taxonomicName id="4C7E3589FF6CFF4AFE6CF95032AEFDA6" authority="Heyne and Taschenberg, 1908" authorityName="Heyne and Taschenberg" authorityYear="1908" box="[484,978,574,599]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="gregaria">
<emphasis id="B90A9218FF6CFF4AFE6CF9503314FDA6" box="[484,616,574,598]" italics="true" pageId="167" pageNumber="166">C. gregaria</emphasis>
<bibRefCitation id="EFEF33FBFF6CFF4AFDE5F95132AEFDA6" author="Heyne, A. &amp; O. Taschenberg" bookContentInfo="262 p." box="[621,978,574,599]" journalOrPublisher="J. F. Schreiber; Esslingen and Munich, Germany" pageId="167" pageNumber="166" refId="ref207668" refString="Heyne, A., and O. Taschenberg. 1908. Die exotischen Kafer in Wort und Bild. J. F. Schreiber; Esslingen and Munich, Germany. 262 p." title="Die exotischen Kafer in Wort und Bild" type="book" year="1908">Heyne and Taschenberg, 1908</bibRefCitation>
</taxonomicName>
(p. 91), but not with
<taxonomicName id="4C7E3589FF6CFF4AFB4AF9503078FD86" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="undata">
<emphasis id="B90A9218FF6CFF4AFB4AF9503444FDA6" box="[1218,1336,574,598]" italics="true" pageId="167" pageNumber="166">C. undata</emphasis>
(
<bibRefCitation id="EFEF33FBFF6CFF4AFACDF9503180FD86" author="Olivier" firstAuthor="Olivier" pageId="167" pageNumber="166" refId="ref218493" refString="Olivier, G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome premier. Baudouin; Paris. [Note. Each genus is separately paginated]." type="book" year="1789">Olivier, 1789</bibRefCitation>
)
</taxonomicName>
(p. 80). We have collected a number of specimens in
<collectingCountry id="F3690E9AFF6CFF4AFC11F9303569FD86" box="[921,1045,606,630]" name="Suriname" pageId="167" pageNumber="166">Suriname</collectingCountry>
, including several males, which were identified as
<taxonomicName id="4C7E3589FF6CFF4AFE2AF910337BFD66" authorityName="Ratcliffe" authorityYear="2008" box="[418,519,638,662]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="B90A9218FF6CFF4AFE2AF910337BFD66" box="[418,519,638,662]" italics="true" pageId="167" pageNumber="166">C. chera</emphasis>
</taxonomicName>
by Ratcliffe. After viewing photos of these specimens, Ponchel (Y. Ponchel, pers. comm. 2015) considers them synonymous with
<taxonomicName id="4C7E3589FF6CFF4AFCD9F9F032B1FD46" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[849,973,670,694]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="undata">
<emphasis id="B90A9218FF6CFF4AFCD9F9F032B1FD46" box="[849,973,670,694]" italics="true" pageId="167" pageNumber="166">C. undata</emphasis>
</taxonomicName>
, which he says has a bicolored and a unicolored morph. Now that we have found male specimens of
<taxonomicName id="4C7E3589FF6CFF4AFC46F9D03549FD26" authorityName="Ratcliffe" authorityYear="2008" box="[974,1077,702,726]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="B90A9218FF6CFF4AFC46F9D03549FD26" box="[974,1077,702,726]" italics="true" pageId="167" pageNumber="166">C. chera</emphasis>
</taxonomicName>
it seems prudent to compare both sexes of
<taxonomicName id="4C7E3589FF6CFF4AFEECF9B030B5FD06" authorityName="Ratcliffe" authorityYear="2008" box="[356,457,734,758]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="B90A9218FF6CFF4AFEECF9B030B5FD06" box="[356,457,734,758]" italics="true" pageId="167" pageNumber="166">C. chera</emphasis>
</taxonomicName>
with those of both color morphs of
<taxonomicName id="4C7E3589FF6CFF4AFCFDF9B03293FD06" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[885,1007,734,758]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="167" pageNumber="166" phylum="Arthropoda" rank="species" species="undata">
<emphasis id="B90A9218FF6CFF4AFCFDF9B03293FD06" box="[885,1007,734,758]" italics="true" pageId="167" pageNumber="166">C. undata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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