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<document id="66BEBE35CBF79AB26AABAFBE23378A61" ID-DOI="10.11646/zootaxa.3626.4.10" ID-GBIF-Dataset="5bb77bb6-2b4b-459e-8957-67c35841c845" ID-ISSN="1175-5326" ID-Zenodo-Dep="220807" ID-ZooBank="D3BC0BEC-1061-4E54-AD15-4320F8639672" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460582709363" checkinUser="plazi" docAuthor="Iwan, Dariusz &amp; Schimrosczyk, David" docDate="2013" docId="03A387BFF8172E53FF5E81770C43F997" docLanguage="en" docName="zt03626p575.pdf" docOrigin="Zootaxa 3626 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helibatus banaszkiewiczae Iwan &amp; Schimrosczyk, 2013, sp. nov." docType="treatment" docVersion="6" lastPageNumber="571" masterDocId="FF9AFFC7F8152E51FFC980500E11FFE9" masterDocTitle="A second species of the genus Helibatus Mulsant &amp; Rey, 1859 (Coleoptera: Tenebrionidae: Opatrina)" masterLastPageNumber="575" masterPageNumber="569" pageNumber="571" updateTime="1698324594669" updateUser="plazi">
<mods:mods id="F204352CA1D243EE1E84FB2913D56D86" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="520F7130B0E6A391CAE8B6F2F87EDEDD">
<mods:title id="CAFA16E3A65AF854AC7D8584B54A9543">A second species of the genus Helibatus Mulsant &amp; Rey, 1859 (Coleoptera: Tenebrionidae: Opatrina)</mods:title>
</mods:titleInfo>
<mods:name id="AA7C85A46B5BD33611C4E1663EEFD8FB" type="personal">
<mods:role id="4FDC8F399429CE5987114CE6DC79D32D">
<mods:roleTerm id="21A271D63A05338D57FDF11615A3FF59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D7C7BB5CF35D328E20C8ADED8DE4FEE5">Iwan, Dariusz</mods:namePart>
</mods:name>
<mods:name id="C29104D7E0ED696B3F6029D54377155C" type="personal">
<mods:role id="DF71859031AFABAF7116430B88E72218">
<mods:roleTerm id="9E70554912D6272CCF29618696EB0260">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="341DD570BC2E3F91A9AD8C5FC399E52B">Schimrosczyk, David</mods:namePart>
</mods:name>
<mods:typeOfResource id="34FB21612C92BC2C7582D1DA433D4022">text</mods:typeOfResource>
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<mods:title id="FB428A57CD061EA7738EB2D5AC2038B0">Zootaxa</mods:title>
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<mods:part id="77ABCC38E68429AC7589F3406E0B80DE">
<mods:date id="2C222A65FDEA2EC74B4F6D910C5C569E">2013</mods:date>
<mods:detail id="4231B32847C625EB0685F39C4D4CAC79" type="volume">
<mods:number id="1202001037AA6C1620F07A88D3BC773B">3626</mods:number>
</mods:detail>
<mods:detail id="47E80F07973CBA20732CC944BD8F98D2" type="issue">
<mods:number id="199A3E673893B16DFD3831AEF07CF926">4</mods:number>
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<mods:classification id="3FB735DD69A4938D0976540D672739D2">journal article</mods:classification>
<mods:identifier id="04868B17CA30E44AEE9B736B0DB3728B" type="DOI">10.11646/zootaxa.3626.4.10</mods:identifier>
<mods:identifier id="58A34F47941AC554043C26D7ECAAA861" type="GBIF-Dataset">5bb77bb6-2b4b-459e-8957-67c35841c845</mods:identifier>
<mods:identifier id="69CC45A5EA8B056A46A970F389294382" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="258D647AD176DEA1B2BCD0E8416D331C" type="Zenodo-Dep">220807</mods:identifier>
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</mods:mods>
<treatment id="03A387BFF8172E53FF5E81770C43F997" ID-DOI="http://doi.org/10.5281/zenodo.6150947" ID-GBIF-Taxon="119563976" ID-Zenodo-Dep="6150947" LSID="urn:lsid:plazi:treatment:03A387BFF8172E53FF5E81770C43F997" httpUri="http://treatment.plazi.org/id/03A387BFF8172E53FF5E81770C43F997" lastPageNumber="571" pageId="2" pageNumber="571">
<subSubSection id="C3106522F8172E53FF5E81770C2EFEA8" box="[151,575,295,321]" pageId="2" pageNumber="571" type="nomenclature">
<paragraph id="8BB536A9F8172E53FF5E81770C2EFEA8" blockId="2.[151,575,295,321]" box="[151,575,295,321]" pageId="2" pageNumber="571">
<heading id="D0FD81C5F8172E53FF5E81770C2EFEA8" bold="true" box="[151,575,295,321]" fontSize="11" level="1" pageId="2" pageNumber="571" reason="1">
<emphasis id="B97EEABBF8172E53FF5E81770C2EFEA8" bold="true" box="[151,575,295,321]" pageId="2" pageNumber="571">
<taxonomicName id="4C0A4D2AF8172E53FF5E81770FCBFEA8" box="[151,474,295,321]" class="Insecta" family="Tenebrionidae" genus="Helibatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="banaszkiewiczae" status="sp. nov.">
<emphasis id="B97EEABBF8172E53FF5E81770FCBFEA8" bold="true" box="[151,474,295,321]" italics="true" pageId="2" pageNumber="571">Helibatus banaszkiewiczae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C0F8172E53FE2881770C2EFEA8" box="[481,575,295,321]" pageId="2" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3106522F8172E53FF5E813F0D51FE43" pageId="2" pageNumber="571" type="etymology">
<paragraph id="8BB536A9F8172E53FF5E813F0D51FE43" blockId="2.[151,1437,366,1663]" pageId="2" pageNumber="571">
<emphasis id="B97EEABBF8172E53FF5E813F0F0FFE6E" bold="true" box="[151,286,367,391]" pageId="2" pageNumber="571">Etymology.</emphasis>
This new species is named in honor of the late Margaret Banaszkiewicz - our colleague and excellent young specialist of
<taxonomicName id="4C0A4D2AF8172E53FEB881C20C00FE43" box="[369,529,402,426]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="571" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
who passed away in 2010.
</paragraph>
</subSubSection>
<subSubSection id="C3106522F8172E53FF0E81E70FFEFE1B" pageId="2" pageNumber="571" type="materials_examined">
<paragraph id="8BB536A9F8172E53FF0E81E70FFEFE1B" blockId="2.[151,1437,366,1663]" pageId="2" pageNumber="571">
<emphasis id="B97EEABBF8172E53FF0E81E70F68FE26" bold="true" box="[199,377,438,463]" pageId="2" pageNumber="571">
<typeStatus id="54B1880BF8172E53FF0E81E70F13FE26" box="[199,258,439,463]" pageId="2" pageNumber="571">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B1880BF8172E53FE4C81E60FE3FE27" box="[389,498,438,462]" pageId="2" pageNumber="571" type="holotype">Holotype</typeStatus>
: 13 &quot;Marble Hall,
<date id="FFB41069F8172E53FD1281E60D3CFE27" box="[731,813,438,462]" pageId="2" pageNumber="571" value="1962-05">V.1962</date>
, TV., Rorke&quot; (TMNH).
<typeStatus id="54B1880BF8172E53FB8481E70AAFFE26" box="[1101,1214,439,463]" pageId="2" pageNumber="571" type="paratype">Paratypes</typeStatus>
: 2Ƥ &quot;Marble Hall,
<date id="FFB41069F8172E53FF5E818A0EF8FE1B" box="[151,233,474,498]" pageId="2" pageNumber="571" value="1962-05">V.1962</date>
, TV., Rorke&quot; (TMNH).
</paragraph>
</subSubSection>
<subSubSection id="C3106522F8172E53FF0E81AE0D6BFDFF" box="[199,890,510,535]" pageId="2" pageNumber="571" type="distribution">
<paragraph id="8BB536A9F8172E53FF0E81AE0D6BFDFF" blockId="2.[151,1437,366,1663]" box="[199,890,510,535]" pageId="2" pageNumber="571">
<emphasis id="B97EEABBF8172E53FF0E81AE0F71FDFE" bold="true" box="[199,352,510,535]" pageId="2" pageNumber="571">Distribution.</emphasis>
<collectingCountry id="F31D7639F8172E53FEAE81AE0C93FDFF" box="[359,642,510,534]" name="South Africa" pageId="2" pageNumber="571">Republic of South Africa</collectingCountry>
(Limpopo Province).
</paragraph>
</subSubSection>
<subSubSection id="C3106522F8172E53FF0E82720B45FD6B" pageId="2" pageNumber="571" type="diagnosis">
<paragraph id="8BB536A9F8172E53FF0E82720B45FD6B" blockId="2.[151,1437,366,1663]" pageId="2" pageNumber="571">
<emphasis id="B97EEABBF8172E53FF0E82720F50FDD2" bold="true" box="[199,321,546,571]" pageId="2" pageNumber="571">Diagnosis.</emphasis>
The species differs from
<taxonomicName id="4C0A4D2AF8172E53FDA282740CC3FDD2" box="[619,722,548,571]" class="Insecta" family="Tenebrionidae" genus="Helibatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B97EEABBF8172E53FDA282740CC3FDD2" box="[619,722,548,571]" italics="true" pageId="2" pageNumber="571">H. morio</emphasis>
</taxonomicName>
in the structure of the metaventral process (simple, not toothlike),
<emphasis id="B97EEABBF8172E53FF1182160EFDFDB7" box="[216,236,582,606]" italics="true" pageId="2" pageNumber="571">cf</emphasis>
. Figs. 10 and 9; punctation on the pronotal disc (punctures fine, never confluent),
<emphasis id="B97EEABBF8172E53FB4382160A8FFDB7" box="[1162,1182,582,606]" italics="true" pageId="2" pageNumber="571">cf</emphasis>
.
<figureCitation id="13312A2CF8172E53FB6582160B2FFDB7" box="[1196,1342,582,606]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[203,1376,196,1949]" captionTargetId="figure@3.[203,1377,195,1949]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Helibatus banaszkiewiczae sp. nov., male (1, 2, 3, 4, 6, 8), H. morio Mulsant &amp; Rey, 1859, male (5, 7). Head, dorsal (1). Mentum (2). Antenna (3). Prosternal process (4). Pronotum (5, 6). Elytral base (7, 8)." httpUri="https://zenodo.org/record/220808/files/figure.png" pageId="2" pageNumber="571">Figs. 6 and 5</figureCitation>
; and the structure of the elytron (intervals almost flat, not convex, and striae never in form of sulci),
<emphasis id="B97EEABBF8172E53FB53823B0ABFFD6A" box="[1178,1198,619,643]" italics="true" pageId="2" pageNumber="571">cf</emphasis>
.
<figureCitation id="13312A2CF8172E53FB75823A0B41FD6B" box="[1212,1360,618,642]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[203,1376,196,1949]" captionTargetId="figure@3.[203,1377,195,1949]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Helibatus banaszkiewiczae sp. nov., male (1, 2, 3, 4, 6, 8), H. morio Mulsant &amp; Rey, 1859, male (5, 7). Head, dorsal (1). Mentum (2). Antenna (3). Prosternal process (4). Pronotum (5, 6). Elytral base (7, 8)." httpUri="https://zenodo.org/record/220808/files/figure.png" pageId="2" pageNumber="571">Figs. 8 and 7</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3106522F8172E53FF0E82DE0C43F997" pageId="2" pageNumber="571" type="description">
<paragraph id="8BB536A9F8172E53FF0E82DE0A35FD07" blockId="2.[151,1437,366,1663]" pageId="2" pageNumber="571">
<emphasis id="B97EEABBF8172E53FF0E82DE0F48FD4E" bold="true" box="[199,345,654,679]" pageId="2" pageNumber="571">Description.</emphasis>
Body length
<emphasis id="B97EEABBF8172E53FE2682C00C18FD4E" box="[495,521,656,679]" italics="true" pageId="2" pageNumber="571">ca</emphasis>
. 9.6–11.0 mm [holotype—11.0 mm] (
<figureCitation id="13312A2CF8172E53FC7A82DE0A17FD4F" box="[947,1030,654,678]" captionStart="FIGURE 14 – 16" captionStartId="5.[151,250,1979,2002]" captionTargetBox="[178,1421,219,1938]" captionTargetId="figure@5.[158,1443,193,1969]" captionTargetPageId="5" captionText="FIGURE 14 – 16. Helibatus morio Mulsant &amp; Rey, 1859, male (14); H. banaszkiewiczae sp. nov., male (15). Map of distribution (16)." httpUri="https://zenodo.org/record/220809/files/figure.png" pageId="2" pageNumber="571">Fig. 15</figureCitation>
). Body surface shining, dark brown, legs and antennae brown. Pronotum length/breadth ratio
<emphasis id="B97EEABBF8172E53FCD082E40D22FD22" box="[793,819,692,715]" italics="true" pageId="2" pageNumber="571">ca</emphasis>
. 0.64, elytra length/breadth ratio
<emphasis id="B97EEABBF8172E53FB6782E40AD8FD22" box="[1198,1225,692,715]" italics="true" pageId="2" pageNumber="571">ca</emphasis>
. 1.24–1.27, length ratio elytra/pronotum
<emphasis id="B97EEABBF8172E53FE4582880FB6FD06" box="[396,423,728,751]" italics="true" pageId="2" pageNumber="571">ca</emphasis>
. 2.00–2.03, breadth ratio elytra/pronotum
<emphasis id="B97EEABBF8172E53FC4D82880D8EFD06" box="[900,927,728,751]" italics="true" pageId="2" pageNumber="571">ca</emphasis>
. 1.02–1.03.
</paragraph>
<paragraph id="8BB536A9F8172E53FF5E82AA0DAEFC2F" blockId="2.[151,1437,366,1663]" pageId="2" pageNumber="571">
Genae protruding outwards, slightly wider than eyes. Clypeus on same level as frons, distinctly emarginate (ratio of length of clypeus/depth of emargination
<emphasis id="B97EEABBF8172E53FDA183700C93FCDE" box="[616,642,800,823]" italics="true" pageId="2" pageNumber="571">ca</emphasis>
. 2.08). Head covered with tubercles, more elongate on frons than on clypeus (
<figureCitation id="13312A2CF8172E53FF3483120F50FCB3" box="[253,321,834,858]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[203,1376,196,1949]" captionTargetId="figure@3.[203,1377,195,1949]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Helibatus banaszkiewiczae sp. nov., male (1, 2, 3, 4, 6, 8), H. morio Mulsant &amp; Rey, 1859, male (5, 7). Head, dorsal (1). Mentum (2). Antenna (3). Prosternal process (4). Pronotum (5, 6). Elytral base (7, 8)." httpUri="https://zenodo.org/record/220808/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
). Antennae (
<figureCitation id="13312A2CF8172E53FE1E83120C0AFCB3" box="[471,539,834,858]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[203,1376,196,1949]" captionTargetId="figure@3.[203,1377,195,1949]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Helibatus banaszkiewiczae sp. nov., male (1, 2, 3, 4, 6, 8), H. morio Mulsant &amp; Rey, 1859, male (5, 7). Head, dorsal (1). Mentum (2). Antenna (3). Prosternal process (4). Pronotum (5, 6). Elytral base (7, 8)." httpUri="https://zenodo.org/record/220808/files/figure.png" pageId="2" pageNumber="571">Fig. 3</figureCitation>
) – ratio of length/width of 2–11 antennomeres as follow: 0.6/0.8; 0.8/0.8; 0.6/ 0.8; 0.6/0.8; 0.6/0.9; 0.6/1.1; 0.7/1.4; 0.6/1.4; 0.7/1.3; 0.9/1.2; antennomeres 7–11 widened, transverse. Lateral margins of mentum together with medial carina form sharp apical spike. Lateral wings of mentum small,visible. Terminal maxillary palpomere moderately widened, securiform (
<figureCitation id="13312A2CF8172E53FCA583FE0DA0FC2F" box="[876,945,942,966]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[203,1376,196,1949]" captionTargetId="figure@3.[203,1377,195,1949]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Helibatus banaszkiewiczae sp. nov., male (1, 2, 3, 4, 6, 8), H. morio Mulsant &amp; Rey, 1859, male (5, 7). Head, dorsal (1). Mentum (2). Antenna (3). Prosternal process (4). Pronotum (5, 6). Elytral base (7, 8)." httpUri="https://zenodo.org/record/220808/files/figure.png" pageId="2" pageNumber="571">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BB536A9F8172E53FF0083830F15FAE4" blockId="2.[151,1437,366,1663]" pageId="2" pageNumber="571">
Pronotum transverse, widest in middle, regularly narrowed to posterior angle. Posterior angles obtusely rectangular. Pronotal disc regularly convex, with distinct and roundish punctures on disc, gradually replaced by elongate tubercles on sides. Pronotal border interrupted in middle of anterior and basal margins. Border of lateral margins entire and very narrow (breadth ratio border/base of 3rd antennomere
<emphasis id="B97EEABBF8172E53FBED84120A54FBB0" box="[1060,1093,1090,1113]" italics="true" pageId="2" pageNumber="571">ca.</emphasis>
0.29). Pronotal base straight, protruding medially slightly beyond level of posterior angles. Lateral gutters of pronotum narrow, bare, without punctures or tubercles (
<figureCitation id="13312A2CF8172E53FE6F84D90FFCFB48" box="[422,493,1161,1185]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[203,1376,196,1949]" captionTargetId="figure@3.[203,1377,195,1949]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Helibatus banaszkiewiczae sp. nov., male (1, 2, 3, 4, 6, 8), H. morio Mulsant &amp; Rey, 1859, male (5, 7). Head, dorsal (1). Mentum (2). Antenna (3). Prosternal process (4). Pronotum (5, 6). Elytral base (7, 8)." httpUri="https://zenodo.org/record/220808/files/figure.png" pageId="2" pageNumber="571">Fig. 6</figureCitation>
). Anterior margin of prosternum bordered. Prosternal process subparallel covered with roundish tubercles and numerous, long, gold-yellow setae. Border of prosternal process disappearing at apex (
<figureCitation id="13312A2CF8172E53FF5684810EF7FB00" box="[159,230,1233,1257]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[203,1376,196,1949]" captionTargetId="figure@3.[203,1377,195,1949]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Helibatus banaszkiewiczae sp. nov., male (1, 2, 3, 4, 6, 8), H. morio Mulsant &amp; Rey, 1859, male (5, 7). Head, dorsal (1). Mentum (2). Antenna (3). Prosternal process (4). Pronotum (5, 6). Elytral base (7, 8)." httpUri="https://zenodo.org/record/220808/files/figure.png" pageId="2" pageNumber="571">Fig. 4</figureCitation>
). Pronotal hypomera covered with elongated tubercles. Border in form of bare stripe, without punctures or tubercles.
</paragraph>
<paragraph id="8BB536A9F8172E53FF0085490DF7FA90" blockId="2.[151,1437,366,1663]" pageId="2" pageNumber="571">
Elytra widest in middle. Elytron with 9 striae. Punctures on striae larger than on intervals. Intervals on disc weakly convex, almost flat; intervals distinctly sparsly punctate, apex and lateral sides of elytra with fine tubercles. Elytral base bordered, humeri rectangular, not protruding outwards (
<figureCitation id="13312A2CF8172E53FC5A85310DC9FA90" box="[915,984,1377,1401]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[203,1376,196,1949]" captionTargetId="figure@3.[203,1377,195,1949]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Helibatus banaszkiewiczae sp. nov., male (1, 2, 3, 4, 6, 8), H. morio Mulsant &amp; Rey, 1859, male (5, 7). Head, dorsal (1). Mentum (2). Antenna (3). Prosternal process (4). Pronotum (5, 6). Elytral base (7, 8)." httpUri="https://zenodo.org/record/220808/files/figure.png" pageId="2" pageNumber="571">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BB536A9F8172E53FF0085D80A4EFA49" blockId="2.[151,1437,366,1663]" box="[201,1119,1413,1440]" pageId="2" pageNumber="571">Wings absent. Epipleuron flat, strongly widened at level of 5th abdominal ventrite.</paragraph>
<paragraph id="8BB536A9F8172E53FF0085FC0C22F9B0" blockId="2.[151,1437,366,1663]" pageId="2" pageNumber="571">Abdominal ventrites distinctly transverse (Fig. 13); apical parts of ventrites 2–5 with regularly distributed, small tubercles, replaced by large punctures towards base; 1st visible abdominal ventrite covered only with small tubercles; last abdominal ventrite flat (Fig. 11). Metaventrite covered with long, gold-yellow setae. Metaventral process simple, without denticulation. Process of 1st abdominal ventrite in males projecting forward beyond metaventrite apical margin (Fig. 10).</paragraph>
<paragraph id="8BB536A9F8172E53FF0086370C43F997" blockId="2.[151,1437,366,1663]" box="[201,594,1638,1663]" pageId="2" pageNumber="571">Legs non-dimorphic (Fig. 12A–C).</paragraph>
</subSubSection>
</treatment>
</document>
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