treatments-xml/data/03/D7/FF/03D7FF1CFF0EFF2BFF48FFEC35E6FCF1.xml
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<taxonomicName id="4C7E3589FF0EFF28FF48FFEC33D7FB69" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[192,683,1153,1177]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="actaeon">
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<emphasis id="B90A9218FF0EFF28FF48FFEC30BFFB69" bold="true" box="[192,451,1154,1177]" italics="true" pageId="197" pageNumber="196">Megasoma actaeon</emphasis>
(
<bibRefCitation id="EFEF33FBFF0EFF28FE5BFFEF33DFFB69" author="Linnaeus" box="[467,675,1153,1177]" firstAuthor="Linnaeus" pageId="197" pageNumber="196" refId="ref212788" refString="Linnaeus, C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii; Stockholm. 823 p." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
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<treatmentCitationGroup id="AB6E6924FF0EFF28FF60FFC63265FB4E" box="[232,793,1192,1214]" pageId="197" pageNumber="196">
<taxonomicName id="4C7E3589FF0EFF28FF60FFC63265FB4E" ID-CoL="6XVSF" authority="Linnaeus 1758: 345" authorityName="Linnaeus" authorityPageNumber="345" authorityYear="1758" box="[232,793,1192,1214]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="actaeon">
<treatmentCitation id="0ADF681BFF0EFF28FF60FFC633D9FB4E" ID-DOI="http://doi.org/10.5281/zenodo.3922358" ID-Zenodo-Dep="3922358" LSID="urn:lsid:zoobank.org:act:9B04709C-1D5B-43C9-B7FC-B4D960C5589F" author="Linnaeus" box="[232,677,1192,1214]" httpUri="http://treatment.plazi.org/id/7CCDE841FD77144E7EA793AF16C341A1" page="345" pageId="197" pageNumber="196" year="1758">
<emphasis id="B90A9218FF0EFF28FF60FFC630B9FB4E" box="[232,453,1192,1214]" italics="true" pageId="197" pageNumber="196">Scarabaeus actaeon</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FE44FFC633D9FB4E" author="Linnaeus" box="[460,677,1192,1214]" firstAuthor="Linnaeus" pageId="197" pageNumber="196" refId="ref212788" refString="Linnaeus, C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii; Stockholm. 823 p." type="book" year="1758">Linnaeus 1758: 345</bibRefCitation>
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(America)
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<treatmentCitationGroup id="AB6E6924FF0EFF28FF60FFAB326FFB2B" box="[232,787,1221,1243]" pageId="197" pageNumber="196">
-
<taxonomicName id="4C7E3589FF0EFF28FF7FFFAB326FFB2B" ID-CoL="6R86G" authority="Felsche 1906: 352" authorityName="Felsche" authorityPageNumber="352" authorityYear="1906" box="[247,787,1221,1243]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="janus">
<treatmentCitation id="0ADF681BFF0EFF28FF7FFFAB33ECFB2B" author="Felsche, C." box="[247,656,1221,1243]" page="352" pageId="197" pageNumber="196" year="1906">
<emphasis id="B90A9218FF0EFF28FF7FFFAB30B8FB2A" box="[247,452,1221,1242]" italics="true" pageId="197" pageNumber="196">Megalosoma janus</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FE43FFAB33ECFB2B" author="Felsche, C." box="[459,656,1221,1243]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="197" pageNumber="196" pagination="349 - 352" part="1906" refId="ref202252" refString="Felsche, C. 1906. Synonymische Bemerkungen uber einige Scarabaeiden aus der Tribus der Dynastini und Beschreibung einer neuen Art. Deutsche Entomologische Zeitschrift 1906 (2): 349 - 352." title="Synonymische Bemerkungen uber einige Scarabaeiden aus der Tribus der Dynastini und Beschreibung einer neuen Art" type="journal article" year="1906">Felsche 1906: 352</bibRefCitation>
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(
<collectingCountry id="F3690E9AFF0EFF28FD17FFAB3277FB2B" box="[671,779,1221,1243]" name="Paraguay" pageId="197" pageNumber="196">Paraguay</collectingCountry>
)
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<paragraph id="8BC14E0AFF0EFF28FF60FF8C32F2FB08" blockId="197.[232,963,1192,1329]" box="[232,910,1250,1272]" pageId="197" pageNumber="196">
<treatmentCitationGroup id="AB6E6924FF0EFF28FF60FF8C32F2FB08" box="[232,910,1250,1272]" pageId="197" pageNumber="196">
-
<taxonomicName id="4C7E3589FF0EFF28FF7FFF8C32F2FB08" authority="Hohne 1923 a: 254" authorityName="Hohne" authorityPageNumber="254" authorityYear="1923" box="[247,910,1250,1272]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="argentinum">
<treatmentCitation id="0ADF681BFF0EFF28FF7FFF8C327BFB08" author="Hohne, W." box="[247,775,1250,1272]" page="254" pageId="197" pageNumber="196" year="1923">
<emphasis id="B90A9218FF0EFF28FF7FFF8C334BFB07" box="[247,567,1250,1271]" italics="true" pageId="197" pageNumber="196">Megasoma janus argentinum</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FDB6FF8C327BFB08" author="Hohne, W." box="[574,775,1250,1272]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="197" pageNumber="196" pagination="252 - 255" part="1923" refId="ref208127" refString="Hohne, W. 1923 a. Neue Dynastiden (Col.). Deutsche Entomologische Zeitschrift 1923 (3): 252 - 255." title="Neue Dynastiden (Col.)" type="journal article" year="1923">Höhne 1923a: 254</bibRefCitation>
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(
<collectingCountry id="F3690E9AFF0EFF28FC9DFF8C32FBFB08" box="[789,903,1250,1272]" name="Argentina" pageId="197" pageNumber="196">Argentina</collectingCountry>
)
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<paragraph id="8BC14E0AFF0EFF28FF60FF913274FAE5" blockId="197.[232,963,1192,1329]" box="[232,776,1278,1301]" pageId="197" pageNumber="196">
<treatmentCitationGroup id="AB6E6924FF0EFF28FF60FF913274FAE5" box="[232,776,1278,1301]" pageId="197" pageNumber="196">
-
<taxonomicName id="4C7E3589FF0EFF28FF7FFF913274FAE5" authority="Nagai 2003: 39" authorityName="Nagai" authorityPageNumber="39" authorityYear="2003" box="[247,776,1278,1301]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="fujitai">
<treatmentCitation id="0ADF681BFF0EFF28FF7FFF9133D7FAE4" author="Nagai" box="[247,683,1278,1301]" page="39" pageId="197" pageNumber="196" year="2003">
<emphasis id="B90A9218FF0EFF28FF7FFF913087FAE3" box="[247,507,1278,1300]" italics="true" pageId="197" pageNumber="196">Megasoma janus fujitai</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FD8AFF9133D7FAE4" author="Nagai" box="[514,683,1279,1301]" firstAuthor="Nagai" pageId="197" pageNumber="196" pagination="35 - 39" refId="ref216675" refString="Nagai, S. 2003. Four new subspecies of the genus Megasoma (Coleoptera, Scarabaeidae) from South America. Gekkan-Mushi 394: 35 - 39." type="journal article" year="2003">Nagai 2003: 39</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF0EFF28FD31FF91327CFAE5" box="[697,768,1279,1301]" name="Brazil" pageId="197" pageNumber="196">Brazil</collectingCountry>
)
</taxonomicName>
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</paragraph>
<paragraph id="8BC14E0AFF0EFF28FF60FE7232BFFAC1" blockId="197.[232,963,1192,1329]" box="[232,963,1307,1329]" pageId="197" pageNumber="196">
<treatmentCitationGroup id="AB6E6924FF0EFF28FF60FE7232BFFAC1" box="[232,963,1307,1329]" pageId="197" pageNumber="196">
-
<taxonomicName id="4C7E3589FF0EFF28FF7FFE7232BFFAC1" authority="Silvestre and Arnaud 2002: 7" authorityName="Silvestre and Arnaud" authorityPageNumber="7" authorityYear="2002" box="[247,963,1307,1329]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="ramirezorum">
<treatmentCitation id="0ADF681BFF0EFF28FF7FFE72322CFAC1" author="Silvestre and Arnaud" box="[247,848,1307,1329]" page="7" pageId="197" pageNumber="196" year="2002">
<emphasis id="B90A9218FF0EFF28FF7FFE72337FFAC1" box="[247,515,1308,1329]" italics="true" pageId="197" pageNumber="196">Megasoma ramirezorum</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FD82FE75322CFAC1" author="Silvestre and Arnaud" box="[522,848,1307,1329]" firstAuthor="Silvestre" pageId="197" pageNumber="196" pagination="7 - 8" refId="ref225031" refString="Silvestre, G., and P. Arnaud. 2002. Description d'une nouvelle espece sud-americaine du genre Megasoma (Coleoptere Dynastidae). Besoiro 8: 7 - 8." type="journal article" year="2002">Silvestre and Arnaud 2002: 7</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF0EFF28FCD6FE7532C0FAC1" box="[862,956,1307,1329]" name="Ecuador" pageId="197" pageNumber="196">Ecuador</collectingCountry>
)
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</paragraph>
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<emphasis id="B90A9218FF0EFF28FF48FE2F3093FAA9" bold="true" box="[192,495,1345,1369]" pageId="197" pageNumber="196">
Distribution.
<collectingCountry id="F3690E9AFF0EFF28FEF4FE2F3095FAA9" box="[380,489,1345,1369]" name="Guyana" pageId="197" pageNumber="196">Guyana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FE70FE2C3394FAAA" author="Erichson, W. F." box="[504,744,1346,1370]" editor="R. Schomburgk" journalOrPublisher="J. J. Weber; Leipzig, Germany" pageId="197" pageNumber="196" pagination="553 - 1260" refId="ref201522" refString="Erichson, W. F. 1848. Insecten. p. 553 - 617. In: R. Schomburgk. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil. J. J. Weber; Leipzig, Germany. p. 553 - 1260." title="Insecten" type="book chapter" volumeTitle="Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil" year="1848">Erichson 1848: 560</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0EFF28FD7EFE2C32AEFAAA" author="Bodkin, G. E." box="[758,978,1346,1370]" journalOrPublisher="The Entomologist's Monthly Magazine" pageId="197" pageNumber="196" pagination="210 - 219" part="55" refId="ref194211" refString="Bodkin, G. E. 1919. Notes on the Coleoptera of British Guiana. The Entomologist's Monthly Magazine 55: 210 - 219." title="Notes on the Coleoptera of British Guiana" type="journal article" year="1919">Bodkin 1919: 214</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0EFF28FC68FE2C35BBFAAA" author="Endrodi, S." box="[992,1223,1346,1370]" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="197" pageNumber="196" pagination="37 - 86" part="23" refId="ref201221" refString="Endrodi, S. 1977 b. Monographie der Dynastinae (Coleoptera). 6. Tribus: Dynastini II. Acta Zoologica Academiae Scientiarum Hungaricae 23 (1 - 2): 37 - 86." title="Monographie der Dynastinae (Coleoptera). 6. Tribus: Dynastini II" type="journal article" year="1977">Endrödi 1977b: 41</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF0EFF28FB5DFE2C3422FAAA" author="Endrodi, S." bookContentInfo="800 p." box="[1237,1374,1346,1370]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="197" pageNumber="196" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 637</bibRefCitation>
, 642;
<bibRefCitation id="EFEF33FBFF0EFF28FF48FE0C3000FA8A" author="Nagai" box="[192,380,1378,1402]" firstAuthor="Nagai" pageId="197" pageNumber="196" pagination="35 - 39" refId="ref216675" refString="Nagai, S. 2003. Four new subspecies of the genus Megasoma (Coleoptera, Scarabaeidae) from South America. Gekkan-Mushi 394: 35 - 39." type="journal article" year="2003">Nagai 2003: 39</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FE3BFE0C33B5FA89" box="[435,713,1378,1401]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="ramirezorum">
<emphasis id="B90A9218FF0EFF28FE3BFE0C33B5FA89" box="[435,713,1378,1401]" italics="true" pageId="197" pageNumber="196">M. janus ramirezorum</emphasis>
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);
<bibRefCitation id="EFEF33FBFF0EFF28FD6BFE0C32B3FA8A" author="Ratcliffe" box="[739,975,1378,1402]" firstAuthor="Ratcliffe" pageId="197" pageNumber="196" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 461</bibRefCitation>
.
<emphasis id="B90A9218FF0EFF28FC54FE0F3517FA89" bold="true" box="[988,1131,1377,1401]" pageId="197" pageNumber="196">
<collectingCountry id="F3690E9AFF0EFF28FC54FE0F3518FA89" box="[988,1124,1377,1401]" name="Suriname" pageId="197" pageNumber="196">Suriname</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FBFBFE0C3435FA8A" author="Endrodi, S." box="[1139,1353,1378,1402]" journalOrPublisher="Folia Entomologica Hungarica" pageId="197" pageNumber="196" pagination="64 - 73" part="6" refId="ref200487" refString="Endrodi, S. 1941. Die Dynastiden-Gattung Megasoma Kirby (Coleopt.). Folia Entomologica Hungarica 6 (3 - 4): 64 - 73." title="Die Dynastiden-Gattung Megasoma Kirby (Coleopt.)" type="journal article" year="1941">Endrödi 1941: 69</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FA08FE0C3072FA69" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FA08FE0C3072FA69" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]); MHCN; NZCS.
<emphasis id="B90A9218FF0EFF28FD9EFEEC3397FA69" bold="true" box="[534,747,1409,1434]" pageId="197" pageNumber="196">
<collectingCountry id="F3690E9AFF0EFF28FD9EFEEC339AFA69" box="[534,742,1409,1434]" name="French Guiana" pageId="197" pageNumber="196">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FD78FEEC32C2FA6A" author="Endrodi, S." box="[752,958,1410,1434]" journalOrPublisher="Folia Entomologica Hungarica" pageId="197" pageNumber="196" pagination="64 - 73" part="6" refId="ref200487" refString="Endrodi, S. 1941. Die Dynastiden-Gattung Megasoma Kirby (Coleopt.). Folia Entomologica Hungarica 6 (3 - 4): 64 - 73." title="Die Dynastiden-Gattung Megasoma Kirby (Coleopt.)" type="journal article" year="1941">Endrödi 1941: 69</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FC65FEEC351DFA69" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1005,1121,1410,1433]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FC65FEEC351DFA69" box="[1005,1121,1410,1433]" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]);
<bibRefCitation id="EFEF33FBFF0EFF28FB26FEEC34E7FA6A" author="Lachaume" box="[1198,1435,1410,1434]" firstAuthor="Lachaume" pageId="197" pageNumber="196" pagination="1 - 85" refId="ref211158" refString="Lachaume, G. 1985. Dynastini 1. Dynastes - Megasoma - Golofa. Les Coleopteres du Monde 5: 1 - 85." type="journal article" year="1985">Lachaume 1985: 34</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0EFF28FF48FECC30A4FA4A" author="Brule, S. &amp; J. Touroult &amp; P. - H. Dalens" bookContentInfo="120 p." box="[192,472,1442,1466]" journalOrPublisher="Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF" pageId="197" pageNumber="196" refId="ref195373" refString="Brule, S., J. Touroult, and P. - H. Dalens (coord.). 2011 a. Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane), 2009 - 2010. Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF. 120 p. + annexes." title="Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane)" type="book" year="2011">Brûlé et al. 2011a: 191</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FD85FECC33FFFA49" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[525,643,1442,1465]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FD85FECC33FFFA49" box="[525,643,1442,1465]" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]), 2014: 182 (as
<taxonomicName id="4C7E3589FF0EFF28FC08FECC328AFA49" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[896,1014,1442,1465]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FC08FECC328AFA49" box="[896,1014,1442,1465]" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]);
<bibRefCitation id="EFEF33FBFF0EFF28FBC0FECC344EFA4A" author="Duranton, M." box="[1096,1330,1442,1466]" journalOrPublisher="Coleopteres, numero special" pageId="197" pageNumber="196" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 22</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0EFF28FAB7FECC305AFA2A" author="Ponchel" firstAuthor="Ponchel" pageId="197" pageNumber="196" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011: 61</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FED1FEAC30B2FA29" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[345,462,1474,1497]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FED1FEAC30B2FA29" box="[345,462,1474,1497]" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]);
<bibRefCitation id="EFEF33FBFF0EFF28FD96FEAC3235FA2A" author="Brule, S. &amp; P. - H. Dalens" bookContentInfo="23 p." box="[542,841,1474,1498]" journalOrPublisher="Rapport de la Societe entomologique Antilles-Guyane" pageId="197" pageNumber="196" refId="ref195220" refString="Brule, S., and P. - H. Dalens. 2012 a. Etude et inventaire entomologique du Mont Itoupe. Mission 2010 pour le compte du Parc Amazonien de Guyane. Rapport de la Societe entomologique Antilles-Guyane. 23 p. + annexes." title="Etude et inventaire entomologique du Mont Itoupe. Mission 2010 pour le compte du Parc Amazonien de Guyane" type="book" year="2012">Brûlé and Dalens 2012a</bibRefCitation>
: annex 4: 18 (as
<taxonomicName id="4C7E3589FF0EFF28FB9BFEAC35F4FA29" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1043,1160,1474,1497]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FB9BFEAC35F4FA29" box="[1043,1160,1474,1497]" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]), 2012b: 37 (as
<taxonomicName id="4C7E3589FF0EFF28FA08FEAC3072FA09" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FA08FEAC3072FA09" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]);
<bibRefCitation id="EFEF33FBFF0EFF28FED4FE8C3343FA0A" author="Dupuis, F." box="[348,575,1506,1530]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="197" pageNumber="196" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 110</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FDF8FE8C3399FA09" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[624,741,1506,1529]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FDF8FE8C3399FA09" box="[624,741,1506,1529]" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]); MHCN.
<emphasis id="B90A9218FF0EFF28FC11FE8F3557FA09" bold="true" box="[921,1067,1505,1529]" pageId="197" pageNumber="196">“Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FBBAFE8C3580FA0A" author="Arrow, G. J." bookContentInfo="124 p." box="[1074,1276,1506,1530]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="197" pageNumber="196" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 99</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0EFF28FA80FE8C3044F9EA" author="Blackwelder, R. E." journalOrPublisher="United States National Museum Bulletin" pageId="197" pageNumber="196" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 259</bibRefCitation>
.
<emphasis id="B90A9218FF0EFF28FEC9FD6F309DF9E9" bold="true" box="[321,481,1537,1561]" pageId="197" pageNumber="196">
<collectingCountry id="F3690E9AFF0EFF28FEC9FD6F30E9F9E9" box="[321,405,1537,1561]" name="Brazil" pageId="197" pageNumber="196">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FE6EFD6C33BEF9EA" author="Endrodi, S." box="[486,706,1538,1562]" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="197" pageNumber="196" pagination="37 - 86" part="23" refId="ref201221" refString="Endrodi, S. 1977 b. Monographie der Dynastinae (Coleoptera). 6. Tribus: Dynastini II. Acta Zoologica Academiae Scientiarum Hungaricae 23 (1 - 2): 37 - 86." title="Monographie der Dynastinae (Coleoptera). 6. Tribus: Dynastini II" type="journal article" year="1977">Endrödi 1977b: 41</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FD79FD6C32AAF9E9" authority="actaeon" box="[753,982,1538,1561]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="actaeon">
<emphasis id="B90A9218FF0EFF28FD79FD6C32AAF9E9" box="[753,982,1538,1561]" italics="true" pageId="197" pageNumber="196">M. actaeon actaeon</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF0EFF28FB8BFD6C35B3F9E9" box="[1027,1231,1538,1561]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="janus">
<emphasis id="B90A9218FF0EFF28FB8BFD6C35B3F9E9" box="[1027,1231,1538,1561]" italics="true" pageId="197" pageNumber="196">M. actaeon janus</emphasis>
</taxonomicName>
), 1985a: 637, 642 (as
<taxonomicName id="4C7E3589FF0EFF28FF62FD4C30AEF9C9" authority="actaeon" box="[234,466,1570,1593]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="actaeon">
<emphasis id="B90A9218FF0EFF28FF62FD4C30AEF9C9" box="[234,466,1570,1593]" italics="true" pageId="197" pageNumber="196">M. actaeon actaeon</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF0EFF28FD89FD4C33ACF9C9" box="[513,720,1570,1593]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="janus">
<emphasis id="B90A9218FF0EFF28FD89FD4C33ACF9C9" box="[513,720,1570,1593]" italics="true" pageId="197" pageNumber="196">M. actaeon janus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FD6EFD4C32AFF9CA" author="Lachaume" box="[742,979,1570,1594]" firstAuthor="Lachaume" pageId="197" pageNumber="196" pagination="1 - 85" refId="ref211158" refString="Lachaume, G. 1985. Dynastini 1. Dynastes - Megasoma - Golofa. Les Coleopteres du Monde 5: 1 - 85." type="journal article" year="1985">Lachaume 1985: 34</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FB8DFD4C3591F9C9" authority="actaeon" box="[1029,1261,1570,1593]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="actaeon">
<emphasis id="B90A9218FF0EFF28FB8DFD4C3591F9C9" box="[1029,1261,1570,1593]" italics="true" pageId="197" pageNumber="196">M. actaeon actaeon</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF0EFF28FA94FD4C3078F9A9" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="janus">
<emphasis id="B90A9218FF0EFF28FA94FD4C3078F9A9" italics="true" pageId="197" pageNumber="196">M. actaeon janus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FE93FD2C33DBF9AA" author="Andreazze, R. &amp; C. R. V. da Fonseca" box="[283,679,1602,1626]" journalOrPublisher="Acta Amazonica" pageId="197" pageNumber="196" pagination="59 - 66" part="28" refId="ref190842" refString="Andreazze, R., and C. R. V. da Fonseca. 1998. Dinastineos (Coleoptera, Scarabaeoidea, Melolonthidae) em uma area de terra firme na Amazonia central, Brasil. Acta Amazonica 28 (1): 59 - 66." title="Dinastineos (Coleoptera, Scarabaeoidea, Melolonthidae) em uma area de terra firme na Amazonia central, Brasil" type="journal article" year="1998">Andreazze and Fonseca 1998: 62</bibRefCitation>
(NA);
<bibRefCitation id="EFEF33FBFF0EFF28FD7FFD2C3527F9A9" author="Silvestre and Arnaud" box="[759,1115,1602,1626]" firstAuthor="Silvestre" pageId="197" pageNumber="196" pagination="7 - 8" refId="ref225031" refString="Silvestre, G., and P. Arnaud. 2002. Description d'une nouvelle espece sud-americaine du genre Megasoma (Coleoptere Dynastidae). Besoiro 8: 7 - 8." type="journal article" year="2002">Silvestre and Arnaud 2002: 7</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FB04FD2C358BF9A9" baseAuthorityName="Felsche" baseAuthorityYear="1906" box="[1164,1271,1602,1625]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="janus">
<emphasis id="B90A9218FF0EFF28FB04FD2C358BF9A9" box="[1164,1271,1602,1625]" italics="true" pageId="197" pageNumber="196">M. janus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FA85FD2C31A1F98A" author="Nagai" firstAuthor="Nagai" pageId="197" pageNumber="196" pagination="35 - 39" refId="ref216675" refString="Nagai, S. 2003. Four new subspecies of the genus Megasoma (Coleoptera, Scarabaeidae) from South America. Gekkan-Mushi 394: 35 - 39." type="journal article" year="2003">Nagai 2003: 39</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FE98FD0C30B2F98A" authorityName="Nagai" authorityYear="2003" box="[272,462,1634,1658]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="fujitai">
<emphasis id="B90A9218FF0EFF28FE98FD0C30B2F98A" box="[272,462,1634,1658]" italics="true" pageId="197" pageNumber="196">M. janus fujitai</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF0EFF28FD88FD0C33CBF989" authority="janus" box="[512,695,1634,1657]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="janus">
<emphasis id="B90A9218FF0EFF28FD88FD0C33CBF989" box="[512,695,1634,1657]" italics="true" pageId="197" pageNumber="196">M. janus janus</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF0EFF28FD60FD0C3287F989" box="[744,1019,1634,1657]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="ramirezorum">
<emphasis id="B90A9218FF0EFF28FD60FD0C3287F989" box="[744,1019,1634,1657]" italics="true" pageId="197" pageNumber="196">M. janus ramirezorum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FB9BFD0C3587F98A" author="Ratcliffe" box="[1043,1275,1634,1658]" firstAuthor="Ratcliffe" pageId="197" pageNumber="196" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 461</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0EFF28FA8FFD0C31A1F96A" author="Moron" firstAuthor="Moron" pageId="197" pageNumber="196" pagination="29 - 36" refId="ref216197" refString="Moron, M. - A. 2005. A new species of Megasoma Kirby (Coleoptera: Scarabaeidae: Dynastinae) from Sinaloa, Mexico. Zootaxa 1037: 29 - 36." type="journal article" year="2005">Morón 2005: 30</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FE9DFDEC30A4F96A" authorityName="Nagai" authorityYear="2003" box="[277,472,1666,1690]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="fujitai">
<emphasis id="B90A9218FF0EFF28FE9DFDEC30A4F96A" box="[277,472,1666,1690]" italics="true" pageId="197" pageNumber="196">M. janus fujitai</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF0EFF28FD87FDEC33B7F969" authority="janus" box="[527,715,1666,1689]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="janus">
<emphasis id="B90A9218FF0EFF28FD87FDEC33B7F969" box="[527,715,1666,1689]" italics="true" pageId="197" pageNumber="196">M. janus janus</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FF0EFF28FD6DFDEC3207F96A" bold="true" box="[741,891,1666,1690]" pageId="197" pageNumber="196">
<collectingCountry id="F3690E9AFF0EFF28FD6DFDEC3209F96A" box="[741,885,1666,1690]" name="Venezuela" pageId="197" pageNumber="196">Venezuela</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FC0DFDEC3507F96A" author="Lachaume" box="[901,1147,1666,1690]" firstAuthor="Lachaume" pageId="197" pageNumber="196" pagination="1 - 85" refId="ref211158" refString="Lachaume, G. 1985. Dynastini 1. Dynastes - Megasoma - Golofa. Les Coleopteres du Monde 5: 1 - 85." type="journal article" year="1985">Lachaume 1985: 34</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0EFF28FB02FDEC343BF96A" author="Nagai" box="[1162,1351,1666,1690]" firstAuthor="Nagai" pageId="197" pageNumber="196" pagination="35 - 39" refId="ref216675" refString="Nagai, S. 2003. Four new subspecies of the genus Megasoma (Coleoptera, Scarabaeidae) from South America. Gekkan-Mushi 394: 35 - 39." type="journal article" year="2003">Nagai 2003: 39</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0EFF28FA08FDEC30D7F949" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="ramirezorum">
<emphasis id="B90A9218FF0EFF28FA08FDEC30D7F949" italics="true" pageId="197" pageNumber="196">M. janus ramirezorum</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FF0EFF28FE4AFDCF3366F949" bold="true" box="[450,538,1697,1721]" pageId="197" pageNumber="196">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF0EFF28FDA9FDCC326CF94A" author="Linnaeus" box="[545,784,1698,1722]" firstAuthor="Linnaeus" pageId="197" pageNumber="196" refId="ref212788" refString="Linnaeus, C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii; Stockholm. 823 p." type="book" year="1758">Linnaeus 1758: 345</bibRefCitation>
(America - as
<taxonomicName id="4C7E3589FF0EFF28FC37FDCF35CCF949" authorityName="Linnaeus" authorityYear="1758" box="[959,1200,1697,1721]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="actaeon">
<emphasis id="B90A9218FF0EFF28FC37FDCF35CCF949" box="[959,1200,1697,1721]" italics="true" pageId="197" pageNumber="196">Scarabaeus actaeon</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FB4FFDCC34DCF94A" author="Felsche, C." box="[1223,1440,1698,1722]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="197" pageNumber="196" pagination="349 - 352" part="1906" refId="ref202252" refString="Felsche, C. 1906. Synonymische Bemerkungen uber einige Scarabaeiden aus der Tribus der Dynastini und Beschreibung einer neuen Art. Deutsche Entomologische Zeitschrift 1906 (2): 349 - 352." title="Synonymische Bemerkungen uber einige Scarabaeiden aus der Tribus der Dynastini und Beschreibung einer neuen Art" type="journal article" year="1906">Felsche 1906: 352</bibRefCitation>
(Paraguay - as
<taxonomicName id="4C7E3589FF0EFF28FEFDFDAF332BF929" authorityName="Felsche" authorityYear="1906" box="[373,599,1729,1753]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="janus">
<emphasis id="B90A9218FF0EFF28FEFDFDAF332BF929" box="[373,599,1729,1753]" italics="true" pageId="197" pageNumber="196">Megalosoma janus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FDE5FDAC323BF92A" author="Hohne, W." box="[621,839,1730,1754]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="197" pageNumber="196" pagination="252 - 255" part="1923" refId="ref208127" refString="Hohne, W. 1923 a. Neue Dynastiden (Col.). Deutsche Entomologische Zeitschrift 1923 (3): 252 - 255." title="Neue Dynastiden (Col.)" type="journal article" year="1923">Höhne 1923a: 254</bibRefCitation>
(Argentina -
<taxonomicName id="4C7E3589FF0EFF28FC6FFDAC359AF929" box="[999,1254,1730,1753]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="argentinum">
<emphasis id="B90A9218FF0EFF28FC6FFDAC359AF929" box="[999,1254,1730,1753]" italics="true" pageId="197" pageNumber="196">M. janus argentinum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FB74FDAC31A1F90A" author="Arrow, G. J." bookContentInfo="124 p." editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="197" pageNumber="196" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 99</bibRefCitation>
(Peru - as
<taxonomicName id="4C7E3589FF0EFF28FED6FD8C309DF909" authorityName="Linnaeus" authorityYear="1758" box="[350,481,1762,1785]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="actaeon">
<emphasis id="B90A9218FF0EFF28FED6FD8C309DF909" box="[350,481,1762,1785]" italics="true" pageId="197" pageNumber="196">M. actaeon</emphasis>
</taxonomicName>
; Argentina, Paraguay - as
<taxonomicName id="4C7E3589FF0EFF28FCADFD8C32ECF909" authorityName="Felsche" authorityYear="1906" box="[805,912,1762,1785]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="janus">
<emphasis id="B90A9218FF0EFF28FCADFD8C32ECF909" box="[805,912,1762,1785]" italics="true" pageId="197" pageNumber="196">M. janus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FC2EFD8C350BF90A" author="Endrodi, S." box="[934,1143,1762,1786]" journalOrPublisher="Folia Entomologica Hungarica" pageId="197" pageNumber="196" pagination="64 - 73" part="6" refId="ref200487" refString="Endrodi, S. 1941. Die Dynastiden-Gattung Megasoma Kirby (Coleopt.). Folia Entomologica Hungarica 6 (3 - 4): 64 - 73." title="Die Dynastiden-Gattung Megasoma Kirby (Coleopt.)" type="journal article" year="1941">Endrödi 1941: 69</bibRefCitation>
, 70 (Peru? - as
<taxonomicName id="4C7E3589FF0EFF28FAA4FD8C34DCF909" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1324,1440,1762,1785]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="acteon">
<emphasis id="B90A9218FF0EFF28FAA4FD8C34DCF909" box="[1324,1440,1762,1785]" italics="true" pageId="197" pageNumber="196">M. acteon</emphasis>
</taxonomicName>
[sic]; Argentina - as
<taxonomicName id="4C7E3589FF0EFF28FE39FC6C33D3F8E9" box="[433,687,1794,1817]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="argentinum">
<emphasis id="B90A9218FF0EFF28FE39FC6C33D3F8E9" box="[433,687,1794,1817]" italics="true" pageId="197" pageNumber="196">M. janus argentinum</emphasis>
</taxonomicName>
; Paraguay - as
<taxonomicName id="4C7E3589FF0EFF28FCE0FC6C3560F8E9" authority="janus" box="[872,1052,1794,1817]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="janus">
<emphasis id="B90A9218FF0EFF28FCE0FC6C3560F8E9" box="[872,1052,1794,1817]" italics="true" pageId="197" pageNumber="196">M. janus janus</emphasis>
</taxonomicName>
), 1977b: 41 (Colombia, Peru - as
<taxonomicName id="4C7E3589FF0EFF28FF48FC4C30D6F8C9" authority="actaeon" box="[192,426,1826,1849]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="actaeon">
<emphasis id="B90A9218FF0EFF28FF48FC4C30D6F8C9" box="[192,426,1826,1849]" italics="true" pageId="197" pageNumber="196">M. actaeon actaeon</emphasis>
</taxonomicName>
; Paraguay - as
<taxonomicName id="4C7E3589FF0EFF28FDE2FC4C3246F8C9" box="[618,826,1826,1849]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="janus">
<emphasis id="B90A9218FF0EFF28FDE2FC4C3246F8C9" box="[618,826,1826,1849]" italics="true" pageId="197" pageNumber="196">M. actaeon janus</emphasis>
</taxonomicName>
), 1985a: 637, 642 (Colombia, Peru - as
<taxonomicName id="4C7E3589FF0EFF28FA93FC4C3061F8A9" authority="actaeon" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="actaeon">
<emphasis id="B90A9218FF0EFF28FA93FC4C3061F8A9" italics="true" pageId="197" pageNumber="196">M. actaeon actaeon</emphasis>
</taxonomicName>
; Paraguay - as
<taxonomicName id="4C7E3589FF0EFF28FE6FFC2C33C1F8A9" box="[487,701,1858,1881]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="janus">
<emphasis id="B90A9218FF0EFF28FE6FFC2C33C1F8A9" box="[487,701,1858,1881]" italics="true" pageId="197" pageNumber="196">M. actaeon janus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FD5FFC2C328EF8AA" author="Blackwelder, R. E." box="[727,1010,1858,1882]" journalOrPublisher="United States National Museum Bulletin" pageId="197" pageNumber="196" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 259</bibRefCitation>
(Peru - as
<taxonomicName id="4C7E3589FF0EFF28FB0BFC2C3477F8A9" authorityName="Linnaeus" authorityYear="1758" box="[1155,1291,1858,1881]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="actaeon">
<emphasis id="B90A9218FF0EFF28FB0BFC2C3477F8A9" box="[1155,1291,1858,1881]" italics="true" pageId="197" pageNumber="196">M. actaeon</emphasis>
</taxonomicName>
; Argentina, Paraguay - as
<taxonomicName id="4C7E3589FF0EFF28FEE5FC0C30ABF889" authorityName="Felsche" authorityYear="1906" box="[365,471,1890,1913]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="species" species="janus">
<emphasis id="B90A9218FF0EFF28FEE5FC0C30ABF889" box="[365,471,1890,1913]" italics="true" pageId="197" pageNumber="196">M. janus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0EFF28FE65FC0C33A5F88A" author="Lachaume" box="[493,729,1890,1914]" firstAuthor="Lachaume" pageId="197" pageNumber="196" pagination="1 - 85" refId="ref211158" refString="Lachaume, G. 1985. Dynastini 1. Dynastes - Megasoma - Golofa. Les Coleopteres du Monde 5: 1 - 85." type="journal article" year="1985">Lachaume 1985: 34</bibRefCitation>
(Bolivia, Colombia, Ecuador, Peru - as
<taxonomicName id="4C7E3589FF0EFF28FB3AFC0C34E5F889" authority="actaeon" box="[1202,1433,1890,1913]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="196" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="actaeon">
<emphasis id="B90A9218FF0EFF28FB3AFC0C34E5F889" box="[1202,1433,1890,1913]" italics="true" pageId="197" pageNumber="196">M. actaeon actaeon</emphasis>
</taxonomicName>
; Argentina, Bolivia, Colombia, Ecuador, Paraguay - as
<taxonomicName id="4C7E3589FF0DFF2BFCDDFBB43558FF01" box="[853,1060,218,241]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="subSpecies" species="actaeon" subSpecies="janus">
<emphasis id="B90A9218FF0DFF2BFCDDFBB43558FF01" box="[853,1060,218,241]" italics="true" pageId="198" pageNumber="197">M. actaeon janus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0DFF2BFBB3FBB734DCFF01" author="Silvestre and Arnaud" box="[1083,1440,217,242]" firstAuthor="Silvestre" pageId="198" pageNumber="197" pagination="7 - 8" refId="ref225031" refString="Silvestre, G., and P. Arnaud. 2002. Description d'une nouvelle espece sud-americaine du genre Megasoma (Coleoptere Dynastidae). Besoiro 8: 7 - 8." type="journal article" year="2002">Silvestre and Arnaud 2002: 7</bibRefCitation>
(Argentina, Bolivia, Paraguay - as
<taxonomicName id="4C7E3589FF0DFF2BFDE2FB9433AAFEE1" authorityName="Felsche" authorityYear="1906" box="[618,726,250,273]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="species" species="janus">
<emphasis id="B90A9218FF0DFF2BFDE2FB9433AAFEE1" box="[618,726,250,273]" italics="true" pageId="198" pageNumber="197">M. janus</emphasis>
</taxonomicName>
; Colombia, Ecuador - as
<taxonomicName id="4C7E3589FF0DFF2BFB80FB9435ACFEE1" authorityName="Silvestre and Arnaud" authorityYear="2002" box="[1032,1232,250,273]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="species" species="ramirezorum">
<emphasis id="B90A9218FF0DFF2BFB80FB9435ACFEE1" box="[1032,1232,250,273]" italics="true" pageId="198" pageNumber="197">M. ramirezorum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0DFF2BFB6FFB9734DCFEE2" author="Nagai" box="[1255,1440,249,274]" firstAuthor="Nagai" pageId="198" pageNumber="197" pagination="35 - 39" refId="ref216675" refString="Nagai, S. 2003. Four new subspecies of the genus Megasoma (Coleoptera, Scarabaeidae) from South America. Gekkan-Mushi 394: 35 - 39." type="journal article" year="2003">Nagai 2003: 39</bibRefCitation>
(Argentina, Bolivia, Paraguay - as
<taxonomicName id="4C7E3589FF0DFF2BFDD7FA74326EFEC1" authority="janus" box="[607,786,282,305]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="janus">
<emphasis id="B90A9218FF0DFF2BFDD7FA74326EFEC1" box="[607,786,282,305]" italics="true" pageId="198" pageNumber="197">M. janus janus</emphasis>
</taxonomicName>
; Colombia, Ecuador, Peru - as
<taxonomicName id="4C7E3589FF0DFF2BFB09FA7434ECFEC1" box="[1153,1424,282,305]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="ramirezorum">
<emphasis id="B90A9218FF0DFF2BFB09FA7434ECFEC1" box="[1153,1424,282,305]" italics="true" pageId="198" pageNumber="197">M. janus ramirezorum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0DFF2BFF48FA5730D7FEA1" author="Ratcliffe" box="[192,427,313,338]" firstAuthor="Ratcliffe" pageId="198" pageNumber="197" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 461</bibRefCitation>
(Colombia, Panama, Paraguay, Peru);
<bibRefCitation id="EFEF33FBFF0DFF2BFC11FA543521FEA2" author="Moron" box="[921,1117,313,338]" firstAuthor="Moron" pageId="198" pageNumber="197" pagination="29 - 36" refId="ref216197" refString="Moron, M. - A. 2005. A new species of Megasoma Kirby (Coleoptera: Scarabaeidae: Dynastinae) from Sinaloa, Mexico. Zootaxa 1037: 29 - 36." type="journal article" year="2005">Morón 2005: 30</bibRefCitation>
(BoliviaPanama - as
<taxonomicName id="4C7E3589FF0DFF2BFA08FA543067FE81" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="species" species="actaeon">
<emphasis id="B90A9218FF0DFF2BFA08FA543067FE81" italics="true" pageId="198" pageNumber="197">M. actaeon</emphasis>
</taxonomicName>
; Argentina, Bolivia, Paraguay - as
<taxonomicName id="4C7E3589FF0DFF2BFD3EFA343218FE81" authority="janus" box="[694,868,346,369]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="janus">
<emphasis id="B90A9218FF0DFF2BFD3EFA343218FE81" box="[694,868,346,369]" italics="true" pageId="198" pageNumber="197">M. janus janus</emphasis>
</taxonomicName>
; Colombia, Ecuador - as
<taxonomicName id="4C7E3589FF0DFF2BFB0EFA3434ECFE81" box="[1158,1424,346,369]" class="Insecta" family="Scarabaeidae" genus="Megalosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="ramirezorum">
<emphasis id="B90A9218FF0DFF2BFB0EFA3434ECFE81" box="[1158,1424,346,369]" italics="true" pageId="198" pageNumber="197">M. janus ramirezorum</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC14E0AFF0DFF2BFF48FAEF33F9FE09" blockId="198.[192,1440,385,505]" pageId="198" pageNumber="197">
<emphasis id="B90A9218FF0DFF2BFF48FAEF305CFE69" bold="true" box="[192,288,385,409]" pageId="198" pageNumber="197">Note 1.</emphasis>
Opinions vary as to whether
<taxonomicName id="4C7E3589FF0DFF2BFD07FAEC3222FE69" baseAuthorityName="Felsche" baseAuthorityYear="1906" box="[655,862,386,409]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="species" species="janus">
<emphasis id="B90A9218FF0DFF2BFD07FAEC3222FE69" box="[655,862,386,409]" italics="true" pageId="198" pageNumber="197">Megasoma janus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7E3589FF0DFF2BFCE5FAEC350DFE69" authorityName="Hohne" authorityYear="1923" box="[877,1137,386,409]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="argentinum">
<emphasis id="B90A9218FF0DFF2BFCE5FAEC350DFE69" box="[877,1137,386,409]" italics="true" pageId="198" pageNumber="197">M. janus argentinum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7E3589FF0DFF2BFB09FAEC343CFE69" authorityName="Nagai" authorityYear="2003" box="[1153,1344,385,409]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="subSpecies" species="janus" subSpecies="fujitai">
<emphasis id="B90A9218FF0DFF2BFB09FAEC343CFE69" box="[1153,1344,385,409]" italics="true" pageId="198" pageNumber="197">M. janus fujitai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E3589FF0DFF2BFA08FAEC301CFE49" authorityName="Silvestre and Arnaud" authorityYear="2002" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="species" species="ramirezorum">
<emphasis id="B90A9218FF0DFF2BFA08FAEC301CFE49" italics="true" pageId="198" pageNumber="197">M. ramirezorum</emphasis>
</taxonomicName>
are valid taxa or synonyms of
<taxonomicName id="4C7E3589FF0DFF2BFD59FACC3228FE49" authorityName="Linnaeus" authorityYear="1758" box="[721,852,418,441]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="species" species="actaeon">
<emphasis id="B90A9218FF0DFF2BFD59FACC3228FE49" box="[721,852,418,441]" italics="true" pageId="198" pageNumber="197">M. actaeon</emphasis>
</taxonomicName>
. Because differences are subtle, we merely mention these taxa here and take no position regarding the validity of these names. See
<bibRefCitation id="EFEF33FBFF0DFF2BFB47FAAC34DCFE2A" author="Moron" box="[1231,1440,449,474]" firstAuthor="Moron" pageId="198" pageNumber="197" pagination="29 - 36" refId="ref216197" refString="Moron, M. - A. 2005. A new species of Megasoma Kirby (Coleoptera: Scarabaeidae: Dynastinae) from Sinaloa, Mexico. Zootaxa 1037: 29 - 36." type="journal article" year="2005">Morón (2005: 30)</bibRefCitation>
for a rough overview of distributions.
</paragraph>
<paragraph id="8BC14E0AFF0DFF2BFF48F96735E6FCF1" blockId="198.[192,1441,521,769]" pageId="198" pageNumber="197">
<emphasis id="B90A9218FF0DFF2BFF48F9673062FDD1" bold="true" box="[192,286,521,545]" pageId="198" pageNumber="197">Note 2.</emphasis>
Five papers (
<bibRefCitation id="EFEF33FBFF0DFF2BFE49F9673326FDD1" author="Prandi" box="[449,602,521,545]" firstAuthor="Prandi" pageId="198" pageNumber="197" pagination="525 - 548" refId="ref220175" refString="Prandi, M. 2016. Una nuova specie di Megasoma Kirby dal Brasile (Coleoptera, Scarabaeidae, Dynastinae). Giornale italiano di Entomologia 14 (61): 525 - 548." type="journal article" year="2016">Prandi 2016</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF0DFF2BFDECF96433CDFDD2" author="Prandi" box="[612,689,522,546]" firstAuthor="Prandi" pageId="198" pageNumber="197" pagination="2 - 12" refId="ref220210" refString="Prandi, M. 2018 a. Synopsis of the species Megasoma actaeon (Linnaeus, 1758) (Coleoptera, Scarabaeidae, Dynastinae). Part 1. Splitting actaeon beetle. Gekkan Mushi 571: 2 - 12." type="journal article" year="2018" yearSuffix="a">2018a</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF0DFF2BFD34F9673274FDD1" author="Prandi" box="[700,776,521,545]" firstAuthor="Prandi" pageId="198" pageNumber="197" pagination="49 - 53" refId="ref220251" refString="Prandi, M. 2018 b. Synopsis of the species Megasoma actaeon (Linnaeus, 1758) (Coleoptera, Scarabaeidae, Dynastinae). Part II. Describing a new species. Gekkan Mushi 574: 49 - 53." type="journal article" year="2018" yearSuffix="b">2018b</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF0DFF2BFC9BF964322DFDD2" author="Prandi, M." box="[787,849,522,546]" journalOrPublisher="Gekkan Mushi" pageId="198" pageNumber="269" pagination="1 - 5" part="575" refId="ref220293" title="Synopsis of the species Megasoma actaeon (Linnaeus, 1758) (Coleoptera, Scarabaeidae, Dynastinae). Part III. Reviewing taxa" type="journal article" year="2019">2019</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0DFF2BFCD4F9643590FDD2" author="Van Meenen and Schouteet" box="[860,1260,521,546]" firstAuthor="Van Meenen" pageId="198" pageNumber="197" refId="ref228175" refString="Van Meenen, J., and T. Schouteet. 2018. Revision of the Megasoma actaeon (Linnaeus, 1758), janus Felsche, 1906, and ramirezorum Silvestre &amp; Arnaud, 2002 complex: description and review. Lambillionea; Chenee, Belgium. 80 p." type="book" year="2018">Van Meenen and Schouteet 2018</bibRefCitation>
) on
<taxonomicName id="4C7E3589FF0DFF2BFA96F9643061FDB1" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="197" phylum="Arthropoda" rank="species" species="actaeon">
<emphasis id="B90A9218FF0DFF2BFA96F9643061FDB1" italics="true" pageId="198" pageNumber="197">Megasoma actaeon</emphasis>
</taxonomicName>
have recently been published, all of them dealing with its purported subspecies, with some new, closely related, taxa being described. It appears these papers are partially overlapping, with at least one taxon (found outside the research area) being newly described twice. We consider the presented characteristics between all these species, subspecies and forms rather minimal given their variation within the newly defined taxonomic bounderies, especially as no genetic evidence is produced. Although the distribution of a taxon described in
<bibRefCitation id="EFEF33FBFF0DFF2BFD12F9A73253FD11" author="Prandi" box="[666,815,713,737]" firstAuthor="Prandi" pageId="198" pageNumber="197" pagination="525 - 548" refId="ref220175" refString="Prandi, M. 2016. Una nuova specie di Megasoma Kirby dal Brasile (Coleoptera, Scarabaeidae, Dynastinae). Giornale italiano di Entomologia 14 (61): 525 - 548." type="journal article" year="2016">Prandi 2016</bibRefCitation>
is within the research area, we have refrained from including it in this checklist until more solid evidence for its validity is provided.
</paragraph>
</subSubSection>
</treatment>
</document>