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<taxonomicName id="4C7E3589FF1BFF3DFF48FF0A33C3FB8C" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[192,703,1124,1148]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFF48FF0A33C3FB8C" bold="true" box="[192,703,1124,1148]" pageId="208" pageNumber="207">
<emphasis id="B90A9218FF1BFF3DFF48FF0A30C6FB8C" bold="true" box="[192,442,1124,1148]" italics="true" pageId="208" pageNumber="207">Euetheola humilis</emphasis>
(
<bibRefCitation id="EFEF33FBFF1BFF3DFE43FF0A33CBFB8C" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[459,695,1124,1148]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="208" pageNumber="207" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister, 1847</bibRefCitation>
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<treatmentCitation id="0ADF681BFF1BFF3DFF60FFE533C3FB50" author="Burmeister, H. C. C." box="[232,703,1163,1185]" page="93" pageId="208" pageNumber="207" year="1847">
<taxonomicName id="4C7E3589FF1BFF3DFF60FFE533C3FB50" authority="Burmeister 1847: 93" authorityName="Burmeister" authorityPageNumber="93" authorityYear="1847" box="[232,703,1163,1185]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFF60FFE530ABFB50" box="[232,471,1163,1184]" italics="true" pageId="208" pageNumber="207">Heteronychus humilis</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFE56FFE533C3FB50" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[478,703,1163,1185]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="208" pageNumber="207" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 93</bibRefCitation>
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(“Guianas,”
<collectingCountry id="F3690E9AFF1BFF3DFCC5FFE532E9FB51" box="[845,917,1163,1185]" name="Brazil" pageId="208" pageNumber="207">Brazil</collectingCountry>
,
<collectingCountry id="F3690E9AFF1BFF3DFC17FFE5357BFB51" box="[927,1031,1163,1185]" name="Colombia" pageId="208" pageNumber="207">Colombia</collectingCountry>
)
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<paragraph id="8BC14E0AFF1BFF3DFF48FFDF335DF97A" blockId="208.[192,1441,1201,1674]" pageId="208" pageNumber="207">
<emphasis id="B90A9218FF1BFF3DFF48FFDF3091FB39" bold="true" box="[192,493,1201,1225]" pageId="208" pageNumber="207">
Distribution.
<collectingCountry id="F3690E9AFF1BFF3DFEF2FFDF309BFB39" box="[378,487,1201,1225]" name="Guyana" pageId="208" pageNumber="207">Guyana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFE7CFFDF339AFB3A" author="Endrodi, S." bookContentInfo="800 p." box="[500,742,1201,1226]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="208" pageNumber="207" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi 1985a: 234</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFD79FFDC3554FB3A" author="Scavo and Joly" box="[753,1064,1201,1226]" firstAuthor="Scavo" pageId="208" pageNumber="207" pagination="173 - 186" refId="ref223627" refString="Scavo, M., and L. J. Joly. 1998. Las especies venezolanas del genero Euetheola (Bates 1888) (Coleoptera, Scarabaeidae, Dynastinae). Boletin de Entomologia Venezolana 13 (2): 173 - 186." type="journal article" year="1998">Scavo and Joly 1998: 177</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFBBCFFDC34E0FB3A" author="Lopez-Garcia" box="[1076,1436,1201,1226]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="208" pageNumber="207" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">López-García et al. 2016: 498</bibRefCitation>
.
<emphasis id="B90A9218FF1BFF3DFF48FFBF3032FB19" bold="true" box="[192,334,1233,1257]" pageId="208" pageNumber="207">
<collectingCountry id="F3690E9AFF1BFF3DFF48FFBF3034FB19" box="[192,328,1233,1257]" name="Suriname" pageId="208" pageNumber="207">Suriname</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFEDBFFBF3348FB1A" author="Endrodi, S." box="[339,564,1233,1258]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="208" pageNumber="207" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 14</bibRefCitation>
.
<emphasis id="B90A9218FF1BFF3DFDB5FFBF326EFB19" bold="true" box="[573,786,1233,1257]" pageId="208" pageNumber="207">
<collectingCountry id="F3690E9AFF1BFF3DFDB5FFBF3271FB19" box="[573,781,1233,1257]" name="French Guiana" pageId="208" pageNumber="207">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFC9FFFBC353AFB1A" author="Scavo and Joly" box="[791,1094,1233,1258]" firstAuthor="Scavo" pageId="208" pageNumber="207" pagination="173 - 186" refId="ref223627" refString="Scavo, M., and L. J. Joly. 1998. Las especies venezolanas del genero Euetheola (Bates 1888) (Coleoptera, Scarabaeidae, Dynastinae). Boletin de Entomologia Venezolana 13 (2): 173 - 186." type="journal article" year="1998">Scavo and Joly 1998: 177</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFBD8FFBC3449FB1A" author="Duranton, M." box="[1104,1333,1234,1258]" journalOrPublisher="Coleopteres, numero special" pageId="208" pageNumber="207" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 19</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFAB7FFBF305AFAF9" author="Ponchel" firstAuthor="Ponchel" pageId="208" pageNumber="207" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011: 61</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1BFF3DFEDEFF9F331DFAF9" box="[342,609,1265,1289]" class="Insecta" family="Scarabaeidae" genus="Eutheola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFEDEFF9F30B8FAF9" box="[342,452,1265,1289]" italics="true" pageId="208" pageNumber="207">Eutheola</emphasis>
[sic]
<emphasis id="B90A9218FF1BFF3DFD8BFF9F331DFAF9" box="[515,609,1265,1289]" italics="true" pageId="208" pageNumber="207">humilis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1BFF3DFDF0FF9F3222FAFA" author="Dupuis, F." box="[632,862,1265,1290]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="208" pageNumber="207" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 110</bibRefCitation>
.
<emphasis id="B90A9218FF1BFF3DFCE0FF9F3286FAF9" bold="true" box="[872,1018,1265,1289]" pageId="208" pageNumber="207">“Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFB89FF9F3585FAFA" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[1025,1273,1265,1290]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="208" pageNumber="207" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 93</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1BFF3DFAA2FF9C3022FAD9" authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFAA2FF9C3022FAD9" italics="true" pageId="208" pageNumber="207">Heteronychus humilis</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FF1BFF3DFEFEFE7F30AEFAD9" bold="true" box="[374,466,1297,1321]" pageId="208" pageNumber="207">
<collectingCountry id="F3690E9AFF1BFF3DFEFEFE7F30B0FAD9" box="[374,460,1297,1321]" name="Brazil" pageId="208" pageNumber="207">Brazil</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFE52FE7F33A9FADA" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[474,725,1297,1322]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="208" pageNumber="207" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 93</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1BFF3DFC81FE7F3569FAD9" authorityName="Burmeister" authorityYear="1847" box="[777,1045,1297,1321]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFC81FE7F3569FAD9" box="[777,1045,1297,1321]" italics="true" pageId="208" pageNumber="207">Heteronychus humilis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1BFF3DFBA5FE7C3587FAD9" author="Arrow, G. J." bookContentInfo="124 p." box="[1069,1275,1297,1322]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="208" pageNumber="207" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 36</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFA80FE7F304BFAB9" author="Blackwelder, R. E." journalOrPublisher="United States National Museum Bulletin" pageId="208" pageNumber="207" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1BFF3DFEE4FE5F3331FAB9" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[364,589,1329,1353]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFEE4FE5F3331FAB9" box="[364,589,1329,1353]" italics="true" pageId="208" pageNumber="207">Euetheola humilis</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF1BFF3DFD09FE5C3239FAB9" box="[641,837,1329,1353]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFD09FE5C3239FAB9" box="[641,837,1329,1353]" italics="true" pageId="208" pageNumber="207">Ligyrus humilis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1BFF3DFCD6FE5F3538FABA" author="Endrodi, S." box="[862,1092,1329,1354]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="208" pageNumber="207" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 14</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF1BFF3DFBD9FE5F35A6FABA" author="Endrodi, S." bookContentInfo="800 p." box="[1105,1242,1329,1354]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="208" pageNumber="207" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 234</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFB6FFE5C3044FA9A" author="Scavo and Joly" firstAuthor="Scavo" pageId="208" pageNumber="207" pagination="173 - 186" refId="ref223627" refString="Scavo, M., and L. J. Joly. 1998. Las especies venezolanas del genero Euetheola (Bates 1888) (Coleoptera, Scarabaeidae, Dynastinae). Boletin de Entomologia Venezolana 13 (2): 173 - 186." type="journal article" year="1998">Scavo and Joly 1998: 177</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFECCFE3C33D6FA9A" author="Lopez-Garcia" box="[324,682,1361,1386]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="208" pageNumber="207" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">López-García et al. 2016: 498</bibRefCitation>
.
<emphasis id="B90A9218FF1BFF3DFD3DFE3F32ECFA99" bold="true" box="[693,912,1361,1385]" pageId="208" pageNumber="207">
<collectingCountry id="F3690E9AFF1BFF3DFD3DFE3F323FFA98" box="[693,835,1361,1384]" name="Venezuela" pageId="208" pageNumber="207">Venezuela</collectingCountry>
(EO):
</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFC1FFE3F35D2FA99" author="Blackwelder, R. E." box="[919,1198,1361,1386]" journalOrPublisher="United States National Museum Bulletin" pageId="208" pageNumber="207" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFB31FE3F34E7FA9A" author="Endrodi, S." box="[1209,1435,1361,1386]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="208" pageNumber="207" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 14</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF1BFF3DFF48FE1F303BFA7A" author="Endrodi, S." bookContentInfo="800 p." box="[192,327,1393,1418]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="208" pageNumber="207" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 234</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1BFF3DFEDBFE1C33F9FA7A" author="Scavo and Joly" box="[339,645,1393,1418]" firstAuthor="Scavo" pageId="208" pageNumber="207" pagination="173 - 186" refId="ref223627" refString="Scavo, M., and L. J. Joly. 1998. Las especies venezolanas del genero Euetheola (Bates 1888) (Coleoptera, Scarabaeidae, Dynastinae). Boletin de Entomologia Venezolana 13 (2): 173 - 186." type="journal article" year="1998">Scavo and Joly 1998: 177</bibRefCitation>
(EO);
<bibRefCitation id="EFEF33FBFF1BFF3DFD5CFE1C3540FA7A" author="Lopez-Garcia" box="[724,1084,1393,1418]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="208" pageNumber="207" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">López-García et al. 2016: 498</bibRefCitation>
.
<emphasis id="B90A9218FF1BFF3DFBCFFE1F35E3FA79" bold="true" box="[1095,1183,1393,1417]" pageId="208" pageNumber="207">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFB2EFE1F34DCFA7A" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[1190,1440,1393,1418]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="208" pageNumber="207" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 93</bibRefCitation>
(Colombia - as
<taxonomicName id="4C7E3589FF1BFF3DFEF1FEFF33F9FA59" authorityName="Burmeister" authorityYear="1847" box="[377,645,1425,1449]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFEF1FEFF33F9FA59" box="[377,645,1425,1449]" italics="true" pageId="208" pageNumber="207">Heteronychus humilis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1BFF3DFD16FEFC3216FA59" author="Arrow, G. J." bookContentInfo="124 p." box="[670,874,1425,1450]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="208" pageNumber="207" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 36</bibRefCitation>
(Mexico - as
<taxonomicName id="4C7E3589FF1BFF3DFB87FEFF3593FA59" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1039,1263,1425,1449]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFB87FEFF3593FA59" box="[1039,1263,1425,1449]" italics="true" pageId="208" pageNumber="207">Euetheola humilis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1BFF3DFA80FEFF3049FA39" author="Blackwelder, R. E." journalOrPublisher="United States National Museum Bulletin" pageId="208" pageNumber="207" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(Mexico, Panama - as
<taxonomicName id="4C7E3589FF1BFF3DFDCEFEDF3259FA39" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[582,805,1457,1481]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFDCEFEDF3259FA39" box="[582,805,1457,1481]" italics="true" pageId="208" pageNumber="207">Euetheola humilis</emphasis>
</taxonomicName>
; Argentina - as
<taxonomicName id="4C7E3589FF1BFF3DFC6FFEDC35D6FA39" box="[999,1194,1457,1481]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFC6FFEDC35D6FA39" box="[999,1194,1457,1481]" italics="true" pageId="208" pageNumber="207">Ligyrus humilis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1BFF3DFB49FEDF34DCFA3A" author="Endrodi, S." box="[1217,1440,1457,1482]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="208" pageNumber="207" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 14</bibRefCitation>
(Argentina, Bolivia, Colombia, Mexico, Nicaragua, Paraguay, U.S.A., Uruguay), 1985a: 234 (Argentina, Bolivia, Colombia, Mexico, Nicaragua, Paraguay, U.S.A., Uruguay);
<bibRefCitation id="EFEF33FBFF1BFF3DFB8EFE9C3440F9FA" author="Scavo and Joly" box="[1030,1340,1521,1546]" firstAuthor="Scavo" pageId="208" pageNumber="207" pagination="173 - 186" refId="ref223627" refString="Scavo, M., and L. J. Joly. 1998. Las especies venezolanas del genero Euetheola (Bates 1888) (Coleoptera, Scarabaeidae, Dynastinae). Boletin de Entomologia Venezolana 13 (2): 173 - 186." type="journal article" year="1998">Scavo and Joly 1998: 177</bibRefCitation>
(Argentina, Bolivia, Colombia, Mexico, Nicaragua, Paraguay, U.S.A., Uruguay);
<bibRefCitation id="EFEF33FBFF1BFF3DFBEEFD7F34DCF9DA" author="Ratcliffe" box="[1126,1440,1553,1578]" etAl="et al." firstAuthor="Ratcliffe" pageId="208" pageNumber="207" pagination="1 - 661" refId="ref222393" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 661." type="journal article" year="2013">Ratcliffe et al. 2013: 278</bibRefCitation>
(ArgentinaU.S.A.);
<bibRefCitation id="EFEF33FBFF1BFF3DFE31FD5C3265F9B9" author="Lopez-Garcia" box="[441,793,1585,1610]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="208" pageNumber="207" pagination="451 - 492" refId="ref212996" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, and G. Amat-Garcia. 2015. The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa 4048 (4): 451 - 492." type="journal article" year="2015">López-García et al. 2015: 463</bibRefCitation>
(ArgentinaU.S.A., Colombia), 2016: 498 (Argentina, Belize, Bolivia, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, U.S.A., Uruguay).
</paragraph>
<paragraph id="8BC14E0AFF1BFF3DFF48FDF73539F8C1" blockId="208.[192,1440,1689,1842]" pageId="208" pageNumber="207">
<emphasis id="B90A9218FF1BFF3DFF48FDF7307BF941" bold="true" box="[192,263,1689,1713]" pageId="208" pageNumber="207">Note.</emphasis>
<taxonomicName id="4C7E3589FF1BFF3DFE84FDF433FDF941" authority="LeConte, 1856" authorityName="LeConte" authorityYear="1856" box="[268,641,1689,1714]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="rugiceps">
<emphasis id="B90A9218FF1BFF3DFE84FDF430ACF941" box="[268,464,1690,1713]" italics="true" pageId="208" pageNumber="207">Ligyrus rugiceps</emphasis>
<bibRefCitation id="EFEF33FBFF1BFF3DFE5DFDF433FDF941" author="LeConte" box="[469,641,1689,1714]" firstAuthor="LeConte" pageId="208" pageNumber="207" pagination="19 - 25" refId="ref212484" refString="LeConte, J. L. 1856. Notice of three genera of Scarabaeidae found in the United States. Journal of the Academy of Natural Sciences of Philadelphia 8: 19 - 25." type="journal article" year="1856">LeConte, 1856</bibRefCitation>
</taxonomicName>
(p. 21) is mentioned as a subspecies of
<taxonomicName id="4C7E3589FF1BFF3DFBCCFDF7303CF921" authority="in Endrodi" authorityName="in Endrodi" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFBCCFDF73460F941" box="[1092,1308,1689,1713]" italics="true" pageId="208" pageNumber="207">Euetheola humilis</emphasis>
in
<bibRefCitation id="EFEF33FBFF1BFF3DFAB6FDF7303CF921" author="Endrodi, S." journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="208" pageNumber="207" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi (1969a: 15</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFEF33FBFF1BFF3DFEC2FDD730B2F921" author="Endrodi, S." bookContentInfo="800 p." box="[330,462,1721,1745]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="208" pageNumber="207" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 235</bibRefCitation>
) and considered a synonym of
<taxonomicName id="4C7E3589FF1BFF3DFCB5FDD43469F922" authority="in Ratcliffe et al. (2013: 278)" authorityName="in Ratcliffe" authorityPageNumber="278" authorityYear="2013" box="[829,1301,1721,1746]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFCB5FDD432C0F921" box="[829,956,1721,1745]" italics="true" pageId="208" pageNumber="207">E. humilis</emphasis>
in
<bibRefCitation id="EFEF33FBFF1BFF3DFC68FDD73469F922" author="Ratcliffe" box="[992,1301,1721,1746]" etAl="et al." firstAuthor="Ratcliffe" pageId="208" pageNumber="207" pagination="1 - 661" refId="ref222393" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 661." type="journal article" year="2013">Ratcliffe et al. (2013: 278)</bibRefCitation>
</taxonomicName>
and López- García et al. (2015: 463, 2016: 498). In
<bibRefCitation id="EFEF33FBFF1BFF3DFD2BFDB7356BF902" author="Ratcliffe and Cave" box="[675,1047,1753,1778]" firstAuthor="Ratcliffe" pageId="208" pageNumber="207" pagination="1 - 293" refId="ref222345" refString="Ratcliffe, B. C., and R. D. Cave. 2017. The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 30: 1 - 293." type="journal article" year="2017">Ratcliffe and Cave (2017: 128)</bibRefCitation>
this synonymy is revoked, with
<taxonomicName id="4C7E3589FF1BFF3DFF48FD943034F8E1" authorityName="LeConte" authorityYear="1856" box="[192,328,1786,1809]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="rugiceps">
<emphasis id="B90A9218FF1BFF3DFF48FD943034F8E1" box="[192,328,1786,1809]" italics="true" pageId="208" pageNumber="207">E. rugiceps</emphasis>
</taxonomicName>
believed to be a species endemic to the
<collectingCountry id="F3690E9AFF1BFF3DFCA5FD943200F8E2" box="[813,892,1786,1810]" name="United States of America" pageId="208" pageNumber="207">U.S.A.</collectingCountry>
Based on
<bibRefCitation id="EFEF33FBFF1BFF3DFC73FD973587F8E2" author="Ratcliffe" box="[1019,1275,1785,1810]" etAl="et al." firstAuthor="Ratcliffe" pageId="208" pageNumber="207" pagination="1 - 661" refId="ref222393" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 661." type="journal article" year="2013">Ratcliffe et al. (2013)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF1BFF3DFAB0FD9730F5F8C2" author="Ratcliffe and Cave" firstAuthor="Ratcliffe" pageId="208" pageNumber="207" pagination="1 - 293" refId="ref222345" refString="Ratcliffe, B. C., and R. D. Cave. 2017. The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 30: 1 - 293." type="journal article" year="2017">Ratcliffe and Cave (2017)</bibRefCitation>
,
<taxonomicName id="4C7E3589FF1BFF3DFE1EFC773309F8C1" box="[406,629,1817,1841]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="207" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B90A9218FF1BFF3DFE1EFC773309F8C1" box="[406,629,1817,1841]" italics="true" pageId="208" pageNumber="207">Euetheola humilis</emphasis>
</taxonomicName>
has its northernmost limit in
<collectingCountry id="F3690E9AFF1BFF3DFC6FFC77353DF8C1" box="[999,1089,1817,1841]" name="Mexico" pageId="208" pageNumber="207">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>