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<mods:title id="A74A21B3912DB3B754464BF82CF038B3">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<treatment id="03D7FF1CFF2AFF0CFF48F91B3502FBDA" ID-DOI="http://doi.org/10.5281/zenodo.3682725" ID-GBIF-Taxon="162246937" ID-Zenodo-Dep="3682725" LSID="urn:lsid:plazi:treatment:03D7FF1CFF2AFF0CFF48F91B3502FBDA" httpUri="http://treatment.plazi.org/id/03D7FF1CFF2AFF0CFF48F91B3502FBDA" lastPageNumber="224" pageId="225" pageNumber="224">
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<paragraph id="8BC14E0AFF2AFF0CFF48F91B33FEFD7D" blockId="225.[192,642,628,653]" box="[192,642,628,653]" pageId="225" pageNumber="224">
<heading id="D089F966FF2AFF0CFF48F91B33FEFD7D" bold="true" box="[192,642,628,653]" fontSize="10" level="2" pageId="225" pageNumber="224" reason="3">
<emphasis id="B90A9218FF2AFF0CFF48F91B33FEFD7D" bold="true" box="[192,642,628,653]" pageId="225" pageNumber="224">
§
<taxonomicName id="4C7E3589FF2AFF0CFF5DF91A33FEFD7D" authority="Prell, 1911" authorityName="Prell" authorityYear="1911" box="[213,642,628,653]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="225" pageNumber="224" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B90A9218FF2AFF0CFF5DF91A3096FD7C" bold="true" box="[213,490,628,652]" italics="true" pageId="225" pageNumber="224">Phileurus excavatus</emphasis>
<bibRefCitation id="EFEF33FBFF2AFF0CFE7AF91B33FEFD7D" author="Prell" box="[498,642,629,653]" firstAuthor="Prell" pageId="225" pageNumber="224" pagination="105 - 107" refId="ref220332" refString="Prell, H. 1911. Beitrage zur Kenntnis der Dynastinen. Entomologische Zeitschrift (Frankfurt) 25 (18): 105 - 107." type="journal article" year="1911">Prell, 1911</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<taxonomicName id="4C7E3589FF2AFF0CFF60F9F533ACFD41" authority="Prell 1911: 106" authorityName="Prell" authorityPageNumber="106" authorityYear="1911" box="[232,720,667,689]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="225" pageNumber="224" phylum="Arthropoda" rank="species" species="excavatus">
<treatmentCitation id="0ADF681BFF2AFF0CFF60F9F53308FD41" author="Prell" box="[232,628,667,689]" page="106" pageId="225" pageNumber="224" year="1911">
<emphasis id="B90A9218FF2AFF0CFF60F9F530BBFD41" box="[232,455,667,689]" italics="true" pageId="225" pageNumber="224">Phileurus excavatus</emphasis>
<bibRefCitation id="EFEF33FBFF2AFF0CFE45F9F53308FD41" author="Prell" box="[461,628,667,689]" firstAuthor="Prell" pageId="225" pageNumber="224" pagination="105 - 107" refId="ref220332" refString="Prell, H. 1911. Beitrage zur Kenntnis der Dynastinen. Entomologische Zeitschrift (Frankfurt) 25 (18): 105 - 107." type="journal article" year="1911">Prell 1911: 106</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF2AFF0CFD0AF9F533B5FD41" box="[642,713,667,689]" name="Brazil" pageId="225" pageNumber="224">Brazil</collectingCountry>
)
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF2AFF0CFF48F9AF3502FBDA" pageId="225" pageNumber="224" type="distribution">
<paragraph id="8BC14E0AFF2AFF0CFF48F9AF35FDFCCA" blockId="225.[192,1441,705,826]" pageId="225" pageNumber="224">
<emphasis id="B90A9218FF2AFF0CFF48F9AF3326FD29" bold="true" box="[192,602,705,730]" pageId="225" pageNumber="224">
Distribution.
<collectingCountry id="F3690E9AFF2AFF0CFEF6F9AC3328FD29" box="[382,596,705,730]" name="French Guiana" pageId="225" pageNumber="224">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF2AFF0CFDECF9AC322FFD2A" author="Duranton, M." box="[612,851,706,730]" journalOrPublisher="Coleopteres, numero special" pageId="225" pageNumber="224" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 25</bibRefCitation>
(?).
<emphasis id="B90A9218FF2AFF0CFC05F9AF3295FD29" bold="true" box="[909,1001,705,729]" pageId="225" pageNumber="224">
<collectingCountry id="F3690E9AFF2AFF0CFC05F9AF329FFD29" box="[909,995,705,729]" name="Brazil" pageId="225" pageNumber="224">Brazil</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF2AFF0CFC7CF9AC35CBFD2A" author="Prell" box="[1012,1207,706,730]" firstAuthor="Prell" pageId="225" pageNumber="224" pagination="105 - 107" refId="ref220332" refString="Prell, H. 1911. Beitrage zur Kenntnis der Dynastinen. Entomologische Zeitschrift (Frankfurt) 25 (18): 105 - 107." type="journal article" year="1911">Prell 1911: 106</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF2AFF0CFB4FF9AC34E7FD2A" author="Arrow, G. J." bookContentInfo="124 p." box="[1223,1435,706,730]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="225" pageNumber="224" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 90</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF2AFF0CFF48F98C30A6FD0A" author="Blackwelder, R. E." box="[192,474,738,762]" journalOrPublisher="United States National Museum Bulletin" pageId="225" pageNumber="224" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 258</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF2AFF0CFE6EF98C33B7FD0A" author="Endrodi, S." box="[486,715,738,762]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="225" pageNumber="224" pagination="73 - 150" part="31" refId="ref201301" refString="Endrodi, S. 1978. Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera). Folia Entomologica Hungarica (N. S.) 31 (2): 73 - 150." title="Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera)" type="journal article" year="1978">Endrödi 1978: 105</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF2AFF0CFD50F98C321DFD0A" author="Endrodi, S." bookContentInfo="800 p." box="[728,865,738,762]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="225" pageNumber="224" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 722</bibRefCitation>
.
<emphasis id="B90A9218FF2AFF0CFCE5F98F32B9FD09" bold="true" box="[877,965,737,761]" pageId="225" pageNumber="224">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF2AFF0CFC45F98C35D3FD0A" author="Endrodi, S." box="[973,1199,738,762]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="225" pageNumber="224" pagination="73 - 150" part="31" refId="ref201301" refString="Endrodi, S. 1978. Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera). Folia Entomologica Hungarica (N. S.) 31 (2): 73 - 150." title="Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera)" type="journal article" year="1978">Endrödi 1978: 105</bibRefCitation>
(
<collectingCountry id="F3690E9AFF2AFF0CFB37F98C344AFD0A" box="[1215,1334,738,762]" name="Colombia" pageId="225" pageNumber="224">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF2AFF0CFACBF98C34E0FD0A" box="[1347,1436,738,762]" name="Mexico" pageId="225" pageNumber="224">Mexico</collectingCountry>
,
<collectingCountry id="F3690E9AFF2AFF0CFF48F86C3054FCE9" box="[192,296,770,793]" name="Panama" pageId="225" pageNumber="224">Panama</collectingCountry>
,
<collectingCountry id="F3690E9AFF2AFF0CFEBAF86C3013FCEA" box="[306,367,770,794]" name="Peru" pageId="225" pageNumber="224">Peru</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF2AFF0CFE0AF86C3310FCEA" author="Endrodi, S." bookContentInfo="800 p." box="[386,620,770,794]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="225" pageNumber="224" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi 1985a: 722</bibRefCitation>
(BrazilMexico);
<bibRefCitation id="EFEF33FBFF2AFF0CFCB7F86C329CFCEA" author="Ponchel" box="[831,992,770,794]" firstAuthor="Ponchel" pageId="225" pageNumber="224" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF2AFF0CFB39F86C3190FCCA" author="Ratcliffe" etAl="et al." firstAuthor="Ratcliffe" pageId="225" pageNumber="224" pagination="1 - 661" refId="ref222393" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 661." type="journal article" year="2013">Ratcliffe et al. 2013: 600</bibRefCitation>
(not in Mesoamerica, only South America);
<bibRefCitation id="EFEF33FBFF2AFF0CFC8EF84C32D3FCCA" author="Dupuis, F." box="[774,943,802,826]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="225" pageNumber="224" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b</bibRefCitation>
(not mentioned).
</paragraph>
<paragraph id="8BC14E0AFF2AFF0CFF48F82430A8FC32" blockId="225.[192,1440,841,962]" pageId="225" pageNumber="224">
<emphasis id="B90A9218FF2AFF0CFF48F824305DFC92" bold="true" box="[192,289,842,866]" pageId="225" pageNumber="224">Note 1.</emphasis>
<bibRefCitation id="EFEF33FBFF2AFF0CFEA1F824335FFC92" author="Duranton, M." box="[297,547,842,866]" journalOrPublisher="Coleopteres, numero special" pageId="225" pageNumber="224" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton (2011: 25)</bibRefCitation>
notes he saw no definite specimens of
<taxonomicName id="4C7E3589FF2AFF0CFB80F8273582FC91" authorityName="Prell" authorityYear="1911" box="[1032,1278,841,865]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="225" pageNumber="224" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B90A9218FF2AFF0CFB80F8273582FC91" box="[1032,1278,841,865]" italics="true" pageId="225" pageNumber="224">Phileurus excavatus</emphasis>
</taxonomicName>
from
<collectingCountry id="F3690E9AFF2AFF0CFAC1F8243062FC72" name="French Guiana" pageId="225" pageNumber="224">French Guiana</collectingCountry>
, and the species is not mentioned in the checklists for
<collectingCountry id="F3690E9AFF2AFF0CFC41F80435F8FC72" box="[969,1156,874,898]" name="French Guiana" pageId="225" pageNumber="224">French Guiana</collectingCountry>
by
<bibRefCitation id="EFEF33FBFF2AFF0CFB3BF8043415FC72" author="Ponchel" box="[1203,1385,874,898]" firstAuthor="Ponchel" pageId="225" pageNumber="224" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel (2011)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF2AFF0CFF48F8E43003FC52" author="Dupuis, F." box="[192,383,906,930]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="225" pageNumber="224" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis (2016b)</bibRefCitation>
. There are no definite records from the research area for this species, and we regard it as not occurring there.
</paragraph>
<paragraph id="8BC14E0AFF2AFF0CFF48F8BC3502FBDA" blockId="225.[192,1441,977,1066]" pageId="225" pageNumber="224">
<emphasis id="B90A9218FF2AFF0CFF48F8BC3061FC1A" bold="true" box="[192,285,978,1002]" pageId="225" pageNumber="224">Note 2.</emphasis>
<bibRefCitation id="EFEF33FBFF2AFF0CFEABF8BC3377FC1A" author="Endrodi, S." box="[291,523,978,1002]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="225" pageNumber="224" pagination="73 - 150" part="31" refId="ref201301" refString="Endrodi, S. 1978. Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera). Folia Entomologica Hungarica (N. S.) 31 (2): 73 - 150." title="Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera)" type="journal article" year="1978">Endrödi (1978: 105</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF2AFF0CFD9EF8BC33E5FC1A" author="Endrodi, S." bookContentInfo="800 p." box="[534,665,978,1002]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="225" pageNumber="224" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 722</bibRefCitation>
) records
<taxonomicName id="4C7E3589FF2AFF0CFC8EF8BF3285FC19" authorityName="Prell" authorityYear="1911" box="[774,1017,977,1001]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="225" pageNumber="224" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B90A9218FF2AFF0CFC8EF8BF3285FC19" box="[774,1017,977,1001]" italics="true" pageId="225" pageNumber="224">Phileurus excavatus</emphasis>
</taxonomicName>
for, among other countries,
<collectingCountry id="F3690E9AFF2AFF0CFAC2F8BC34DCFC1A" box="[1354,1440,978,1002]" name="Mexico" pageId="225" pageNumber="224">Mexico</collectingCountry>
and
<collectingCountry id="F3690E9AFF2AFF0CFF7DF89C3022FBF9" box="[245,350,1010,1033]" name="Panama" pageId="225" pageNumber="224">Panama</collectingCountry>
.
<bibRefCitation id="EFEF33FBFF2AFF0CFEE1F89C33DDFBFA" author="Ratcliffe" box="[361,673,1010,1034]" etAl="et al." firstAuthor="Ratcliffe" pageId="225" pageNumber="224" pagination="1 - 661" refId="ref222393" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 661." type="journal article" year="2013">Ratcliffe et al. (2013: 600)</bibRefCitation>
mention this, but state they have not seen any specimens from Central America. They consider this species to be restricted to South America.
</paragraph>
</subSubSection>
</treatment>
</document>