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<treatment id="03D7FF1CFF03FF25FF48F95C35C1FA28" ID-DOI="http://doi.org/10.5281/zenodo.3682337" ID-GBIF-Taxon="162246832" ID-Zenodo-Dep="3682337" LSID="urn:lsid:plazi:treatment:03D7FF1CFF03FF25FF48F95C35C1FA28" httpUri="http://treatment.plazi.org/id/03D7FF1CFF03FF25FF48F95C35C1FA28" lastPageNumber="199" pageId="200" pageNumber="199">
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<paragraph id="8BC14E0AFF03FF25FF48F95C3334FDB9" blockId="200.[192,584,561,585]" box="[192,584,561,585]" pageId="200" pageNumber="199">
<heading id="D089F966FF03FF25FF48F95C3334FDB9" bold="true" box="[192,584,561,585]" fontSize="10" level="2" pageId="200" pageNumber="199" reason="3">
<taxonomicName id="4C7E3589FF03FF25FF48F95C3334FDB9" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[192,584,561,585]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<emphasis id="B90A9218FF03FF25FF48F95C3334FDB9" bold="true" box="[192,584,561,585]" pageId="200" pageNumber="199">
<emphasis id="B90A9218FF03FF25FF48F95C3027FDB9" bold="true" box="[192,347,562,585]" italics="true" pageId="200" pageNumber="199">Enema pan</emphasis>
(
<bibRefCitation id="EFEF33FBFF03FF25FEE3F95F3343FDB9" author="Fabricius, J. C." bookContentInfo="832 p." box="[363,575,561,585]" journalOrPublisher="Kortii; Flensburg and Leipzig, Germany" pageId="200" pageNumber="199" refId="ref201756" refString="Fabricius, J. C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus. Kortii; Flensburg and Leipzig, Germany. 832 p." title="Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus" type="book" year="1775">Fabricius, 1775</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3641D81FF03FF25FF60F9363293FD58" pageId="200" pageNumber="199" type="reference_group">
<paragraph id="8BC14E0AFF03FF25FF60F93633CCFD9E" blockId="200.[232,1007,600,680]" box="[232,688,600,622]" pageId="200" pageNumber="199">
<treatmentCitationGroup id="AB6E6924FF03FF25FF60F93633CCFD9E" box="[232,688,600,622]" pageId="200" pageNumber="199">
<taxonomicName id="4C7E3589FF03FF25FF60F93633CCFD9E" ID-CoL="4V2CK" authority="Fabricius 1775: 5" authorityName="Fabricius" authorityPageNumber="5" authorityYear="1775" box="[232,688,600,622]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<treatmentCitation id="0ADF681BFF03FF25FF60F9363320FD9E" author="Fabricius, J. C." box="[232,604,600,622]" page="5" pageId="200" pageNumber="199" year="1775">
<emphasis id="B90A9218FF03FF25FF60F93630E6FD9D" box="[232,410,600,621]" italics="true" pageId="200" pageNumber="199">Scarabaeus pan</emphasis>
<bibRefCitation id="EFEF33FBFF03FF25FE29F9363320FD9E" author="Fabricius, J. C." bookContentInfo="832 p." box="[417,604,600,622]" journalOrPublisher="Kortii; Flensburg and Leipzig, Germany" pageId="200" pageNumber="199" refId="ref201756" refString="Fabricius, J. C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus. Kortii; Flensburg and Leipzig, Germany. 832 p." title="Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus" type="book" year="1775">Fabricius 1775: 5</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF03FF25FDE3F93633D4FD9E" box="[619,680,600,622]" name="India" pageId="200" pageNumber="199">India</collectingCountry>
)
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC14E0AFF03FF25FF60F91B3262FD7B" blockId="200.[232,1007,600,680]" box="[232,798,628,651]" pageId="200" pageNumber="199">
<treatmentCitationGroup id="AB6E6924FF03FF25FF60F91B3262FD7B" box="[232,798,628,651]" pageId="200" pageNumber="199">
=
<taxonomicName id="4C7E3589FF03FF25FF75F91A3262FD7B" ID-CoL="6XW7C" authority="Fabricius 1775: 5" authorityName="Fabricius" authorityPageNumber="5" authorityYear="1775" box="[253,798,628,651]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="chorinaeus">
<treatmentCitation id="0ADF681BFF03FF25FF75F91A33BDFD7B" author="Fabricius, J. C." box="[253,705,628,651]" page="5" pageId="200" pageNumber="199" year="1775">
<emphasis id="B90A9218FF03FF25FF75F91A3083FD79" box="[253,511,628,649]" italics="true" pageId="200" pageNumber="199">Scarabaeus chorinaeus</emphasis>
<bibRefCitation id="EFEF33FBFF03FF25FD8DF91B33BDFD7B" author="Fabricius, J. C." bookContentInfo="832 p." box="[517,705,629,651]" journalOrPublisher="Kortii; Flensburg and Leipzig, Germany" pageId="200" pageNumber="199" refId="ref201756" refString="Fabricius, J. C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus. Kortii; Flensburg and Leipzig, Germany. 832 p." title="Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus" type="book" year="1775">Fabricius 1775: 5</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF03FF25FD58F91B326BFD7B" box="[720,791,629,651]" name="Brazil" pageId="200" pageNumber="199">Brazil</collectingCountry>
)
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC14E0AFF03FF25FF60F9FC3293FD58" blockId="200.[232,1007,600,680]" box="[232,1007,657,680]" pageId="200" pageNumber="199">
<treatmentCitationGroup id="AB6E6924FF03FF25FF60F9FC3293FD58" box="[232,1007,657,680]" pageId="200" pageNumber="199">
=
<treatmentCitation id="0ADF681BFF03FF25FF75F9FC339DFD57" author="Burmeister, H. C. C." box="[253,737,657,679]" page="234" pageId="200" pageNumber="199" year="1847">
<taxonomicName id="4C7E3589FF03FF25FF75F9FC339DFD57" authority="Burmeister 1847: 234" authorityName="Burmeister" authorityPageNumber="234" authorityYear="1847" box="[253,737,657,679]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="infundibulum">
<emphasis id="B90A9218FF03FF25FF75F9FC3097FD56" box="[253,491,657,679]" italics="true" pageId="200" pageNumber="199">Enema infundibulum</emphasis>
<bibRefCitation id="EFEF33FBFF03FF25FE79F9FF339DFD57" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[497,737,657,679]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="200" pageNumber="199" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 234</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF03FF25FD67F9FF3249FD57" box="[751,821,657,679]" name="Brazil" pageId="200" pageNumber="199">Brazil</collectingCountry>
-
<typeStatus id="54C5F0A8FF03FF25FCC2F9FC3294FD58" box="[842,1000,658,680]" pageId="200" pageNumber="199" type="lectotype">LECTOTYPE</typeStatus>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF03FF25FF48F9D935C1FA28" pageId="200" pageNumber="199" type="distribution">
<paragraph id="8BC14E0AFF03FF25FF48F9D93227FB00" blockId="200.[192,1440,695,1264]" pageId="200" pageNumber="199">
<emphasis id="B90A9218FF03FF25FF48F9D93090FD3F" bold="true" box="[192,492,695,719]" pageId="200" pageNumber="199">
Distribution.
<collectingCountry id="F3690E9AFF03FF25FEF1F9D9309AFD3F" box="[377,486,695,719]" name="Guyana" pageId="200" pageNumber="199">Guyana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF03FF25FE7AF9D633A6FD20" author="Erichson, W. F." box="[498,730,696,720]" editor="R. Schomburgk" journalOrPublisher="J. J. Weber; Leipzig, Germany" pageId="200" pageNumber="199" pagination="553 - 1260" refId="ref201522" refString="Erichson, W. F. 1848. Insecten. p. 553 - 617. In: R. Schomburgk. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil. J. J. Weber; Leipzig, Germany. p. 553 - 1260." title="Insecten" type="book chapter" volumeTitle="Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil" year="1848">Erichson 1848: 561</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF03FF25FC82F9D632AAFD20" authorityName="Burmeister" authorityYear="1847" box="[778,982,696,720]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="infundibulum">
<emphasis id="B90A9218FF03FF25FC82F9D632AAFD20" box="[778,982,696,720]" italics="true" pageId="200" pageNumber="199">E. infundibulum</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FF03FF25FC65F9D635BFFD3F" bold="true" box="[1005,1219,695,720]" pageId="200" pageNumber="199">
<collectingCountry id="F3690E9AFF03FF25FC65F9D635C1FD3F" box="[1005,1213,695,720]" name="French Guiana" pageId="200" pageNumber="199">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF03FF25FB41F9D634E7FD20" author="Endrodi, S." box="[1225,1435,696,720]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="200" pageNumber="199" pagination="9 - 174" part="29" refId="ref201147" refString="Endrodi, S. 1976. Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae). Folia Entomologica Hungarica (N. S.) 29 (2): 9 - 174." title="Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae)" type="journal article" year="1976">Endrödi 1976: 32</bibRefCitation>
, 61 (as
<taxonomicName id="4C7E3589FF03FF25FE83F9B63024FD1F" authorityName="Fabricius" authorityYear="1775" box="[267,344,728,751]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<emphasis id="B90A9218FF03FF25FE83F9B63024FD1F" box="[267,344,728,751]" italics="true" pageId="200" pageNumber="199">E. pan</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF03FF25FE0DF9B633E5FD00" authorityName="Fabricius" authorityYear="1775" box="[389,665,728,752]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="chorinaeus">
<emphasis id="B90A9218FF03FF25FE0DF9B633E5FD00" box="[389,665,728,752]" italics="true" pageId="200" pageNumber="199">Scarabaeus chorinaeus</emphasis>
</taxonomicName>
(treated as a synonym of
<taxonomicName id="4C7E3589FF03FF25FC4DF9B635EAFD00" authorityName="Endrodi" authorityYear="1976" box="[965,1174,728,752]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="lycaon">
<emphasis id="B90A9218FF03FF25FC4DF9B635EAFD00" box="[965,1174,728,752]" italics="true" pageId="200" pageNumber="199">Megaceras lycaon</emphasis>
</taxonomicName>
(now
<taxonomicName id="4C7E3589FF03FF25FB53F9B63403FD00" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1243,1407,728,752]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="philoctetes">
<emphasis id="B90A9218FF03FF25FB53F9B63403FD00" box="[1243,1407,728,752]" italics="true" pageId="200" pageNumber="199">M. philoctetes</emphasis>
</taxonomicName>
))), 1985a: 579, 580;
<bibRefCitation id="EFEF33FBFF03FF25FE19F99633D6FCE0" author="Dechambre, R. - P." box="[401,682,760,784]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="200" pageNumber="199" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FD3FF99632D5FCE0" author="Lachaume" box="[695,937,760,784]" firstAuthor="Lachaume" pageId="200" pageNumber="199" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 46</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FC3EF99635ADFCE0" author="Brule, S. &amp; J. Touroult &amp; P. - H. Dalens" bookContentInfo="120 p." box="[950,1233,760,784]" journalOrPublisher="Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF" pageId="200" pageNumber="199" refId="ref195373" refString="Brule, S., J. Touroult, and P. - H. Dalens (coord.). 2011 a. Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane), 2009 - 2010. Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF. 120 p. + annexes." title="Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane)" type="book" year="2011">Brûlé et al. 2011a: 191</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FB56F996319CFCC0" author="Duranton, M." journalOrPublisher="Coleopteres, numero special" pageId="200" pageNumber="199" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 22</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FF66F87A33C5FCC0" author="Gasca-Alvarez, H. J. &amp; B. C. Ratcliffe" box="[238,697,788,816]" journalOrPublisher="Zootaxa" pageId="200" pageNumber="199" pagination="21 - 40" part="3090" refId="ref203834" refString="Gasca-Alvarez, H. J., and B. C. Ratcliffe. 2011. An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms. Zootaxa 3090: 21 - 40." title="An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms" type="journal article" year="2011">Gasca-Álvarez and Ratcliffe 2011: 25</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FD4FF87632E0FCC0" author="Ponchel" box="[711,924,792,816]" firstAuthor="Ponchel" pageId="200" pageNumber="199" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011: 61</bibRefCitation>
.
<emphasis id="B90A9218FF03FF25FC21F8793547FCDF" bold="true" box="[937,1083,791,815]" pageId="200" pageNumber="199">“Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFF03FF25FBCCF876342CFCC0" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[1092,1360,792,816]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="200" pageNumber="199" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 234</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF03FF25FA0DF8763015FCA0" authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="infundibulum">
<emphasis id="B90A9218FF03FF25FA0DF8763015FCA0" italics="true" pageId="200" pageNumber="199">E. infundibulum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF03FF25FEF6F85633F2FCA0" author="Blackwelder, R. E." box="[382,654,824,848]" journalOrPublisher="United States National Museum Bulletin" pageId="200" pageNumber="199" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 256</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF03FF25FD34F85632F8FCA0" authorityName="Burmeister" authorityYear="1847" box="[700,900,824,848]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="infundibulum">
<emphasis id="B90A9218FF03FF25FD34F85632F8FCA0" box="[700,900,824,848]" italics="true" pageId="200" pageNumber="199">E. infundibulum</emphasis>
</taxonomicName>
(treated as a synonym of
<taxonomicName id="4C7E3589FF03FF25FB39F85634FBFCBF" box="[1201,1415,824,847]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="variety" species="pan" variety="enema">
<emphasis id="B90A9218FF03FF25FB39F8563583FCBF" box="[1201,1279,824,847]" italics="true" pageId="200" pageNumber="199">E. pan</emphasis>
var.
<emphasis id="B90A9218FF03FF25FAB1F85634FBFCBF" box="[1337,1415,824,847]" italics="true" pageId="200" pageNumber="199">enema</emphasis>
</taxonomicName>
)).
<emphasis id="B90A9218FF03FF25FF48F839301CFC9F" bold="true" box="[192,352,855,879]" pageId="200" pageNumber="199">
<collectingCountry id="F3690E9AFF03FF25FF48F8393068FC9F" box="[192,276,855,879]" name="Brazil" pageId="200" pageNumber="199">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFF03FF25FEEDF8363349FC80" author="Fabricius, J. C." bookContentInfo="832 p." box="[357,565,856,880]" journalOrPublisher="Kortii; Flensburg and Leipzig, Germany" pageId="200" pageNumber="199" refId="ref201756" refString="Fabricius, J. C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus. Kortii; Flensburg and Leipzig, Germany. 832 p." title="Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus" type="book" year="1775">Fabricius 1775: 5</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF03FF25FDECF8363200FC80" authorityName="Fabricius" authorityYear="1775" box="[612,892,856,880]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="chorinaeus">
<emphasis id="B90A9218FF03FF25FDECF8363200FC80" box="[612,892,856,880]" italics="true" pageId="200" pageNumber="199">Scarabaeus chorinaeus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF03FF25FC19F83635EAFC80" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[913,1174,856,880]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="200" pageNumber="199" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 234</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF03FF25FB4DF83634ECFC80" authorityName="Burmeister" authorityYear="1847" box="[1221,1424,856,880]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="infundibulum">
<emphasis id="B90A9218FF03FF25FB4DF83634ECFC80" box="[1221,1424,856,880]" italics="true" pageId="200" pageNumber="199">E. infundibulum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF03FF25FF48F81630ECFC60" author="Arrow, G. J." bookContentInfo="124 p." box="[192,400,888,912]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="200" pageNumber="199" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 65</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FE16F81633C5FC60" author="Blackwelder, R. E." box="[414,697,888,912]" journalOrPublisher="United States National Museum Bulletin" pageId="200" pageNumber="199" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 256</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FD4FF81632E7FC60" author="Endrodi, S." box="[711,923,888,912]" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="200" pageNumber="199" pagination="11 - 17" part="19" refId="ref200988" refString="Endrodi, S. 1973. Designation von Lectotypen aus der Sammlung von H. Burmeister: Oryctini und Dynastini (Coleoptera: Melolonthidae). Acta Zoologica Academiae Scientiarum Hungaricae 19 (1 - 2): 11 - 17." title="Designation von Lectotypen aus der Sammlung von H. Burmeister: Oryctini und Dynastini (Coleoptera: Melolonthidae)" type="journal article" year="1973">Endrödi 1973: 15</bibRefCitation>
(designation
<typeStatus id="54C5F0A8FF03FF25FBCDF8163591FC60" box="[1093,1261,888,912]" pageId="200" pageNumber="199" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="4C7E3589FF03FF25FB7EF8163055FC40" authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="infundibulum">
<emphasis id="B90A9218FF03FF25FB7EF8163055FC40" italics="true" pageId="200" pageNumber="199">Enema infundibulum</emphasis>
</taxonomicName>
), 1976: 32, 61, 1985a: 579, 580;
<bibRefCitation id="EFEF33FBFF03FF25FD38F8F632BBFC40" author="Dechambre, R. - P." box="[688,967,920,944]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="200" pageNumber="199" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FC5BF8F635B8FC40" author="Lachaume" box="[979,1220,920,944]" firstAuthor="Lachaume" pageId="200" pageNumber="199" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 46</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF03FF25FB58F8FA302CFC20" author="Gasca-Alvarez, H. J. &amp; N. O. Aguiar" journalOrPublisher="Acta Zoologica Mexicana (N. S.)" pageId="200" pageNumber="199" pagination="221 - 224" part="24" refId="ref203674" refString="Gasca-Alvarez, H. J., and N. O. Aguiar. 2008. Primer registro de Amblyodus taurus Westwood, 1878 (Scarabaeidae: Dynastinae: Phileurini) en Brasil. Acta Zoologica Mexicana (N. S.) 24 (3): 221 - 224." title="Primer registro de Amblyodus taurus Westwood, 1878 (Scarabaeidae: Dynastinae: Phileurini) en Brasil" type="journal article" year="2008">Gasca-Álvarez et al. 2008: 13</bibRefCitation>
(NA);
<bibRefCitation id="EFEF33FBFF03FF25FE2CF8DA3205FC20" author="Gasca-Alvarez, H. J. &amp; C. R. V. da Fonseca" box="[420,889,948,976]" journalOrPublisher="Acta Amazonica" pageId="200" pageNumber="199" pagination="719 - 722" part="39" refId="ref203726" refString="Gasca-Alvarez, H. J., and C. R. V. da Fonseca. 2009. Oryctini (Coleoptera: Scarabaeidae: Dynastinae) da colecao de invertebrados do Instituto Nacional de Pesquisas da Amazonia (INPA), Manaus, Amazonas, Brasil. Acta Amazonica 39 (3): 719 - 722." title="Oryctini (Coleoptera: Scarabaeidae: Dynastinae) da colecao de invertebrados do Instituto Nacional de Pesquisas da Amazonia (INPA), Manaus, Amazonas, Brasil" type="journal article" year="2009">Gasca-Álvarez and Fonseca 2009: 721</bibRefCitation>
(NA);
<bibRefCitation id="EFEF33FBFF03FF25FC46F8DA34E0FC20" author="Gasca-Alvarez, H. J. &amp; B. C. Ratcliffe" box="[974,1436,948,976]" journalOrPublisher="Zootaxa" pageId="200" pageNumber="199" pagination="21 - 40" part="3090" refId="ref203834" refString="Gasca-Alvarez, H. J., and B. C. Ratcliffe. 2011. An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms. Zootaxa 3090: 21 - 40." title="An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms" type="journal article" year="2011">Gasca-Álvarez and Ratcliffe 2011: 25</bibRefCitation>
.
<emphasis id="B90A9218FF03FF25FF48F8B93064FC1F" bold="true" box="[192,280,983,1007]" pageId="200" pageNumber="199">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF03FF25FE97F8B6308EFC00" author="Fabricius, J. C." bookContentInfo="832 p." box="[287,498,984,1008]" journalOrPublisher="Kortii; Flensburg and Leipzig, Germany" pageId="200" pageNumber="199" refId="ref201756" refString="Fabricius, J. C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus. Kortii; Flensburg and Leipzig, Germany. 832 p." title="Systema entomologiae, sistens insectorum classes, ordines, genera, species, adjectis synonymis, locis, descriptionibus, observationibus" type="book" year="1775">Fabricius 1775: 5</bibRefCitation>
(
<collectingCountry id="F3690E9AFF03FF25FD89F8B6333FFC00" box="[513,579,984,1008]" name="India" pageId="200" pageNumber="199">India</collectingCountry>
- as
<taxonomicName id="4C7E3589FF03FF25FDF4F8B6323CFC1F" authorityName="Fabricius" authorityYear="1775" box="[636,832,984,1008]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<emphasis id="B90A9218FF03FF25FDF4F8B6323CFC1F" box="[636,832,984,1008]" italics="true" pageId="200" pageNumber="199">Scarabaeus pan</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF03FF25FCDEF8B63563FC00" author="Arrow, G. J." bookContentInfo="124 p." box="[854,1055,984,1008]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="200" pageNumber="199" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 65</bibRefCitation>
(
<collectingCountry id="F3690E9AFF03FF25FBA7F8B635DAFC00" box="[1071,1190,984,1008]" name="Colombia" pageId="200" pageNumber="199">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FB39F8B6344BFC00" box="[1201,1335,984,1008]" name="Costa Rica" pageId="200" pageNumber="199">Costa Rica</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FACBF8B634E0FC00" box="[1347,1436,984,1008]" name="Mexico" pageId="200" pageNumber="199">Mexico</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FF48F896303CFBE0" box="[192,320,1016,1040]" name="Nicaragua" pageId="200" pageNumber="199">Nicaragua</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FEC1F89630C3FBE0" box="[329,447,1016,1040]" name="Paraguay" pageId="200" pageNumber="199">Paraguay</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FE41F8963379FBE0" box="[457,517,1016,1040]" name="Peru" pageId="200" pageNumber="199">Peru</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF03FF25FD9FF896325DFBE0" author="Blackwelder, R. E." box="[535,801,1016,1040]" journalOrPublisher="United States National Museum Bulletin" pageId="200" pageNumber="199" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 256</bibRefCitation>
(
<collectingCountry id="F3690E9AFF03FF25FCA7F89632D7FBE0" box="[815,939,1016,1040]" name="Argentina" pageId="200" pageNumber="199">Argentina</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FC3CF896355BFBE0" box="[948,1063,1016,1040]" name="Colombia" pageId="200" pageNumber="199">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FBB9F89635CDFBE0" box="[1073,1201,1016,1040]" name="Costa Rica" pageId="200" pageNumber="199">Costa Rica</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FB33F896346EFBE0" box="[1211,1298,1016,1040]" name="Mexico" pageId="200" pageNumber="199">Mexico</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FA93F89634E7FBE0" box="[1307,1435,1016,1040]" name="Nicaragua" pageId="200" pageNumber="199">Nicaragua</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FF48FF763044FBC0" box="[192,312,1048,1072]" name="Paraguay" pageId="200" pageNumber="199">Paraguay</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FECAFF763003FBC0" box="[322,383,1048,1072]" name="Peru" pageId="200" pageNumber="199">Peru</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF03FF25FE1AFF76331EFBC0" author="Endrodi, S." box="[402,610,1048,1072]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="200" pageNumber="199" pagination="9 - 174" part="29" refId="ref201147" refString="Endrodi, S. 1976. Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae). Folia Entomologica Hungarica (N. S.) 29 (2): 9 - 174." title="Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae)" type="journal article" year="1976">Endrödi 1976: 32</bibRefCitation>
, 61 (
<collectingCountry id="F3690E9AFF03FF25FD1EFF763393FBC0" box="[662,751,1048,1072]" name="Bolivia" pageId="200" pageNumber="199">Bolivia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FD72FF76320CFBC0" box="[762,880,1048,1072]" name="Colombia" pageId="200" pageNumber="199">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FCF1FF76329DFBC0" box="[889,993,1048,1072]" name="Ecuador" pageId="200" pageNumber="199">Ecuador</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FC63FF76351FFBC0" box="[1003,1123,1048,1072]" name="Paraguay" pageId="200" pageNumber="199">Paraguay</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FBE6FF7635D5FBDF" box="[1134,1193,1048,1071]" name="Peru" pageId="200" pageNumber="199">Peru</collectingCountry>
- as
<taxonomicName id="4C7E3589FF03FF25FB54FF763456FBDF" authorityName="Fabricius" authorityYear="1775" box="[1244,1322,1048,1071]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<emphasis id="B90A9218FF03FF25FB54FF763456FBDF" box="[1244,1322,1048,1071]" italics="true" pageId="200" pageNumber="199">E. pan</emphasis>
</taxonomicName>
;
<collectingCountry id="F3690E9AFF03FF25FABFFF76340EFBDF" box="[1335,1394,1048,1071]" name="Peru" pageId="200" pageNumber="199">Peru</collectingCountry>
- as
<taxonomicName id="4C7E3589FF03FF25FF48FF5630A7FBA0" authorityName="Fabricius" authorityYear="1775" box="[192,475,1080,1104]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="chorinaeus">
<emphasis id="B90A9218FF03FF25FF48FF5630A7FBA0" box="[192,475,1080,1104]" italics="true" pageId="200" pageNumber="199">Scarabaeus chorinaeus</emphasis>
</taxonomicName>
(treated as a synonym of
<taxonomicName id="4C7E3589FF03FF25FC95FF563289FBA0" authorityName="Endrodi" authorityYear="1976" box="[797,1013,1080,1104]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="lycaon">
<emphasis id="B90A9218FF03FF25FC95FF563289FBA0" box="[797,1013,1080,1104]" italics="true" pageId="200" pageNumber="199">Megaceras lycaon</emphasis>
</taxonomicName>
(now
<taxonomicName id="4C7E3589FF03FF25FBC8FF563595FBA0" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1088,1257,1080,1104]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="philoctetes">
<emphasis id="B90A9218FF03FF25FBC8FF563595FBA0" box="[1088,1257,1080,1104]" italics="true" pageId="200" pageNumber="199">M. philoctetes</emphasis>
</taxonomicName>
))), 1985a: 579, 580 (
<collectingCountry id="F3690E9AFF03FF25FF73FF363029FB80" box="[251,341,1112,1136]" name="Bolivia" pageId="200" pageNumber="199">Bolivia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FEE8FF3630ABFB80" box="[352,471,1112,1136]" name="Colombia" pageId="200" pageNumber="199">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FE69FF363336FB80" box="[481,586,1112,1136]" name="Ecuador" pageId="200" pageNumber="199">Ecuador</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FDDDFF3633B2FB80" box="[597,718,1112,1136]" name="Paraguay" pageId="200" pageNumber="199">Paraguay</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FD51FF36326BFB80" box="[729,791,1112,1136]" name="Peru" pageId="200" pageNumber="199">Peru</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF03FF25FCA3FF363542FB80" author="Dechambre, R. - P." box="[811,1086,1112,1136]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="200" pageNumber="199" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
(
<collectingCountry id="F3690E9AFF03FF25FBC4FF3635DAFB80" box="[1100,1190,1112,1136]" name="Bolivia" pageId="200" pageNumber="199">Bolivia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FB39FF363454FB80" box="[1201,1320,1112,1136]" name="Colombia" pageId="200" pageNumber="199">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FABAFF3634E7FB80" box="[1330,1435,1112,1136]" name="Ecuador" pageId="200" pageNumber="199">Ecuador</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FF48FF163045FB60" box="[192,313,1144,1168]" name="Paraguay" pageId="200" pageNumber="199">Paraguay</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FECEFF1630F8FB60" box="[326,388,1144,1168]" name="Peru" pageId="200" pageNumber="199">Peru</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF03FF25FE11FF1633F4FB60" author="Lachaume" box="[409,648,1144,1168]" firstAuthor="Lachaume" pageId="200" pageNumber="199" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 46</bibRefCitation>
(
<collectingCountry id="F3690E9AFF03FF25FD10FF16338EFB60" box="[664,754,1144,1168]" name="Bolivia" pageId="200" pageNumber="199">Bolivia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FD77FF16320AFB60" box="[767,886,1144,1168]" name="Colombia" pageId="200" pageNumber="199">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FC09FF163296FB60" box="[897,1002,1144,1168]" name="Ecuador" pageId="200" pageNumber="199">Ecuador</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FC7FFF16350CFB60" box="[1015,1136,1144,1168]" name="Paraguay" pageId="200" pageNumber="199">Paraguay</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FBF4FF1635C6FB60" box="[1148,1210,1144,1168]" name="Peru" pageId="200" pageNumber="199">Peru</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF03FF25FB47FF1A3032FB40" author="Gasca-Alvarez, H. J. &amp; N. O. Aguiar" journalOrPublisher="Acta Zoologica Mexicana (N. S.)" pageId="200" pageNumber="199" pagination="221 - 224" part="24" refId="ref203674" refString="Gasca-Alvarez, H. J., and N. O. Aguiar. 2008. Primer registro de Amblyodus taurus Westwood, 1878 (Scarabaeidae: Dynastinae: Phileurini) en Brasil. Acta Zoologica Mexicana (N. S.) 24 (3): 221 - 224." title="Primer registro de Amblyodus taurus Westwood, 1878 (Scarabaeidae: Dynastinae: Phileurini) en Brasil" type="journal article" year="2008">Gasca-Álvarez et al. 2008: 13</bibRefCitation>
(Mexico–Paraguay);
<bibRefCitation id="EFEF33FBFF03FF25FDDBFFFA3566FB40" author="Gasca-Alvarez, H. J. &amp; B. C. Ratcliffe" box="[595,1050,1172,1200]" journalOrPublisher="Zootaxa" pageId="200" pageNumber="199" pagination="21 - 40" part="3090" refId="ref203834" refString="Gasca-Alvarez, H. J., and B. C. Ratcliffe. 2011. An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms. Zootaxa 3090: 21 - 40." title="An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms" type="journal article" year="2011">Gasca-Álvarez and Ratcliffe 2011: 25</bibRefCitation>
(
<collectingCountry id="F3690E9AFF03FF25FBA3FFF635F9FB40" box="[1067,1157,1176,1200]" name="Bolivia" pageId="200" pageNumber="199">Bolivia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FB1AFFF63475FB40" box="[1170,1289,1176,1200]" name="Colombia" pageId="200" pageNumber="199">Colombia</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FA9DFFF634E7FB40" box="[1301,1435,1176,1200]" name="Costa Rica" pageId="200" pageNumber="199">Costa Rica</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FF48FFD63055FB20" box="[192,297,1208,1232]" name="Ecuador" pageId="200" pageNumber="199">Ecuador</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FEBDFFD630CEFB20" box="[309,434,1208,1232]" name="Honduras" pageId="200" pageNumber="199">Honduras</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FE36FFD6336BFB20" box="[446,535,1208,1232]" name="Mexico" pageId="200" pageNumber="199">Mexico</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FDABFFD633DAFB20" box="[547,678,1208,1232]" name="Nicaragua" pageId="200" pageNumber="199">Nicaragua</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FD3AFFD63266FB3F" box="[690,794,1208,1231]" name="Panama" pageId="200" pageNumber="199">Panama</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FCAFFFD632DCFB20" box="[807,928,1208,1232]" name="Paraguay" pageId="200" pageNumber="199">Paraguay</collectingCountry>
,
<collectingCountry id="F3690E9AFF03FF25FC25FFD63297FB20" box="[941,1003,1208,1232]" name="Peru" pageId="200" pageNumber="199">Peru</collectingCountry>
);
<bibRefCitation id="EFEF33FBFF03FF25FB88FFD63456FB20" author="Ratcliffe" box="[1024,1322,1208,1232]" etAl="et al." firstAuthor="Ratcliffe" pageId="200" pageNumber="199" pagination="1 - 661" refId="ref222393" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 661." type="journal article" year="2013">Ratcliffe et al. 2013: 369</bibRefCitation>
(Mexico– South America);
<bibRefCitation id="EFEF33FBFF03FF25FE06FFB6330FFB00" author="Dupuis, F." box="[398,627,1240,1264]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="200" pageNumber="199" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 108</bibRefCitation>
(see
<taxonomicName id="4C7E3589FF03FF25FD39FFB63236FB00" authorityName="Chevrolat" authorityYear="1843" box="[689,842,1240,1264]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="endymion">
<emphasis id="B90A9218FF03FF25FD39FFB63236FB00" box="[689,842,1240,1264]" italics="true" pageId="200" pageNumber="199">E. endymion</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC14E0AFF03FF25FF48FE6E35C1FA28" blockId="200.[192,1440,1280,1496]" pageId="200" pageNumber="199">
<emphasis id="B90A9218FF03FF25FF48FE6E3074FAE8" bold="true" box="[192,264,1280,1304]" pageId="200" pageNumber="199">Note.</emphasis>
According to
<bibRefCitation id="EFEF33FBFF03FF25FE27FE6E33D4FAE8" author="Dupuis, F." box="[431,680,1280,1304]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="200" pageNumber="199" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis (2016b: 108)</bibRefCitation>
,
<taxonomicName id="4C7E3589FF03FF25FD3BFE6E323CFAE7" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[691,832,1280,1303]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<emphasis id="B90A9218FF03FF25FD3BFE6E323CFAE7" box="[691,832,1280,1303]" italics="true" pageId="200" pageNumber="199">Enema pan</emphasis>
</taxonomicName>
does not occur in
<collectingCountry id="F3690E9AFF03FF25FB92FE6E35AAFAE8" box="[1050,1238,1280,1304]" name="French Guiana" pageId="200" pageNumber="199">French Guiana</collectingCountry>
, with all records of this species in
<collectingCountry id="F3690E9AFF03FF25FE1BFE4E3330FAC8" box="[403,588,1312,1336]" name="French Guiana" pageId="200" pageNumber="199">French Guiana</collectingCountry>
being based on misidentifications of
<taxonomicName id="4C7E3589FF03FF25FB99FE4E35D7FAC8" authorityName="Chevrolat" authorityYear="1843" box="[1041,1195,1312,1336]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="endymion">
<emphasis id="B90A9218FF03FF25FB99FE4E35D7FAC8" box="[1041,1195,1312,1336]" italics="true" pageId="200" pageNumber="199">E. endymion</emphasis>
</taxonomicName>
. We doubt if
<taxonomicName id="4C7E3589FF03FF25FAC7FE4E34DCFAC7" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[1359,1440,1312,1335]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<emphasis id="B90A9218FF03FF25FAC7FE4E34DCFAC7" box="[1359,1440,1312,1335]" italics="true" pageId="200" pageNumber="199">E. pan</emphasis>
</taxonomicName>
is present in the Guianas, especially since we are also unaware of any records from
<collectingCountry id="F3690E9AFF03FF25FB26FE2E344DFAA8" box="[1198,1329,1344,1368]" name="Venezuela" pageId="200" pageNumber="199">Venezuela</collectingCountry>
. It would be interesting to see if the old record from
<collectingCountry id="F3690E9AFF03FF25FD36FE0E3263FA88" box="[702,799,1376,1400]" name="Guyana" pageId="200" pageNumber="199">Guyana</collectingCountry>
in
<bibRefCitation id="EFEF33FBFF03FF25FCCCFE0E3541FA88" author="Erichson, W. F." box="[836,1085,1376,1400]" editor="R. Schomburgk" journalOrPublisher="J. J. Weber; Leipzig, Germany" pageId="200" pageNumber="199" pagination="553 - 1260" refId="ref201522" refString="Erichson, W. F. 1848. Insecten. p. 553 - 617. In: R. Schomburgk. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil. J. J. Weber; Leipzig, Germany. p. 553 - 1260." title="Insecten" type="book chapter" volumeTitle="Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil" year="1848">Erichson (1848: 561)</bibRefCitation>
and the specimens identified as
<taxonomicName id="4C7E3589FF03FF25FF6BFEEE3048FA67" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[227,308,1408,1431]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<emphasis id="B90A9218FF03FF25FF6BFEEE3048FA67" box="[227,308,1408,1431]" italics="true" pageId="200" pageNumber="199">E. pan</emphasis>
</taxonomicName>
found in
<collectingCountry id="F3690E9AFF03FF25FE21FEEE3088FA68" box="[425,500,1408,1432]" name="Brazil" pageId="200" pageNumber="199">Brazil</collectingCountry>
north of the Amazon River actually belong to this species, or if they are also misidentified
<taxonomicName id="4C7E3589FF03FF25FEE2FECE3378FA48" authorityName="Chevrolat" authorityYear="1843" box="[362,516,1440,1464]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="endymion">
<emphasis id="B90A9218FF03FF25FEE2FECE3378FA48" box="[362,516,1440,1464]" italics="true" pageId="200" pageNumber="199">E. endymion</emphasis>
</taxonomicName>
. Because of the current identification of the specimens from the Brazilian part of the research area, we regard
<taxonomicName id="4C7E3589FF03FF25FDF7FEAE33ACFA27" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[639,720,1472,1495]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="199" phylum="Arthropoda" rank="species" species="pan">
<emphasis id="B90A9218FF03FF25FDF7FEAE33ACFA27" box="[639,720,1472,1495]" italics="true" pageId="200" pageNumber="199">E. pan</emphasis>
</taxonomicName>
as potentially occurring in the Guianas.
</paragraph>
</subSubSection>
</treatment>
</document>
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