treatments-xml/data/03/A3/87/03A387B9FF46FF6DFDF01E8FFD9EA443.xml
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<mods:title id="FD12DB45D727DC46DB42C2B0724A632D">Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)</mods:title>
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<mods:namePart id="5FBA5C9EA397AFD2FE397D4B01AF2700">Zallot, Enrico</mods:namePart>
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<mods:affiliation id="EEF90EC550F1B495B1544F7E8ABBB371">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. &amp; Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands.</mods:affiliation>
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<emphasis id="B97EEABDFF46FF6BFDF01E8FFD72A5C2" bold="true" box="[553,711,976,1003]" italics="true" pageId="130" pageNumber="131">Cochlostoma</emphasis>
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(
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<figureCitation id="13312A2AFF46FF6BFD851EAAFD77A239" box="[604,706,1013,1040]" captionStart="Fig" captionStartId="118.[189,232,1856,1882]" captionTargetBox="[189,1398,1147,1827]" captionTargetId="figure-199@118.[189,1398,1147,1827]" captionTargetPageId="118" captionText="Fig. 105. Distribution of samples with undetermined position in the phylogenetic trees: white= Cochlostoma (T.) macei (Bourguignat, 1870); purple =C. (T.) sturanii (Wagner, 1897); orange=C. (T.) kleciaki (Braun, 1887); yellow=C. (T.) arnautorum (Wagner, 1906); red =C. (T.) atlanticum (Bourguignat, 1868); green =C. (T.) euboicum (Westerlund, 1885); blue =C. (T.) martensianum (Möllendorff, 1873); magenta= C. (T.) simrothi (Caziot, 1908); cyan =C. (T.) fontqueri Haas, 1924." figureDoi="http://doi.org/10.5281/zenodo.10853918" httpUri="https://zenodo.org/record/10853918/files/figure.png" pageId="130" pageNumber="131">Figs 105</figureCitation>
(orange dots), 114116
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<emphasis id="B97EEABDFF46FF6BFF641966FE3BA27B" box="[189,398,1080,1106]" italics="true" pageId="130" pageNumber="131">Pomatias kleciaki</emphasis>
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; “
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Bttg
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80”;
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<emphasis id="B97EEABDFF46FF6BFF6419B2FE3BA32E" bold="true" box="[189,398,1261,1287]" pageId="130" pageNumber="131">Other specimens</emphasis>
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<location id="8ED56074FF46FF6BFACF1842FF40A372" LSID="urn:lsid:plazi:treatment:03A387B9FF46FF6DFDF01E8FFD9EA443:8ED56074FF46FF6BFACF1842FF40A372" country="Croatia" latitude="42.9637" longLatPrecision="6" longitude="17.3084" name="Zupanje Selo" pageId="130" pageNumber="131">Županje Selo</location>
;
<geoCoordinate id="EE3E5068FF46FF6BFED8181EFE3EA375" box="[257,395,1345,1372]" degrees="42.9637" direction="north" orientation="latitude" pageId="130" pageNumber="131" precision="5" value="42.9637">42.9637° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FF46FF6BFE4E181EFDABA375" box="[407,542,1345,1372]" degrees="17.3084" direction="east" orientation="longitude" pageId="130" pageNumber="131" precision="5" value="17.3084">17.3084° E</geoCoordinate>
; 1985;
<collectorName id="26FF5379FF46FF6BFDAC181EFD1CA372" box="[629,681,1345,1371]" pageId="130" pageNumber="131">Kiss</collectorName>
and
<collectorName id="26FF5379FF46FF6BFD3B181DFCFCA372" box="[738,841,1345,1372]" pageId="130" pageNumber="131">L. Pintér</collectorName>
leg.;
<specimenCode id="DBAC9ED4FF46FF6BFC52181EFBF4A375" box="[907,1089,1345,1372]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="130" pageNumber="131">HNHM105302</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB536AFFF46FF6BFF6418DBFEE8A3B4" blockId="130.[189,349,1411,1438]" box="[189,349,1411,1438]" pageId="130" pageNumber="131">
<heading id="D0FD81C3FF46FF6BFF6418DBFEE8A3B4" bold="true" box="[189,349,1411,1438]" fontSize="11" level="3" pageId="130" pageNumber="131" reason="3">
<emphasis id="B97EEABDFF46FF6BFF6418DBFEE8A3B4" bold="true" box="[189,349,1411,1438]" pageId="130" pageNumber="131">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFF46FF6BFF6418EBFCDAA3E4" blockId="130.[189,879,1459,1486]" box="[189,879,1459,1486]" pageId="130" pageNumber="131">
<collectingCountry id="F31D763FFF46FF6BFF6418EBFE89A3E7" box="[189,316,1460,1486]" name="Croatia" pageId="130" pageNumber="131">CROATIA</collectingCountry>
• “Lesina insula” (Hvar island,
<collectingRegion id="49CEF84DFF46FF6BFD6818ECFCD1A3E4" box="[689,868,1459,1485]" country="Croatia" name="Splitsko-Dalmatinska" pageId="130" pageNumber="131">Split-Dalmatia</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3106524FF46FF6BFF6418A9FCB2A1C1" pageId="130" pageNumber="131" type="description">
<paragraph id="8BB536AFFF46FF6BFF6418A9FEFBA039" blockId="130.[189,334,1526,1552]" box="[189,334,1526,1552]" pageId="130" pageNumber="131">
<heading id="D0FD81C3FF46FF6BFF6418A9FEFBA039" bold="true" box="[189,334,1526,1552]" fontSize="11" level="3" pageId="130" pageNumber="131" reason="3">
<emphasis id="B97EEABDFF46FF6BFF6418A9FEFBA039" bold="true" box="[189,334,1526,1552]" pageId="130" pageNumber="131">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFF46FF6BFF641B79FCEBA0A0" blockId="130.[189,1398,1574,1673]" pageId="130" pageNumber="131">
<smallCapsWord id="8D53A073FF46FF6BFF641B79FEB4A016" baselines="1593,1594" box="[189,257,1574,1600]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="130" pageNumber="131">SHELL</smallCapsWord>
. (based on the male
<typeStatus id="54B1880DFF46FF6BFE2C1B79FDD3A069" box="[501,614,1574,1600]" pageId="130" pageNumber="131" type="lectotype">lectotype</typeStatus>
). Teleoconch spotless. Rather strong ribs becoming less strong and closely spaced toward aperture. Very strong lip with indented columellar lobe presumably (
<typeStatus id="54B1880DFF46FF6BFAD01B15FAC3A04D" box="[1289,1398,1610,1636]" pageId="130" pageNumber="131" type="lectotype">lectotype</typeStatus>
only seen in frontal and lateral view) covering umbilicus.
</paragraph>
<paragraph id="8BB536AFFF46FF6BFF641BEDFD71A110" blockId="130.[189,1398,1713,1849]" pageId="130" pageNumber="131">
<smallCapsWord id="8D53A073FF46FF6BFF641BEDFED9A0E3" baselines="1733,1733" box="[189,364,1714,1740]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="130" pageNumber="131">MEASUREMENTS</smallCapsWord>
.
<specimenCount id="9D0CFD26FF46FF6BFE5E1BEDFE28A0E5" box="[391,413,1714,1740]" count="1" pageId="130" pageNumber="131" type="female"></specimenCount>
MIZPASW7082: whorls=8.1, H =
<quantity id="4CF29B4AFF46FF6BFC871BEEFC72A0E5" box="[862,967,1713,1740]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="130" pageNumber="131" unit="mm" value="8.9">8.9 mm</quantity>
, H/W=2.6, roundness= 0.13, ribs inclin.=63°, ribs per mm 1
<superScript id="7C7F9BE7FF46FF6BFDDF1B8AFDA4A0CA" attach="left" box="[518,529,1749,1763]" fontSize="6" pageId="130" pageNumber="131">st</superScript>
whorl = 6, ribs per mm 4
<superScript id="7C7F9BE7FF46FF6BFC9F1B8AFCE6A0CA" attach="left" box="[838,851,1749,1763]" fontSize="6" pageId="130" pageNumber="131">th</superScript>
whorl = 10.
<specimenCount id="9D0CFD26FF46FF6BFC301B89FB9BA0D9" box="[1001,1070,1750,1776]" count="3" pageId="130" pageNumber="131" type="female">3 ♀♀</specimenCount>
Županje Selo: whorls 7.2 7.7, H =
<quantity id="4CF29B4AFF46FF6BFEFA1BA5FE0CA13C" box="[291,441,1786,1813]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.4" metricValueMin="6.1" pageId="130" pageNumber="131" unit="mm" value="6.25" valueMax="6.4" valueMin="6.1">6.16.4 mm</quantity>
, H/W= 2.442.63, roundness =0.150.17, ribs inclin.=5762, ribs per mm 1
<superScript id="7C7F9BE7FF46FF6BFAB21BA5FAC3A121" attach="none" box="[1387,1398,1786,1800]" fontSize="6" pageId="130" pageNumber="131">st</superScript>
whorl =812, ribs per mm 4
<superScript id="7C7F9BE7FF46FF6BFDDF1A41FDA6A105" attach="left" box="[518,531,1822,1836]" fontSize="6" pageId="130" pageNumber="131">th</superScript>
whorl= 1017.
</paragraph>
<paragraph id="8BB536AFFF46FF6BFF641A3DFCB2A1C1" blockId="130.[189,1399,1889,2024]" pageId="130" pageNumber="131">
<smallCapsWord id="8D53A073FF46FF6BFF641A3DFEA1A153" baselines="1909,1909" box="[189,276,1890,1916]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="130" pageNumber="131">FEMALE</smallCapsWord>
<smallCapsWord id="8D53A073FF46FF6BFEFB1A38FECCA153" baselines="1909" box="[290,377,1895,1914]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genital" pageId="130" pageNumber="131">GENITAL</smallCapsWord>
<smallCapsWord id="8D53A073FF46FF6BFE511A38FE69A153" baselines="1909" box="[392,476,1895,1914]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="organs" pageId="130" pageNumber="131">ORGANS</smallCapsWord>
. Poorly preserved specimens from Županje Selo were dissected. Slightly posterior pedunculus of bursa copulatrix, club-shaped, elongated, tortuous seminal receptacle and short loops clustered close to apex of seminal receptacle, junction of uterus gland far from connection between distal oviduct and pedunculus of bursa copulatrix.
</paragraph>
</subSubSection>
<caption id="DF756627FF47FF6AFF641AD6FE3EA1EF" ID-DOI="http://doi.org/10.5281/zenodo.10853937" ID-Zenodo-Dep="10853937" httpUri="https://zenodo.org/record/10853937/files/figure.png" pageId="131" pageNumber="132" startId="131.[189,232,1929,1955]" targetBox="[189,1398,265,1884]" targetPageId="131">
<paragraph id="8BB536AFFF47FF6AFF641AD6FE3EA1EF" blockId="131.[189,1399,1929,1990]" pageId="131" pageNumber="132">
<emphasis id="B97EEABDFF47FF6AFF641AD6FE93A18A" bold="true" box="[189,294,1929,1955]" pageId="131" pageNumber="132">Fig. 114.</emphasis>
<taxonomicName id="4C0A4D2CFF47FF6AFEF41AD6FCAEA18A" authority="(Wagner, 1906)" baseAuthorityName="Wagner" baseAuthorityYear="1906" box="[301,795,1929,1955]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="131" pageNumber="132" phylum="Mollusca" rank="species" species="kleciaki" subGenus="Turritus">
<emphasis id="B97EEABDFF47FF6AFEF41AD6FE7DA18A" box="[301,456,1929,1955]" italics="true" pageId="131" pageNumber="132">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFF47FF6AFE001AD5FE5AA18A" box="[473,495,1930,1955]" italics="true" pageId="131" pageNumber="132">T.</emphasis>
)
<emphasis id="B97EEABDFF47FF6AFE261AD6FDEFA18A" box="[511,602,1929,1955]" italics="true" pageId="131" pageNumber="132">kleciaki</emphasis>
(
<bibRefCitation id="EF9B4B5EFF47FF6AFDB01AD6FCA4A18A" author="Wagner" box="[617,785,1929,1955]" firstAuthor="Wagner" pageId="131" pageNumber="132" pagination="121 - 140" refId="ref38295" refString="Wagner A. J. 1906. Neue Formen und Fundorte der Genera Pomatias Studer und Auritus Westerlund. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 38 (2 - 3): 92 - 101, 121 - 140." type="journal article" year="1906">Wagner, 1906</bibRefCitation>
)
</taxonomicName>
, lectotype (male), SMF-160836/1, Hvar Island, HR (SMF-160836/1).
</paragraph>
</caption>
<subSubSection id="C3106524FF40FF6DFF641C56FD9EA443" pageId="132" pageNumber="133" type="discussion">
<paragraph id="8BB536AFFF40FF6DFF641C56FE9AA70A" blockId="132.[189,303,265,291]" box="[189,303,265,291]" pageId="132" pageNumber="133">
<heading id="D0FD81C3FF40FF6DFF641C56FE9AA70A" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="132" pageNumber="133" reason="3">
<emphasis id="B97EEABDFF40FF6DFF641C56FE9AA70A" bold="true" box="[189,303,265,291]" pageId="132" pageNumber="133">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFF40FF6DFF641C68FD9EA443" blockId="132.[189,1399,311,619]" pageId="132" pageNumber="133">
The
<typeStatus id="54B1880DFF40FF6DFF2F1C68FED7A778" box="[246,354,311,337]" pageId="132" pageNumber="133" type="lectotype">lectotype</typeStatus>
is a male specimen. The female specimen in Wagners collection (
<figureCitation id="13312A2AFF40FF6DFB7B1C68FABCA77B" box="[1186,1289,311,338]" captionStart="Fig" captionStartId="132.[189,230,1999,2025]" captionTargetBox="[196,1387,657,1976]" captionTargetId="figure-138@132.[196,1387,657,1976]" captionTargetPageId="132" captionText="Fig.115. Cochlostoma(T.)kleciaki(Wagner,1906),MIZPAS-W7082,2-Makarska,HR(MIZPAS-W7082)." figureDoi="http://doi.org/10.5281/zenodo.10853941" httpUri="https://zenodo.org/record/10853941/files/figure.png" pageId="132" pageNumber="133">Fig. 115</figureCitation>
) is from Makarska, a Croatian city on the coast opposite Hvar Island. The female genital morphology of the specimens from Županje Selo, Pelješac peninsula, south of Hvar Island (
<collectingCountry id="F31D763FFF40FF6DFBE21C22FB22A7BE" box="[1083,1175,381,407]" name="Croatia" pageId="132" pageNumber="133">Croatia</collectingCountry>
), here assumed as representative of the species, show clade B features. The similarity with
<taxonomicName id="4C0A4D2CFF40FF6DFBE51CFEFB53A793" authorityName="Cochl." baseAuthorityName="Westerlund" baseAuthorityYear="1879" box="[1084,1254,417,443]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="132" pageNumber="133" phylum="Mollusca" rank="species" species="nanum" subGenus="Turritus">
<emphasis id="B97EEABDFF40FF6DFBE51CFEFBE2A792" box="[1084,1111,417,443]" italics="true" pageId="132" pageNumber="133">C.</emphasis>
(
<emphasis id="B97EEABDFF40FF6DFBB51CFEFB37A793" box="[1132,1154,417,442]" italics="true" pageId="132" pageNumber="133">T.</emphasis>
)
<emphasis id="B97EEABDFF40FF6DFB4F1CFEFB53A793" box="[1174,1254,417,442]" italics="true" pageId="132" pageNumber="133">nanum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A4D2CFF40FF6DFAFE1CFEFE97A7F7" authorityName="Cochl." baseAuthorityName="Hirc" baseAuthorityYear="1881" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="132" pageNumber="133" phylum="Mollusca" rank="species" species="stossichi" subGenus="Turritus">
<emphasis id="B97EEABDFF40FF6DFAFE1CFEFAF7A792" box="[1319,1346,417,443]" italics="true" pageId="132" pageNumber="133">C.</emphasis>
(
<emphasis id="B97EEABDFF40FF6DFA8E1CFEFAD8A793" box="[1367,1389,417,442]" italics="true" pageId="132" pageNumber="133">T.</emphasis>
)
<emphasis id="B97EEABDFF40FF6DFF641C9BFE97A7F7" box="[189,290,452,478]" italics="true" pageId="132" pageNumber="133">stossichi</emphasis>
</taxonomicName>
was also noticed by Braun who remarked it in the original description (1887: 110111: “Die Form steht den beiden croatischen Arten
<taxonomicName id="4C0A4D2CFF40FF6DFD7A1CB7FCB2A428" authorityName="Westerlund" authorityYear="1879" box="[675,775,488,513]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="132" pageNumber="133" phylum="Mollusca" rank="species" species="nanus">
<emphasis id="B97EEABDFF40FF6DFD7A1CB7FCB2A428" box="[675,775,488,513]" italics="true" pageId="132" pageNumber="133">P. nanus</emphasis>
</taxonomicName>
West, von Udbina bei Gospic und
<taxonomicName id="4C0A4D2CFF40FF6DFB7C1CB7FAC2A428" authority="Cless." authorityName="Cless." box="[1189,1399,487,513]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="132" pageNumber="133" phylum="Mollusca" rank="species" species="stossichi">
<emphasis id="B97EEABDFF40FF6DFB7C1CB7FA9DA428" box="[1189,1320,487,513]" italics="true" pageId="132" pageNumber="133">P. stossichi</emphasis>
Cless.
</taxonomicName>
vom Mte. Risniak in Südcroatien sehr nahe, dürfte aber vorläufig schon wegen des weitentlegenen Fundortes nicht ohne weiteres mit einer derselben identificiert werden.” To be noted that the localities cited by Braun are not correct).
</paragraph>
</subSubSection>
</treatment>
</document>