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<document id="1F4867F2633C6D15ADF9F0F2DF0B7F06" ID-CLB-Dataset="56043" ID-DOI="10.5852/ejt.2022.813.1729" ID-GBIF-Dataset="ac406bb4-11b2-425e-b793-599560c503d9" ID-ISSN="2118-9773" ID-Zenodo-Dep="6448751" ID-ZooBank="687DE2C9-28A4-43E6-A47D-DE2D2839AB60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649696887367" checkinUser="felipe" docAuthor="Cole, Mary L. &amp; Herbert, David G." docDate="2022" docId="03A387ECFB7C7A4ACDFD31B00318FB02" docLanguage="en" docName="ejt-813_1-32.pdf" docOrigin="European Journal of Taxonomy 813" docSource="http://dx.doi.org/10.5852/ejt.2022.813.1729" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gulella maraisi Cole &amp; Herbert 2022, sp. nov." docType="treatment" docUuid="1D02CEC4-33FC-48DD-86EF-592869390159" docUuidSource="ZooBank" docVersion="9" lastPageNumber="28" masterDocId="FF9AFF94FB647A51CF7633470125FFEF" masterDocTitle="Eight new species of Gulella Pfeiffer, 1856 from the south-east coast of South Africa (Gastropoda: Streptaxidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="25" updateTime="1699319808271" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9BBC1C9442B4DF8520FDAB99D3D5DD1B">Eight new species of Gulella Pfeiffer, 1856 from the south-east coast of South Africa (Gastropoda: Streptaxidae)</mods:title>
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<mods:namePart id="62B9DBF7B1376741B7403A9E0EB906F5">Cole, Mary L.</mods:namePart>
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<mods:affiliation id="F8918A92A9815078AE1D5C1998ABCB00">East London Museum, 319 Oxford St, East London, 5201, South Africa and Department of Zoology and Entomology, Rhodes University, Makhanda, 6140, South Africa.</mods:affiliation>
<mods:nameIdentifier id="F6A7A7641CC623F20802E4C4631961DE" type="email">marybursey@elmuseum.za.org</mods:nameIdentifier>
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<mods:namePart id="F31A3583D7069FA2A5572416F470AD42">Herbert, David G.</mods:namePart>
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<mods:affiliation id="3946ADCD2A2679E75D3DE342EF92C40A">Department of Natural Sciences, National Museum Wales, Cardiff, United Kingdom and University of KwaZulu-Natal, P. Bag. X 01, Scottsville, 3209, South Africa.</mods:affiliation>
<mods:nameIdentifier id="ABD4415F97BB1ED526BFEEAE51894CDB" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="2A6E688A6733A08D5DDDBDEDACC5E4B3">European Journal of Taxonomy</mods:title>
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<mods:date id="16B28811A64CEAE0FE7445AE5331B73D">2022</mods:date>
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<mods:number id="AEA170E1510297198E3AD283CFC19224">2022-04-11</mods:number>
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<treatment id="03A387ECFB7C7A4ACDFD31B00318FB02" ID-DOI="http://doi.org/10.5281/zenodo.6453005" ID-GBIF-Taxon="194587699" ID-Zenodo-Dep="6453005" LSID="urn:lsid:zoobank.org:act:1D02CEC4-33FC-48DD-86EF-592869390159" httpUri="http://treatment.plazi.org/id/03A387ECFB7C7A4ACDFD31B00318FB02" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
<subSubSection id="C3106571FB7C7A49CDFD31B0028CFCFD" box="[651,937,759,786]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BB536FAFB7C7A49CDFD31B0028CFCFD" blockId="24.[371,1217,759,856]" box="[651,937,759,786]" pageId="24" pageNumber="25">
<heading id="D0FD8196FB7C7A49CDFD31B0028CFCFD" box="[651,937,759,786]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="4C0A4D79FB7C7A49CDFD31B00262FCFE" ID-CoL="8ZFY4" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[651,839,759,786]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7C7A49CDFD31B00262FCFE" bold="true" box="[651,839,759,786]" italics="true" pageId="24" pageNumber="25">Gulella maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7C7A49CC3831BF028CFCFD" box="[846,937,760,786]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB7C7A49CE05305D025DFCB7" pageId="24" pageNumber="25" type="description">
<paragraph id="8BB536FAFB7C7A49CE05305D05E4FCDA" blockId="24.[371,1217,759,856]" box="[371,1217,794,821]" pageId="24" pageNumber="25">
<uri id="FF9B3AF8FB7C7A49CE05305D05E4FCDA" box="[371,1217,794,821]" pageId="24" pageNumber="25">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BB536FAFB7C7A49CDCA307A025DFCB7" blockId="24.[371,1217,759,856]" box="[700,888,829,856]" pageId="24" pageNumber="25">
<figureCitation id="13312A7FFB7C7A49CDCA307A0233FCB7" box="[700,790,829,856]" captionStart="Fig" captionStartId="24.[188,231,1963,1989]" captionTargetBox="[293,1319,1104,1920]" captionTargetId="figure-262@24.[293,1319,1104,1920]" captionTargetPageId="24" captionText="Fig. 10. Distribution map of Gulella abbotti sp. nov. (purple triangles) and G. maraisi sp. nov. (orange circles). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.6448779" httpUri="https://zenodo.org/record/6448779/files/figure.png" pageId="24" pageNumber="25">Figs 10</figureCitation>
,
<figureCitation id="13312A7FFB7C7A49CC543079025DFCB7" box="[802,888,830,856]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="24" pageNumber="25">11A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB7C7A48CFCB303802C5FEA9" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BB536FAFB7C7A49CFCB30380013FC76" blockId="24.[189,310,895,921]" box="[189,310,895,921]" pageId="24" pageNumber="25">
<heading id="D0FD8196FB7C7A49CFCB30380013FC76" bold="true" box="[189,310,895,921]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B97EEAE8FB7C7A49CFCB30380013FC76" bold="true" box="[189,310,895,921]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB7C7A49CFCB30EA0452FC04" blockId="24.[189,1399,941,1003]" pageId="24" pageNumber="25">Shell small, cylindrical; sculptured by axial ribs that extend to mid-whorl on spire whorls and from suture to suture on last whorl; aperture little obstructed by teeth; dentition typically three-fold, including</paragraph>
<caption id="DF756672FB7C7A49CFCA34EC0322F806" ID-DOI="http://doi.org/10.5281/zenodo.6448779" ID-Zenodo-Dep="6448779" httpUri="https://zenodo.org/record/6448779/files/figure.png" pageId="24" pageNumber="25" startId="24.[188,231,1963,1989]" targetBox="[293,1319,1104,1920]" targetPageId="24">
<paragraph id="8BB536FAFB7C7A49CFCA34EC0322F806" blockId="24.[188,1399,1963,2025]" pageId="24" pageNumber="25">
<emphasis id="B97EEAE8FB7C7A49CFCA34EC003CF82A" bold="true" box="[188,281,1963,1989]" pageId="24" pageNumber="25">Fig. 10.</emphasis>
Distribution map of
<taxonomicName id="4C0A4D79FB7C7A49CD6434EC03E0F82A" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[530,709,1963,1989]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="abbotti" status="sp. nov.">
<emphasis id="B97EEAE8FB7C7A49CD6434EC03E0F82A" box="[530,709,1963,1989]" italics="true" pageId="24" pageNumber="25">Gulella abbotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7C7A49CDBB34EB020DF829" box="[717,808,1964,1990]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
(purple triangles) and
<taxonomicName id="4C0A4D79FB7C7A49CB4334EB0596F82A" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[1077,1203,1964,1990]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7C7A49CB4334EB0596F82A" box="[1077,1203,1964,1990]" italics="true" pageId="24" pageNumber="25">G. maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7C7A49CBCC34EB0430F829" box="[1210,1301,1964,1990]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
(orange circles). Contour at 1000 m.
</paragraph>
</caption>
<paragraph id="8BB536FAFB7D7A48CFCB324E02C5FEA9" blockId="25.[189,1398,265,327]" pageId="25" pageNumber="26">a parietal lamella, a simple labral tooth and a quadrate columella lamella; sometimes with an additional small basal tooth just left of centre; umbilicus small, elongate-ovate.</paragraph>
</subSubSection>
<subSubSection id="C3106571FB7D7A48CFCB322902B2FE36" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BB536FAFB7D7A48CFCB32290061FE67" blockId="25.[189,324,366,392]" box="[189,324,366,392]" pageId="25" pageNumber="26">
<heading id="D0FD8196FB7D7A48CFCB32290061FE67" bold="true" box="[189,324,366,392]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B97EEAE8FB7D7A48CFCB32290061FE67" bold="true" box="[189,324,366,392]" pageId="25" pageNumber="26">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB7D7A48CFCB32DB02B2FE36" blockId="25.[189,1399,412,474]" pageId="25" pageNumber="26">
Named for Dr Johan P. Marais, a very productive conchologist who collected much material for the
<collectingRegion id="49CEF818FB7D7A48CFCB32F80050FE36" box="[189,373,447,473]" country="South Africa" name="KwaZulu-Natal" pageId="25" pageNumber="26">KwaZulu-Natal</collectingRegion>
Museum, including specimens of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB7D7A48CFCB314702F4FACC" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BB536FAFB7D7A48CFCB3147008FFDF5" blockId="25.[189,426,512,538]" box="[189,426,512,538]" pageId="25" pageNumber="26">
<heading id="D0FD8196FB7D7A48CFCB3147008FFDF5" bold="true" box="[189,426,512,538]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B97EEAE8FB7D7A48CFCB3147008FFDF5" bold="true" box="[189,426,512,538]" pageId="25" pageNumber="26">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB7D7A48CF9C316803EAFD7F" blockId="25.[185,1399,559,656]" pageId="25" pageNumber="26">
<materialsCitation id="3B623CA7FB7D7A48CF9C316803EAFD7F" ID-GBIF-Occurrence="3750850347" collectingDate="1996-06" collectionCode="NMSA" collectorName="J. P. Marais" country="South Africa" latitude="-31.083" location="KwaZulu-Natal" longLatPrecision="72" longitude="30.197" pageId="25" pageNumber="26" specimenCode="P1676/T4520" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<emphasis id="B97EEAE8FB7D7A48CF9C31680079FDA6" bold="true" box="[234,348,559,585]" pageId="25" pageNumber="26">
<typeStatus id="54B18858FB7D7A48CF9C31680079FDA6" box="[234,348,559,585]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F31D766AFB7D7A48CFCB311400B5FD82" box="[189,400,595,621]" name="South Africa" pageId="25" pageNumber="26">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B97EEAE8FB7D7A48CED9311403A2FD82" bold="true" box="[431,647,595,621]" pageId="25" pageNumber="26">
<collectingRegion id="49CEF818FB7D7A48CED931140351FD82" box="[431,628,595,621]" country="South Africa" name="KwaZulu-Natal" pageId="25" pageNumber="26">KwaZulu-Natal</collectingRegion>
•
</emphasis>
<locationDeviation id="9A072FD8FB7D7A48CDE63114051DFD83" box="[656,1080,594,621]" pageId="25" pageNumber="26">Mtamvuna River mouth beach drift</locationDeviation>
;
<geoCoordinate id="EE3E503DFB7D7A48CB3331140590FD82" box="[1093,1205,595,621]" degrees="31.083" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="55" value="-31.083">31.083°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB7D7A48CBB23114041CFD82" box="[1220,1337,595,621]" degrees="30.197" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="55" value="30.197">30.197° E</geoCoordinate>
;
<date id="FFB4103AFB7D7A48CA30311401D2FD7F" pageId="25" pageNumber="26" value="1996-06">
<collectingDate id="EFF0E9D2FB7D7A48CA30311401D2FD7F" pageId="25" pageNumber="26" value="1996-06">Jun. 1996</collectingDate>
</date>
; leg.
<collectorName id="26FF532CFB7D7A48CE40313100E4FD7F" box="[310,449,630,656]" pageId="25" pageNumber="26">J. P. Marais</collectorName>
;
<collectionCode id="ED1BAE3FFB7D7A48CEBB31310305FD7F" box="[461,544,630,656]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB7D7A48CD51313203EEFD7F" box="[551,715,629,656]" pageId="25" pageNumber="26">P1676/T4520</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB536FAFB7D7A48CF9C31F00323FBE1" blockId="25.[185,1399,695,1038]" pageId="25" pageNumber="26">
<materialsCitation id="3B623CA7FB7D7A48CF9C31F00239FCF7" ID-GBIF-Occurrence="3750850330" collectingDate="1996-06" collectionCode="NMSA" collectorName="J. P. Marais" country="South Africa" latitude="-31.083" location="KwaZulu-Natal" longLatPrecision="72" longitude="30.197" pageId="25" pageNumber="26" specimenCode="V3993/T4485" specimenCount="3" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<emphasis id="B97EEAE8FB7D7A48CF9C31F00042FD3E" bold="true" box="[234,359,695,721]" pageId="25" pageNumber="26">
<typeStatus id="54B18858FB7D7A48CF9C31F00042FD3E" box="[234,359,695,721]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
(listed south to north)
<collectingCountry id="F31D766AFB7D7A48CFCB319D00AEFD1B" box="[189,395,730,756]" name="South Africa" pageId="25" pageNumber="26">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B97EEAE8FB7D7A48CEE8319D0346FD1B" bold="true" box="[414,611,730,756]" pageId="25" pageNumber="26">
<collectingRegion id="49CEF818FB7D7A48CEE8319D0346FD1B" box="[414,611,730,756]" country="South Africa" name="KwaZulu-Natal" pageId="25" pageNumber="26">KwaZulu-Natal</collectingRegion>
</emphasis>
•
<specimenCount id="9D0CFD73FB7D7A48CD0D319D03F6FD1A" box="[635,723,730,757]" pageId="25" pageNumber="26">3specs</specimenCount>
.;
<locationDeviation id="9A072FD8FB7D7A48CD97319C0552FD1B" box="[737,1143,730,757]" pageId="25" pageNumber="26">Mtamvuna River mouth beach drift</locationDeviation>
;
<geoCoordinate id="EE3E503DFB7D7A48CBF6319D05D7FD1B" box="[1152,1266,730,756]" degrees="31.083" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="55" value="-31.083">31.083°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB7D7A48CB8A319D0454FD1A" box="[1276,1393,730,757]" degrees="30.197" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="55" value="30.197">30.197°E</geoCoordinate>
;
<date id="FFB4103AFB7D7A48CFCF31B90017FCF8" box="[185,306,765,792]" pageId="25" pageNumber="26" value="1996-06">
<collectingDate id="EFF0E9D2FB7D7A48CFCF31B90017FCF8" box="[185,306,765,792]" pageId="25" pageNumber="26" value="1996-06">Jun. 1996</collectingDate>
</date>
;
<collectorName id="26FF532CFB7D7A48CE3431B900F5FCF8" box="[322,464,765,792]" pageId="25" pageNumber="26">J. P. Marais</collectorName>
leg.;
<collectionCode id="ED1BAE3FFB7D7A48CD6C31B90348FCF7" box="[538,621,766,792]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB7D7A48CD0E31BA0239FCF7" box="[632,796,765,792]" pageId="25" pageNumber="26">V3993/T4485</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB7D7A48CC4A31B90382FCD4" ID-GBIF-Occurrence="3750850416" collectingDate="1996-06" collectionCode="NMSA" collectorName="J. P. Marais" country="South Africa" latitude="-31.083" location="Mtamvuna River" longLatPrecision="72" longitude="30.197" pageId="25" pageNumber="26" specimenCode="NHMUK 20210077, V3993" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="9D0CFD73FB7D7A48CC4A31B902B0FCF7" box="[828,917,766,792]" pageId="25" pageNumber="26" type="generic">1 spec.</specimenCount>
; same collection data as for preceding;
<specimenCode id="DBAC9E81FB7D7A48CFCB30660089FCD4" box="[189,428,801,827]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="25" pageNumber="26" type="Museum">NHMUK 20210077</specimenCode>
, prev.
<collectionCode id="ED1BAE3FFB7D7A48CE8A3066036AFCD4" box="[508,591,801,827]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB7D7A48CD2130660382FCD4" box="[599,679,801,827]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">V3993</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB7D7A48CDB430660369FC6E" ID-GBIF-Occurrence="3750850395" collectingDate="2008-05-06" collectionCode="NMSA" collectorName="D. Herbert &amp; L. Davis" country="South Africa" county="Port Shepstone" elevation="130" latitude="-30.65898" location="Hlokohloko Valley" longLatPrecision="1" longitude="30.34292" municipality="Simuma area" pageId="25" pageNumber="26" specimenCode="W6277/T4486" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="9D0CFD73FB7D7A48CDB43066023DFCD4" box="[706,792,801,827]" pageId="25" pageNumber="26" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="62D44E76FB7D7A48CC53306602FFFCD4" box="[805,986,801,827]" pageId="25" pageNumber="26">Port Shepstone</collectingCounty>
,
<collectingMunicipality id="6BD1AC80FB7D7A48CC91306605A0FCD4" box="[999,1157,801,827]" pageId="25" pageNumber="26">Simuma area</collectingMunicipality>
,
<location id="8ED56021FB7D7A48CBE430660457FCD4" LSID="urn:lsid:plazi:treatment:03A387ECFB7C7A4ACDFD31B00318FB02:8ED56021FB7D7A48CBE430660457FCD4" box="[1170,1394,801,827]" country="South Africa" county="Port Shepstone" latitude="-30.65898" longLatPrecision="1" longitude="30.34292" municipality="Simuma area" name="Hlokohloko Valley" pageId="25" pageNumber="26" stateProvince="KwaZulu-Natal">Hlokohloko Valley</location>
,
<location id="8ED56021FB7D7A48CFC8300300C4FCB1" LSID="urn:lsid:plazi:treatment:03A387ECFB7C7A4ACDFD31B00318FB02:8ED56021FB7D7A48CFC8300300C4FCB1" box="[190,481,836,862]" country="South Africa" county="Port Shepstone" latitude="-30.65898" longLatPrecision="1" longitude="30.34292" municipality="Simuma area" name="degraded riverine forest" pageId="25" pageNumber="26" stateProvince="KwaZulu-Natal">degraded riverine forest</location>
;
<geoCoordinate id="EE3E503DFB7D7A48CE863003035BFCB1" box="[496,638,836,863]" degrees="30.65898" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-30.65898">30.65898°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB7D7A48CDF830030205FCB1" box="[654,800,836,862]" degrees="30.34292" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="30.34292">30.34292° E</geoCoordinate>
, ca
<elevation id="0027D1C9FB7D7A48CC22300302C0FCB1" box="[852,997,836,862]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="26" unit="m" value="130.0">
<quantity id="4CF29B1FFB7D7A48CC2230030287FCB1" box="[852,930,836,862]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="26" unit="m" value="130.0">130 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB7D7A48CC80300305A8FCB1" box="[1014,1165,836,862]" pageId="25" pageNumber="26" value="2008-05-06">
<collectingDate id="EFF0E9D2FB7D7A48CC80300305A8FCB1" box="[1014,1165,836,862]" pageId="25" pageNumber="26" value="2008-05-06">6 May 2008</collectingDate>
</date>
;
<collectorName id="26FF532CFB7D7A48CBEB3003043BFCB1" box="[1181,1310,836,862]" pageId="25" pageNumber="26">D. Herbert</collectorName>
and
<collectorName id="26FF532CFB7D7A48CA2B30030024FC6E" pageId="25" pageNumber="26">L. Davis</collectorName>
leg.;
<collectionCode id="ED1BAE3FFB7D7A48CE30302000BCFC6E" box="[326,409,871,897]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB7D7A48CED730200369FC6E" box="[417,588,871,897]" pageId="25" pageNumber="26">W6277/T4486</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB7D7A48CD1E3020036EFC27" ID-GBIF-Occurrence="3750850313" collectingDate="2011-03-05" collectionCode="NMSA" collectorName="D. Herbert &amp; L. Davis" country="South Africa" county="Port Shepstone" elevation="175" latitude="-30.6631" location="Hlokohloko Valley" longLatPrecision="1" longitude="30.33685" municipality="Simuma area" pageId="25" pageNumber="26" specimenCode="W7831/T4487" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="9D0CFD73FB7D7A48CD1E3020039BFC6D" box="[616,702,871,898]" pageId="25" pageNumber="26" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="62D44E76FB7D7A48CDBD302F02A4FC6E" box="[715,897,871,898]" pageId="25" pageNumber="26">Port Shepstone</collectingCounty>
,
<collectingMunicipality id="6BD1AC80FB7D7A48CCFB30200509FC6D" box="[909,1068,871,898]" pageId="25" pageNumber="26">Simuma area</collectingMunicipality>
,
<location id="8ED56021FB7D7A48CB4E3020043CFC6E" LSID="urn:lsid:plazi:treatment:03A387ECFB7C7A4ACDFD31B00318FB02:8ED56021FB7D7A48CB4E3020043CFC6E" box="[1080,1305,871,897]" country="South Africa" county="Port Shepstone" latitude="-30.6631" longLatPrecision="1" longitude="30.33685" municipality="Simuma area" name="Hlokohloko Valley" pageId="25" pageNumber="26" stateProvince="KwaZulu-Natal">Hlokohloko Valley</location>
, stn 11- 001,
<location id="8ED56021FB7D7A48CF8D30CD008DFC4B" LSID="urn:lsid:plazi:treatment:03A387ECFB7C7A4ACDFD31B00318FB02:8ED56021FB7D7A48CF8D30CD008DFC4B" box="[251,424,906,932]" country="South Africa" county="Port Shepstone" latitude="-30.6631" longLatPrecision="1" longitude="30.33685" municipality="Simuma area" name="riverine forest" pageId="25" pageNumber="26" stateProvince="KwaZulu-Natal">riverine forest</location>
;
<geoCoordinate id="EE3E503DFB7D7A48CEC030CD0360FC4B" box="[438,581,906,932]" degrees="30.66310" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-30.6631">30.66310°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB7D7A48CD2330CD03C2FC4A" box="[597,743,906,933]" degrees="30.33685" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="30.33685">30.33685° E</geoCoordinate>
, ca
<elevation id="0027D1C9FB7D7A48CC6D30CD02C3FC4B" box="[795,998,906,933]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="25" pageNumber="26" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">
<quantity id="4CF29B1FFB7D7A48CC6D30CD0286FC4A" box="[795,931,906,933]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="25" pageNumber="26" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">150–200 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB7D7A48CC8130CD05ABFC4B" box="[1015,1166,906,933]" pageId="25" pageNumber="26" value="2011-03-05">
<collectingDate id="EFF0E9D2FB7D7A48CC8130CD05ABFC4B" box="[1015,1166,906,933]" pageId="25" pageNumber="26" value="2011-03-05">5 Mar. 2011</collectingDate>
</date>
;
<collectorName id="26FF532CFB7D7A48CBEB30CC043BFC4B" box="[1181,1310,906,933]" pageId="25" pageNumber="26">D. Herbert</collectorName>
and
<collectorName id="26FF532CFB7D7A48CA2B30CC0024FC28" pageId="25" pageNumber="26">L. Davis</collectorName>
leg.;
<collectionCode id="ED1BAE3FFB7D7A48CE3330E900BDFC27" box="[325,408,942,968]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB7D7A48CED630E9036EFC27" box="[416,587,942,968]" pageId="25" pageNumber="26">W7831/T4487</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB7D7A48CD1030E90327FBE1" ID-GBIF-Occurrence="3750850415" collectedFrom="in leaf-litter" collectingDate="2012-02" collectionCode="NMSA" collectorName="J. Harvey" country="South Africa" county="Port Shepstone area" elevation="237" latitude="-30.6683" location="woodland / forest" longLatPrecision="6" longitude="30.347" municipality="Marble Delta" pageId="25" pageNumber="26" specimenCode="W8786/T4483" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="9D0CFD73FB7D7A48CD1030E90399FC27" box="[614,700,942,968]" pageId="25" pageNumber="26" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="62D44E76FB7D7A48CDBF30E90292FC27" box="[713,951,941,968]" pageId="25" pageNumber="26">Port Shepstone area</collectingCounty>
,
<collectingMunicipality id="6BD1AC80FB7D7A48CCB230EA0546FC28" box="[964,1123,941,967]" pageId="25" pageNumber="26">Marble Delta</collectingMunicipality>
,
<locationDeviation id="9A072FD8FB7D7A48CB0630EA01CBFC04" pageId="25" pageNumber="26">South side of Simuma Hill</locationDeviation>
,
<location id="8ED56021FB7D7A48CF8A30960098FC04" LSID="urn:lsid:plazi:treatment:03A387ECFB7C7A4ACDFD31B00318FB02:8ED56021FB7D7A48CF8A30960098FC04" box="[252,445,977,1003]" country="South Africa" county="Port Shepstone area" latitude="-30.6683" longLatPrecision="6" longitude="30.347" municipality="Marble Delta" name="woodland / forest" pageId="25" pageNumber="26" stateProvince="KwaZulu-Natal">woodland/forest</location>
,
<collectedFrom id="38167507FB7D7A48CEBD3096037CFC04" box="[459,601,977,1003]" pageId="25" pageNumber="26">in leaf-litter</collectedFrom>
, stn S1;
<geoCoordinate id="EE3E503DFB7D7A48CDB73096021BFC04" box="[705,830,977,1003]" degrees="30.6683" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="5" value="-30.6683">30.6683°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB7D7A48CC3B309602F6FC04" box="[845,979,977,1003]" degrees="30.3470" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="5" value="30.347">30.3470° E</geoCoordinate>
;
<elevation id="0027D1C9FB7D7A48CC963096054BFC04" box="[992,1134,977,1003]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="25" pageNumber="26" unit="m" value="237.0">
<quantity id="4CF29B1FFB7D7A48CC9630960509FC04" box="[992,1068,977,1003]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="25" pageNumber="26" unit="m" value="237.0">237 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB7D7A48CB0B309605D2FC04" box="[1149,1271,977,1003]" pageId="25" pageNumber="26" value="2012-02">
<collectingDate id="EFF0E9D2FB7D7A48CB0B309605D2FC04" box="[1149,1271,977,1003]" pageId="25" pageNumber="26" value="2012-02">Feb. 2012</collectingDate>
</date>
;
<collectorName id="26FF532CFB7D7A48CA7330960453FC04" box="[1285,1398,977,1003]" pageId="25" pageNumber="26">J. Harvey</collectorName>
leg.;
<collectionCode id="ED1BAE3FFB7D7A48CF8F30B30069FBE1" box="[249,332,1012,1038]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB7D7A48CE2530B30327FBE1" box="[339,514,1012,1038]" pageId="25" pageNumber="26">W8786/T4483</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB536FAFB7D7A48CF9C37720083FBA0" blockId="25.[186,1399,1077,1315]" box="[234,422,1077,1103]" pageId="25" pageNumber="26">
<emphasis id="B97EEAE8FB7D7A48CF9C37720083FBA0" bold="true" box="[234,422,1077,1103]" pageId="25" pageNumber="26">Other material</emphasis>
</paragraph>
<paragraph id="8BB536FAFB7D7A48CFCB371E02F4FACC" blockId="25.[186,1399,1077,1315]" pageId="25" pageNumber="26">
<materialsCitation id="3B623CA7FB7D7A48CFCB371E02A3FB79" ID-GBIF-Occurrence="3750850363" collectingDate="1996-06" collectionCode="NMSA" collectorName="J. P. Marais" country="South Africa" latitude="-31.083" location="KwaZulu-Natal" longLatPrecision="72" longitude="30.197" pageId="25" pageNumber="26" specimenCode="P1613, V3993" specimenCount="2" stateProvince="KwaZulu-Natal">
<collectingCountry id="F31D766AFB7D7A48CFCB371E00AEFB9C" box="[189,395,1113,1139]" name="South Africa" pageId="25" pageNumber="26">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B97EEAE8FB7D7A48CEE8371E0346FB9C" bold="true" box="[414,611,1113,1139]" pageId="25" pageNumber="26">
<collectingRegion id="49CEF818FB7D7A48CEE8371E0346FB9C" box="[414,611,1113,1139]" country="South Africa" name="KwaZulu-Natal" pageId="25" pageNumber="26">KwaZulu-Natal</collectingRegion>
</emphasis>
•
<specimenCount id="9D0CFD73FB7D7A48CD0D371E03F6FB9C" box="[635,723,1113,1139]" pageId="25" pageNumber="26">2specs</specimenCount>
.;
<locationDeviation id="9A072FD8FB7D7A48CD97371E0552FB9D" box="[737,1143,1112,1139]" pageId="25" pageNumber="26">Mtamvuna River mouth beach drift</locationDeviation>
;
<geoCoordinate id="EE3E503DFB7D7A48CBF6371E05D7FB9C" box="[1152,1266,1113,1139]" degrees="31.083" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="55" value="-31.083">31.083°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB7D7A48CB8A371E0454FB9C" box="[1276,1393,1113,1139]" degrees="30.197" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="55" value="30.197">30.197°E</geoCoordinate>
;
<date id="FFB4103AFB7D7A48CFCC373B0008FB79" box="[186,301,1148,1174]" pageId="25" pageNumber="26" value="1996-06">
<collectingDate id="EFF0E9D2FB7D7A48CFCC373B0008FB79" box="[186,301,1148,1174]" pageId="25" pageNumber="26" value="1996-06">Jun. 1996</collectingDate>
</date>
;
<collectorName id="26FF532CFB7D7A48CE40373B009DFB79" box="[310,440,1148,1174]" pageId="25" pageNumber="26">J. P. Marais</collectorName>
leg.;
<collectionCode id="ED1BAE3FFB7D7A48CE83373B036DFB79" box="[501,584,1148,1174]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB7D7A48CD3B373B03BDFB79" box="[589,664,1148,1174]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">P1613</specimenCode>
prev.
<collectionCode id="ED1BAE3FFB7D7A48CDAD373B020BFB79" box="[731,814,1148,1174]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB7D7A48CC45373B02A3FB79" box="[819,902,1148,1174]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="26" type="Museum">V3993</specimenCode>
</materialsCitation>
.
<materialsCitation id="3B623CA7FB7D7A48CCF9373B0212FB32" ID-GBIF-Occurrence="3750850350" collectingDate="2011-02-15" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" county="Mkambati Nature Reserve" latitude="-31.2738" location="Mkambati Nature Reserve" longLatPrecision="6" longitude="30.0236" municipality="East" pageId="25" pageNumber="26" specimenCode="ELMD 18754" specimenCount="1" stateProvince="Eastern Cape">
<emphasis id="B97EEAE8FB7D7A48CCF9373B0512FB7A" bold="true" box="[911,1079,1147,1174]" pageId="25" pageNumber="26">
<collectingRegion id="49CEF818FB7D7A48CCF9373B0512FB7A" box="[911,1079,1147,1174]" country="South Africa" name="Eastern Cape" pageId="25" pageNumber="26">Eastern Cape</collectingRegion>
</emphasis>
•
<specimenCount id="9D0CFD73FB7D7A48CB3D373B05B8FB79" box="[1099,1181,1148,1174]" pageId="25" pageNumber="26" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="62D44E76FB7D7A48CBD1373B003EFB55" pageId="25" pageNumber="26">Mkambati Nature Reserve</collectingCounty>
,
<locationDeviation id="9A072FD8FB7D7A48CE5A37E703BFFB56" box="[300,666,1183,1210]" pageId="25" pageNumber="26">East bank of Mkambati River</locationDeviation>
,
<locationDeviation id="9A072FD8FB7D7A48CDDC37D805F8FB56" box="[682,1245,1183,1209]" pageId="25" pageNumber="26">between Mkambati and Strandloper waterfalls</locationDeviation>
;
<geoCoordinate id="EE3E503DFB7D7A48CB9837D80454FB56" box="[1262,1393,1183,1209]" degrees="31.2738" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="5" value="-31.2738">31.2738° S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB7D7A48CFCB37850064FB32" box="[189,321,1218,1245]" degrees="30.0236" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="5" value="30.0236">30.0236° E</geoCoordinate>
;
<date id="FFB4103AFB7D7A48CE3B378500CDFB33" box="[333,488,1218,1245]" pageId="25" pageNumber="26" value="2011-02-15">
<collectingDate id="EFF0E9D2FB7D7A48CE3B378500CDFB33" box="[333,488,1218,1245]" pageId="25" pageNumber="26" value="2011-02-15">15 Feb. 2011</collectingDate>
</date>
;
<collectorName id="26FF532CFB7D7A48CE8537840376FB33" box="[499,595,1218,1245]" pageId="25" pageNumber="26">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB7D7A48CDE237840212FB32" box="[660,823,1218,1245]" pageId="25" pageNumber="26">ELMD 18754</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB7D7A48CC393785036DFAEF" ID-GBIF-Occurrence="3750850404" collectingDate="2019-03-17" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" county="Mkambati Nature Reserve" latitude="-31.2738" location="Mkambati River" longLatPrecision="6" longitude="30.0236" municipality="East" pageId="25" pageNumber="26" specimenCode="ELMD 18475" specimenCount="1" stateProvince="Eastern Cape">
<specimenCount id="9D0CFD73FB7D7A48CC3937850286FB32" box="[847,931,1218,1245]" pageId="25" pageNumber="26" type="generic">1 spec.</specimenCount>
; same locality as for preceding;
<date id="FFB4103AFB7D7A48CA6A378501DFFB10" pageId="25" pageNumber="26" value="2019-03-17">
<collectingDate id="EFF0E9D2FB7D7A48CA6A378501DFFB10" pageId="25" pageNumber="26" value="2019-03-17">17 Mar. 2019</collectingDate>
</date>
;
<collectorName id="26FF532CFB7D7A48CE7037A10040FB10" box="[262,357,1253,1280]" pageId="25" pageNumber="26">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB7D7A48CED037A1036DFAEF" box="[422,584,1253,1280]" pageId="25" pageNumber="26">ELMD 18475</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB7D7A48CD1637A202E8FACC" ID-GBIF-Occurrence="3750850407" collectingDate="2011-02-16" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" county="Mkambati Nature Reserve" latitude="-31.3" location="gorge of Daza River" longLatPrecision="6" longitude="29.9879" municipality="Daza River" pageId="25" pageNumber="26" specimenCode="ELMD 18756" specimenCount="4" stateProvince="Eastern Cape">
<specimenCount id="9D0CFD73FB7D7A48CD1637A2039EFAEF" box="[608,699,1253,1280]" pageId="25" pageNumber="26">4 specs</specimenCount>
.;
<collectingCounty id="62D44E76FB7D7A48CDBD37A20524FAEF" box="[715,1025,1253,1280]" pageId="25" pageNumber="26">Mkambati Nature Reserve</collectingCounty>
,
<location id="8ED56021FB7D7A48CB7B37A105DEFB10" LSID="urn:lsid:plazi:treatment:03A387ECFB7C7A4ACDFD31B00318FB02:8ED56021FB7D7A48CB7B37A105DEFB10" box="[1037,1275,1253,1280]" country="South Africa" county="Mkambati Nature Reserve" latitude="-31.3" longLatPrecision="6" longitude="29.9879" municipality="Daza River" name="gorge of Daza River" pageId="25" pageNumber="26" stateProvince="Eastern Cape">gorge of Daza River</location>
,
<location id="8ED56021FB7D7A48CA7337A10457FB10" LSID="urn:lsid:plazi:treatment:03A387ECFB7C7A4ACDFD31B00318FB02:8ED56021FB7D7A48CA7337A10457FB10" box="[1285,1394,1253,1280]" country="South Africa" county="Mkambati Nature Reserve" latitude="-31.3" longLatPrecision="6" longitude="29.9879" municipality="Daza River" name="East side" pageId="25" pageNumber="26" stateProvince="Eastern Cape">East side</location>
;
<geoCoordinate id="EE3E503DFB7D7A48CFCA364E001BFACC" box="[188,318,1289,1315]" degrees="31.3000" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="5" value="-31.3">31.3000° S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB7D7A48CE3D364E00EBFACC" box="[331,462,1289,1315]" degrees="29.9879" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="5" value="29.9879">29.9879°E</geoCoordinate>
;
<date id="FFB4103AFB7D7A48CEAD364F0352FACC" box="[475,631,1288,1315]" pageId="25" pageNumber="26" value="2011-02-16">
<collectingDate id="EFF0E9D2FB7D7A48CEAD364F0352FACC" box="[475,631,1288,1315]" pageId="25" pageNumber="26" value="2011-02-16">16 Feb. 2011</collectingDate>
</date>
;
<collectorName id="26FF532CFB7D7A48CDF5364E03C6FACD" box="[643,739,1288,1315]" pageId="25" pageNumber="26">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB7D7A48CC53364E02E8FACC" box="[805,973,1288,1315]" pageId="25" pageNumber="26">ELMD 18756</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB7D7A48CFCB360D0478F8F9" pageId="25" pageNumber="26" type="description">
<paragraph id="8BB536FAFB7D7A48CFCB360D006BFA8B" blockId="25.[189,334,1354,1380]" box="[189,334,1354,1380]" pageId="25" pageNumber="26">
<heading id="D0FD8196FB7D7A48CFCB360D006BFA8B" bold="true" box="[189,334,1354,1380]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B97EEAE8FB7D7A48CFCB360D006BFA8B" bold="true" box="[189,334,1354,1380]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB7D7A48CFCB363E0478F8F9" blockId="25.[187,1400,1400,1814]" pageId="25" pageNumber="26">
SHELL (
<figureCitation id="13312A7FFB7D7A48CE65363F004FFA7C" box="[275,362,1400,1427]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
). Shell small, cylindrical, length
<quantity id="4CF29B1FFB7D7A48CD82363F02A5FA7C" box="[756,896,1400,1427]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.2" metricValueMin="3.5" pageId="25" pageNumber="26" unit="mm" value="3.85" valueMax="4.2" valueMin="3.5">3.5–4.2 mm</quantity>
, width
<quantity id="4CF29B1FFB7D7A48CCAE363F0540FA7C" box="[984,1125,1400,1427]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="25" pageNumber="26" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
, L:W 2.3–2.6 (n = 11). Protoconch
<quantity id="4CF29B1FFB7D7A48CE3A36DB0082FA59" box="[332,423,1436,1462]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="25" pageNumber="26" unit="mm" value="1.1">1.1 mm</quantity>
in diameter, comprising approx. 2.5 whorls, smooth; junction between protoconch and teleoconch distinct. Teleoconch comprising approx. 5 whorls; whorls moderately convex; sculptured by subsutural axial riblets extending to mid-whorl on earlier whorls, but strong on last whorl and running from suture to suture (
<figureCitation id="13312A7FFB7D7A48CEB535420374F9CF" box="[451,593,1541,1568]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="25" pageNumber="26">Fig. 11A, B</figureCitation>
); axial pleats prominent on base, running into umbilicus (
<figureCitation id="13312A7FFB7D7A48CB8D35420442F9F0" box="[1275,1383,1541,1567]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="25" pageNumber="26">Fig. 11D</figureCitation>
). Peristome thickened, interrupted in parietal region and with a hiatus behind parietal lamella (
<figureCitation id="13312A7FFB7D7A48CB89356F044CF9AC" box="[1279,1385,1576,1603]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="25" pageNumber="26">Fig. 11C</figureCitation>
). Aperture little obstructed by teeth; apertural dentition three- or four-fold (
<figureCitation id="13312A7FFB7D7A48CB47350B05E5F989" box="[1073,1216,1612,1638]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="25" pageNumber="26">Fig. 11C, E</figureCitation>
): 1) an oblique parietal lamella which begins above insertion of labrum and curves and runs into aperture; 2) a simple in-running, ridge-like mid-labral tooth; 3) a small basal tooth just to left of centre near the lip edge in the
<collectingRegion id="49CEF818FB7D7A48CE7C35F1008DF920" box="[266,424,1717,1744]" country="South Africa" name="Eastern Cape" pageId="25" pageNumber="26">Eastern Cape</collectingRegion>
population (
<figureCitation id="13312A7FFB7D7A48CD3735F2038EF920" box="[577,683,1717,1743]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="25" pageNumber="26">Fig. 11E</figureCitation>
); 4) a quadrate columella lamella. Labral tooth corresponds with a shallow pit behind outer lip (
<figureCitation id="13312A7FFB7D7A48CD1C359F03F3F91C" box="[618,726,1752,1779]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="25" pageNumber="26">Fig. 11B</figureCitation>
). Umbilicus very small, elongate-ovate (
<figureCitation id="13312A7FFB7D7A48CBCF359F0403F91C" box="[1209,1318,1752,1779]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[211,1377,281,1845]" captionTargetId="figure-74@26.[203,1385,265,1848]" captionTargetPageId="26" captionText="Fig. 11. Gulella maraisi sp. nov., holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm. A. Aperture view. B. Side view. C. View into aperture. D. Oblique view of base showing umbilicus. E. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="25" pageNumber="26">Fig. 11D</figureCitation>
). Shell translucent, uniformly milky-white when fresh, reddish-pink dried tissue of animal visible internally.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB7D7A48CFCB347A03F9F846" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BB536FAFB7D7A48CFCB347A00E1F8B8" blockId="25.[189,452,1853,1879]" box="[189,452,1853,1879]" pageId="25" pageNumber="26">
<emphasis id="B97EEAE8FB7D7A48CFCB347A0073F8B8" bold="true" box="[189,342,1853,1879]" pageId="25" pageNumber="26">Distribution</emphasis>
(
<figureCitation id="13312A7FFB7D7A48CE10347A009EF8B8" box="[358,443,1853,1879]" captionStart="Fig" captionStartId="24.[188,231,1963,1989]" captionTargetBox="[293,1319,1104,1920]" captionTargetId="figure-262@24.[293,1319,1104,1920]" captionTargetPageId="24" captionText="Fig. 10. Distribution map of Gulella abbotti sp. nov. (purple triangles) and G. maraisi sp. nov. (orange circles). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.6448779" httpUri="https://zenodo.org/record/6448779/files/figure.png" pageId="25" pageNumber="26">Fig. 10</figureCitation>
)
</paragraph>
<paragraph id="8BB536FAFB7D7A48CFCB342B03F9F846" blockId="25.[189,1398,1900,1961]" pageId="25" pageNumber="26">
Recorded at isolated localities in southern
<collectingRegion id="49CEF818FB7D7A48CDC5342B024EF869" box="[691,875,1900,1926]" country="South Africa" name="KwaZulu-Natal" pageId="25" pageNumber="26">KwaZulu-Natal</collectingRegion>
and northeast
<collectingRegion id="49CEF818FB7D7A48CB6E342B0591F869" box="[1048,1204,1900,1926]" country="South Africa" name="Eastern Cape" pageId="25" pageNumber="26">Eastern Cape</collectingRegion>
provinces, from the coast up to approx.
<quantity id="4CF29B1FFB7D7A48CEB934C8033FF846" box="[463,538,1935,1961]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="26" unit="m" value="130.0">130 m</quantity>
above sea level.
</paragraph>
</subSubSection>
<caption id="DF756672FB7E7A4BCFCB34230013F806" ID-DOI="http://doi.org/10.5281/zenodo.6448781" ID-Zenodo-Dep="6448781" httpUri="https://zenodo.org/record/6448781/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1892,1918]" targetBox="[211,1377,281,1845]" targetPageId="26">
<paragraph id="8BB536FAFB7E7A4BCFCB34230013F806" blockId="26.[189,1399,1892,2025]" pageId="26" pageNumber="27">
<emphasis id="B97EEAE8FB7E7A4BCFCB3423003EF890" bold="true" box="[189,283,1892,1919]" pageId="26" pageNumber="27">Fig. 11.</emphasis>
<taxonomicName id="4C0A4D79FB7E7A4BCE50342200C6F890" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[294,483,1893,1919]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7E7A4BCE50342200C6F890" box="[294,483,1893,1919]" italics="true" pageId="26" pageNumber="27">Gulella maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7E7A4BCE9B3422036BF890" box="[493,590,1893,1919]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMSA P1676/T4520), length 4.12 mm, width 1.63 mm.
<emphasis id="B97EEAE8FB7E7A4BCFCB34CF01F7F84D" bold="true" box="[189,210,1928,1954]" pageId="26" pageNumber="27">A</emphasis>
. Aperture view.
<emphasis id="B97EEAE8FB7E7A4BCEEB34CF0095F84D" bold="true" box="[413,432,1928,1954]" pageId="26" pageNumber="27">B</emphasis>
. Side view.
<emphasis id="B97EEAE8FB7E7A4BCD3134CF0379F84D" bold="true" box="[583,604,1928,1954]" pageId="26" pageNumber="27">C</emphasis>
. View into aperture.
<emphasis id="B97EEAE8FB7E7A4BCC2B34CF0257F84D" bold="true" box="[861,882,1928,1954]" pageId="26" pageNumber="27">D</emphasis>
. Oblique view of base showing umbilicus.
<emphasis id="B97EEAE8FB7E7A4BCFCB34EC01F4F82A" bold="true" box="[189,209,1963,1989]" pageId="26" pageNumber="27">E</emphasis>
. Specimen from Eastern Cape population (ELMD 18754), length 3.57 mm, width 1.55 mm. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3106571FB7F7A4ACFCB324F0056FE9B" pageId="27" pageNumber="28" type="biology_ecology">
<paragraph id="8BB536FAFB7F7A4ACFCB324F0038FECD" blockId="27.[189,285,264,290]" box="[189,285,264,290]" pageId="27" pageNumber="28">
<heading id="D0FD8196FB7F7A4ACFCB324F0038FECD" bold="true" box="[189,285,264,290]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B97EEAE8FB7F7A4ACFCB324F0038FECD" bold="true" box="[189,285,264,290]" pageId="27" pageNumber="28">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB7F7A4ACFCB32700056FE9B" blockId="27.[189,1398,311,372]" pageId="27" pageNumber="28">
<collectingRegion id="49CEF818FB7F7A4ACFCB32700050FEBE" box="[189,373,311,337]" country="South Africa" name="KwaZulu-Natal" pageId="27" pageNumber="28">KwaZulu-Natal</collectingRegion>
Scarp Forest and Pondoland Scarp Forest (
<bibRefCitation id="EF9B4B0BFB7F7A4ACCF632700553FEBE" author="Mucina L. &amp; Abbott A. &amp; Tichy L." box="[896,1142,311,337]" pageId="27" pageNumber="28" pagination="91 - 124" refId="ref16214" refString="Mucina L., Abbott A. &amp; Tichy L. 2018 a. Classification of Pondoland scarp forests. In: Mucina L. (ed.) Vegetation Survey and Classification of Subtropical Forests of Southern Africa: 91 - 124. Geobotany Studies series. Springer, Cham, Switzerland. https: // doi. org / 10.1007 / 978 - 3 - 319 - 67831 - 3 _ 5" type="book chapter" year="2018">
Mucina
<emphasis id="B97EEAE8FB7F7A4ACC91327F0501FEBE" box="[999,1060,311,337]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2018a
</bibRefCitation>
,
<bibRefCitation id="EF9B4B0BFB7F7A4ACBF2327005F7FEBD" author="Mucina L. &amp; Lotter M. C. &amp; Tichy L. &amp; Siebert S. J. &amp; Scott-Shaw C. R." box="[1156,1234,311,338]" pageId="27" pageNumber="28" pagination="125 - 225" refId="ref16288" refString="Mucina L., Lotter M. C., Tichy L., Siebert S. J. &amp; Scott-Shaw C. R. 2018 b. Classification of the Eastern scarp forests. In: Mucina L. (ed.) Vegetation Survey and Classification of Subtropical Forests of Southern Africa: 125 - 225. Geobotany Studies series. Springer, Cham, Switzerland. https: // doi. org / 10.1007 / 978 - 3 - 319 - 67831 - 3 _ 6" type="book chapter" year="2018">2018b</bibRefCitation>
); in leaf-litter and under logs.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB7F7A4ACFCB32DB037BFC23" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BB536FAFB7F7A4ACFCB32DB000AFE59" blockId="27.[189,303,412,438]" box="[189,303,412,438]" pageId="27" pageNumber="28">
<heading id="D0FD8196FB7F7A4ACFCB32DB000AFE59" bold="true" box="[189,303,412,438]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B97EEAE8FB7F7A4ACFCB32DB000AFE59" bold="true" box="[189,303,412,438]" pageId="27" pageNumber="28">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB7F7A4ACFCB328D03CBFD9E" blockId="27.[189,1401,458,626]" pageId="27" pageNumber="28">
Specimens collected at Mkambati Nature Reserve are slightly smaller, appear to have one less whorl and have a small basal tooth to the left of centre, absent in all the specimens collected in KwaZulu- Natal. In other respects specimens from both areas appear to be identical. The
<collectingRegion id="49CEF818FB7F7A4ACB20315605D4FDC4" box="[1110,1265,529,555]" country="South Africa" name="Eastern Cape" pageId="27" pageNumber="28">Eastern Cape</collectingRegion>
population is considered here to be
<taxonomicName id="4C0A4D79FB7F7A4ACE9E31730381FDA1" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[488,676,564,590]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7F7A4ACE9E31730381FDA1" box="[488,676,564,590]" italics="true" pageId="27" pageNumber="28">Gulella maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7F7A4ACDD83173022BFDA1" box="[686,782,564,590]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
, but due to the differences mentioned above, these specimens are not included in the
<typeStatus id="54B18858FB7F7A4ACD3A311F035AFD9D" box="[588,639,600,626]" pageId="27" pageNumber="28">type</typeStatus>
material.
</paragraph>
<paragraph id="8BB536FAFB7F7A4ACFCB31DF037BFC23" blockId="27.[188,1399,664,972]" pageId="27" pageNumber="28">
<taxonomicName id="4C0A4D79FB7F7A4ACFCB31DF005EFD5D" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[189,379,664,690]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7F7A4ACFCB31DF005EFD5D" box="[189,379,664,690]" italics="true" pageId="27" pageNumber="28">Gulella maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7F7A4ACEFE31DE00C2FD5C" box="[392,487,665,691]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
superficially resembles
<taxonomicName id="4C0A4D79FB7F7A4ACC6531DE02BBFD5D" baseAuthorityName="Sturany" baseAuthorityYear="1898" box="[787,926,664,691]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="pentheri">
<emphasis id="B97EEAE8FB7F7A4ACC6531DE02BBFD5D" box="[787,926,664,691]" italics="true" pageId="27" pageNumber="28">G. pentheri</emphasis>
</taxonomicName>
, but is larger and has axial sculpture, while
<taxonomicName id="4C0A4D79FB7F7A4ACE7D31FB00B3FD39" baseAuthorityName="Sturany" baseAuthorityYear="1898" box="[267,406,700,726]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="pentheri">
<emphasis id="B97EEAE8FB7F7A4ACE7D31FB00B3FD39" box="[267,406,700,726]" italics="true" pageId="27" pageNumber="28">G. pentheri</emphasis>
</taxonomicName>
is narrower and smooth. The parietal lamella of
<taxonomicName id="4C0A4D79FB7F7A4ACC8931FB05A7FD3A" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[1023,1154,700,726]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7F7A4ACC8931FB05A7FD3A" box="[1023,1154,700,726]" italics="true" pageId="27" pageNumber="28">G. maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7F7A4ACBF831FB05CBFD39" box="[1166,1262,700,726]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
extends to the edge of the labrum and curves strongly before running into the aperture, a feature not present in
<taxonomicName id="4C0A4D79FB7F7A4ACFCB30450067FCF3" baseAuthorityName="Sturany" baseAuthorityYear="1898" box="[189,322,770,796]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="pentheri">
<emphasis id="B97EEAE8FB7F7A4ACFCB30450067FCF3" box="[189,322,770,796]" italics="true" pageId="27" pageNumber="28">G. pentheri</emphasis>
</taxonomicName>
. In respect of its apertural dentition and the short axial riblets,
<taxonomicName id="4C0A4D79FB7F7A4ACB6B304505BFFCF3" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[1053,1178,770,796]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7F7A4ACB6B304505BFFCF3" box="[1053,1178,770,796]" italics="true" pageId="27" pageNumber="28">G. maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7F7A4ACBD7304405DEFCF2" box="[1185,1275,771,797]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C0A4D79FB7F7A4ACFCA306100ADFCD0" baseAuthorityName="Burnup" baseAuthorityYear="1914" box="[188,392,805,832]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="inhluzaniensis">
<emphasis id="B97EEAE8FB7F7A4ACFCA306100ADFCD0" box="[188,392,805,832]" italics="true" pageId="27" pageNumber="28">G. inhluzaniensis</emphasis>
</taxonomicName>
, but in the present species the columella lamella is much larger and the last whorl has much stronger axial ribs.
<taxonomicName id="4C0A4D79FB7F7A4ACE84300F038BFC8D" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[498,686,840,866]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7F7A4ACE84300F038BFC8D" box="[498,686,840,866]" italics="true" pageId="27" pageNumber="28">Gulella maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7F7A4ACDC3300E0234FC8C" box="[693,785,841,867]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
also resembles
<taxonomicName id="4C0A4D79FB7F7A4ACCA2300E0568FC8D" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[980,1101,840,867]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="abbotti" status="sp. nov.">
<emphasis id="B97EEAE8FB7F7A4ACCA2300E0568FC8D" box="[980,1101,840,867]" italics="true" pageId="27" pageNumber="28">G. abbotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7F7A4ACB22300E0596FC8C" box="[1108,1203,841,867]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
, but the latter is smaller, narrower and has a more substantial labral tooth as well as a basal tooth. Species in the
<taxonomicName id="4C0A4D79FB7F7A4ACA68302B0020FC46" baseAuthorityName="Craven" baseAuthorityYear="1881" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="infans">
<emphasis id="B97EEAE8FB7F7A4ACA68302B0020FC46" italics="true" pageId="27" pageNumber="28">Gulella infans</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EF9B4B0BFB7F7A4ACE1530C803A2FC46" author="Herbert D. &amp; Kilburn D." box="[355,647,911,937]" pageId="27" pageNumber="28" refId="ref15954" refString="Herbert D. &amp; Kilburn D. 2004. Field Guide to the Land Snails and Slugs of Eastern South Africa. Natal Museum, Pietermaritzburg, South Africa." type="book" year="2004">Herbert &amp; Kilburn 2004</bibRefCitation>
) are larger, generally broader, have a less prominent columella lamella and weaker axial sculpture.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB7F7A4ACFCB30B40318FB02" pageId="27" pageNumber="28" type="conservation">
<paragraph id="8BB536FAFB7F7A4ACFCB30B40040FBE2" blockId="27.[189,357,1011,1037]" box="[189,357,1011,1037]" pageId="27" pageNumber="28">
<heading id="D0FD8196FB7F7A4ACFCB30B40040FBE2" bold="true" box="[189,357,1011,1037]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B97EEAE8FB7F7A4ACFCB30B40040FBE2" bold="true" box="[189,357,1011,1037]" pageId="27" pageNumber="28">Conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB7F7A4ACFCB37650318FB02" blockId="27.[188,1401,1058,1261]" pageId="27" pageNumber="28">
<taxonomicName id="4C0A4D79FB7F7A4ACFCB37650052FBD3" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[189,375,1058,1084]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="maraisi" status="sp. nov.">
<emphasis id="B97EEAE8FB7F7A4ACFCB37650052FBD3" box="[189,375,1058,1084]" italics="true" pageId="27" pageNumber="28">Gulella maraisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB7F7A4ACE09376400FEFBD2" box="[383,475,1059,1085]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
appears to be a very rare species with few records. It has been collected alive at only two localities approx.
<quantity id="4CF29B1FFB7F7A4ACD5037020357FBB0" box="[550,626,1093,1120]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="27" pageNumber="28" unit="km" value="75.0">75 km</quantity>
apart. One of these is under threat from mining since the lower Hlokohloko Valley is being sacrificed as a waste rock dump. It has not been found in Oribi Gorge Nature Reserve immediately to the west. The other locality is in a protected area, the Mkambati Nature Reserve. It has been collected in beach drift at the mouth of the Mtamvuna River, but as yet not in the Mtamvuna Gorge Nature Reserve upstream.
</paragraph>
</subSubSection>
</treatment>
</document>
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