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§
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<emphasis id="B90A9218FF17FF31FF5DFEE2334AFA54" bold="true" box="[213,566,1420,1444]" italics="true" pageId="220" pageNumber="219">Hemiphileurus variolosus</emphasis>
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<emphasis id="B90A9218FF17FF31FF60FEDD30B0FA38" box="[232,460,1459,1480]" italics="true" pageId="220" pageNumber="219">Phileurus variolosus</emphasis>
<bibRefCitation id="EFEF33FBFF17FF31FE5BFEDD33BDFA39" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[467,705,1459,1481]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="220" pageNumber="219" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 156</bibRefCitation>
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Distribution.
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:
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<bibRefCitation id="EFEF33FBFF17FF31FDD3FEB7324EFA02" author="Endrodi, S." box="[603,818,1497,1522]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="220" pageNumber="219" pagination="73 - 150" part="31" refId="ref201301" refString="Endrodi, S. 1978. Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera). Folia Entomologica Hungarica (N. S.) 31 (2): 73 - 150." title="Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera)" type="journal article" year="1978">Endrödi 1978: 90</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF17FF31FCB7FEB73554FA02" author="Ratcliffe" box="[831,1064,1497,1522]" firstAuthor="Ratcliffe" pageId="220" pageNumber="219" pagination="433 - 443" refId="ref221542" refString="Ratcliffe, B. C. 2001. New species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) from Mexico, Guatemala, Colombia, and Brazil. The Coleopterists Bulletin 55 (4): 433 - 443." type="journal article" year="2001">Ratcliffe 2001: 442</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF17FF31FBBDFEB734E0FA02" author="Ratcliffe and Cave" box="[1077,1436,1497,1522]" firstAuthor="Ratcliffe" pageId="220" pageNumber="219" pagination="1 - 424" refId="ref222249" refString="Ratcliffe, B. C., and R. D. Cave. 2006. The dynastine scarab beetles of Honduras, Nicaragua, and El Salvador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 21: 1 - 424." type="journal article" year="2006">Ratcliffe and Cave 2006: 322</bibRefCitation>
.
<emphasis id="B90A9218FF17FF31FF48FE97302AF9E1" bold="true" box="[192,342,1529,1553]" pageId="220" pageNumber="219">
<collectingCountry id="F3690E9AFF17FF31FF48FE97302CF9E1" box="[192,336,1529,1553]" name="Venezuela" pageId="220" pageNumber="219">Venezuela</collectingCountry>
:
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<bibRefCitation id="EFEF33FBFF17FF31FED6FE943350F9E2" author="Arrow, G. J." bookContentInfo="124 p." box="[350,556,1529,1554]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="220" pageNumber="219" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 88</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF17FF31FDB1FE97322FF9E1" author="Blackwelder, R. E." box="[569,851,1529,1554]" journalOrPublisher="United States National Museum Bulletin" pageId="220" pageNumber="219" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 258</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF17FF31FCD7FE973549F9E2" author="Endrodi, S." box="[863,1077,1529,1554]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="220" pageNumber="219" pagination="73 - 150" part="31" refId="ref201301" refString="Endrodi, S. 1978. Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera). Folia Entomologica Hungarica (N. S.) 31 (2): 73 - 150." title="Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera)" type="journal article" year="1978">Endrödi 1978: 90</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF17FF31FBCAFE973450F9E2" author="Ratcliffe" box="[1090,1324,1529,1554]" firstAuthor="Ratcliffe" pageId="220" pageNumber="219" pagination="433 - 443" refId="ref221542" refString="Ratcliffe, B. C. 2001. New species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) from Mexico, Guatemala, Colombia, and Brazil. The Coleopterists Bulletin 55 (4): 433 - 443." type="journal article" year="2001">Ratcliffe 2001: 442</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF17FF31FAB0FE9730C8F9C2" author="Ratcliffe and Cave" firstAuthor="Ratcliffe" pageId="220" pageNumber="219" pagination="1 - 424" refId="ref222249" refString="Ratcliffe, B. C., and R. D. Cave. 2006. The dynastine scarab beetles of Honduras, Nicaragua, and El Salvador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 21: 1 - 424." type="journal article" year="2006">Ratcliffe and Cave 2006: 322</bibRefCitation>
.
<emphasis id="B90A9218FF17FF31FE37FD77336BF9C1" bold="true" box="[447,535,1561,1585]" pageId="220" pageNumber="219">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF17FF31FD96FD77325BF9C1" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[542,807,1561,1586]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="220" pageNumber="219" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 156</bibRefCitation>
(Colombia - as
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<emphasis id="B90A9218FF17FF31FC6DFD77359CF9C1" box="[997,1248,1561,1585]" italics="true" pageId="220" pageNumber="219">Phileurus variolosus</emphasis>
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1978: 90
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<emphasis id="B90A9218FF17FF31FE29FD0F33E4F989" box="[417,664,1633,1657]" italics="true" pageId="220" pageNumber="219">Phileurus variolosus</emphasis>
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), 1985a: 715 (
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<collectingCountry id="F3690E9AFF17FF31FD17FDEF326AF969" box="[671,790,1665,1689]" name="Colombia" pageId="220" pageNumber="219">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FCA8FDEC32D8F969" box="[800,932,1665,1690]" name="Costa Rica" pageId="220" pageNumber="219">Costa Rica</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FC26FDEF356BF969" box="[942,1047,1665,1689]" name="Ecuador" pageId="220" pageNumber="219">Ecuador</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FBA9FDEC35F5F969" box="[1057,1161,1666,1689]" name="Panama" pageId="220" pageNumber="219">Panama</collectingCountry>
</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FB1CFDEF34E8F969" box="[1172,1428,1665,1689]" name="Trinidad and Tobago" pageId="220" pageNumber="219">Trinidad and Tobago</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF17FF31FF48FDCF3366F94A" author="Ratcliffe and Cave" box="[192,538,1697,1722]" firstAuthor="Ratcliffe" pageId="220" pageNumber="219" pagination="1 - 424" refId="ref222249" refString="Ratcliffe, B. C., and R. D. Cave. 2006. The dynastine scarab beetles of Honduras, Nicaragua, and El Salvador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 21: 1 - 424." type="journal article" year="2006">Ratcliffe and Cave 2006: 322</bibRefCitation>
(
<collectingCountry id="F3690E9AFF17FF31FDA0FDCF3455F949" box="[552,1321,1697,1722]" isEnumeration="true" pageId="220" pageNumber="219">
<collectingCountry id="F3690E9AFF17FF31FDA0FDCF33E3F949" box="[552,671,1697,1721]" name="Colombia" pageId="220" pageNumber="219">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FD22FDCC3252F949" box="[682,814,1697,1722]" name="Costa Rica" pageId="220" pageNumber="219">Costa Rica</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FCB1FDCF32DEF949" box="[825,930,1697,1721]" name="Ecuador" pageId="220" pageNumber="219">Ecuador</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FC24FDCF3555F949" box="[940,1065,1697,1721]" name="Honduras" pageId="220" pageNumber="219">Honduras</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FBBCFDCF35CBF949" box="[1076,1207,1697,1721]" name="Nicaragua" pageId="220" pageNumber="219">Nicaragua</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FB49FDCC3455F949" box="[1217,1321,1698,1721]" name="Panama" pageId="220" pageNumber="219">Panama</collectingCountry>
</collectingCountry>
,
<collectingCountry id="F3690E9AFF17FF31FABCFDCF302EF929" name="Trinidad and Tobago" pageId="220" pageNumber="219">Trinidad and Tobago</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF17FF31FEEDFDAF3374F92A" author="Ponchel" box="[357,520,1729,1754]" firstAuthor="Ponchel" pageId="220" pageNumber="219" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF17FF31FD68FDAF32F6F929" author="Dupuis, F." box="[736,906,1729,1753]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="220" pageNumber="219" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b</bibRefCitation>
(not mentioned).
</paragraph>
<paragraph id="8BC14E0AFF17FF31FF48FD8735D6F871" blockId="220.[192,1440,1769,1921]" pageId="220" pageNumber="219">
<emphasis id="B90A9218FF17FF31FF48FD873074F8F1" bold="true" box="[192,264,1769,1793]" pageId="220" pageNumber="219">Note.</emphasis>
<bibRefCitation id="EFEF33FBFF17FF31FE86FD873092F8F2" author="Endrodi, S." box="[270,494,1769,1794]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="220" pageNumber="219" pagination="73 - 150" part="31" refId="ref201301" refString="Endrodi, S. 1978. Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera). Folia Entomologica Hungarica (N. S.) 31 (2): 73 - 150." title="Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera)" type="journal article" year="1978">Endrödi (1978: 90)</bibRefCitation>
records
<taxonomicName id="4C7E3589FF17FF31FDDAFD8732F0F8F1" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[594,908,1769,1793]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="220" pageNumber="219" phylum="Arthropoda" rank="species" species="variolosus">
<emphasis id="B90A9218FF17FF31FDDAFD8732F0F8F1" box="[594,908,1769,1793]" italics="true" pageId="220" pageNumber="219">Hemiphileurus variolosus</emphasis>
</taxonomicName>
for
<collectingCountry id="F3690E9AFF17FF31FC31FD87350FF8F1" box="[953,1139,1769,1793]" name="French Guiana" pageId="220" pageNumber="219">French Guiana</collectingCountry>
, and this record is copied in
<bibRefCitation id="EFEF33FBFF17FF31FF69FC6730A7F8D2" author="Ratcliffe" box="[225,475,1801,1826]" firstAuthor="Ratcliffe" pageId="220" pageNumber="219" pagination="433 - 443" refId="ref221542" refString="Ratcliffe, B. C. 2001. New species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) from Mexico, Guatemala, Colombia, and Brazil. The Coleopterists Bulletin 55 (4): 433 - 443." type="journal article" year="2001">Ratcliffe (2001: 442)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF17FF31FD93FC6732E9F8D2" author="Ratcliffe and Cave" box="[539,917,1801,1826]" firstAuthor="Ratcliffe" pageId="220" pageNumber="219" pagination="1 - 424" refId="ref222249" refString="Ratcliffe, B. C., and R. D. Cave. 2006. The dynastine scarab beetles of Honduras, Nicaragua, and El Salvador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 21: 1 - 424." type="journal article" year="2006">Ratcliffe and Cave (2006: 322)</bibRefCitation>
. However, we are not aware of any recent records from the research area. The species seems to have a more western distribution. Because of this, we assume the specimen from
<collectingCountry id="F3690E9AFF17FF31FDBEFC27338CF891" box="[566,752,1865,1889]" name="French Guiana" pageId="220" pageNumber="219">French Guiana</collectingCountry>
in the MNM (= MNCN) as mentioned in
<bibRefCitation id="EFEF33FBFF17FF31FB61FC2734E3F892" author="Endrodi, S." box="[1257,1439,1865,1890]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="220" pageNumber="219" pagination="73 - 150" part="31" refId="ref201301" refString="Endrodi, S. 1978. Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera). Folia Entomologica Hungarica (N. S.) 31 (2): 73 - 150." title="Monographie der Dynastinae. 8. Tribus: Phileurini, amerikanische Arten II. (Coleoptera)" type="journal article" year="1978">Endrödi (1978)</bibRefCitation>
has been mislabeled. We regard this species as not occurring in the research area.
</paragraph>
</subSubSection>
</treatment>
</document>